Tosapusia neudorfensis ( Schaffer, 1898 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 57-58

publication ID	https://doi.org/ 10.11646/zootaxa.4982.1.1
publication LSID	lsid:zoobank.org:pub:58388FB8-128A-4381-83D1-3C508D0D3873
DOI	https://doi.org/10.5281/zenodo.5044044
persistent identifier	https://treatment.plazi.org/id/D446F043-FFCD-FFE4-8C92-FAFCFCFE0434
treatment provided by	Plazi
scientific name	Tosapusia neudorfensis ( Schaffer, 1898 )
status	comb. nov.

Treatment

Tosapusia neudorfensis ( Schaffer, 1898) View in CoL nov. comb.

Figs 19A View FIGURE 19 1 –A View FIGURE 1 2 View FIGURE 2 , B 1 –B View FIGURE 1 2 View FIGURE 2 , C 1 –C View FIGURE 1 2 View FIGURE 2 , D 1 –D View FIGURE 1 2 View FIGURE 2 , 18A View FIGURE 18

* Mitra Neudorfensis View in CoL nov. spec. — Schaffer 1898: 542, fig. 5.

V [exillum]. (U [romitra].) neudorfense (Schff.) View in CoL — Sieber 1958a: 153.

Vexillum svagrovskyi View in CoL sp. nov. — Biskupič 2020: 77, figs. 2–5.

Type material. The syntypes have been deposited in the collection of Natural History Museum Vienna ( Schaffer, 1898) but cannot be located .

Additional material. SNM-PM Z 40040, SL: 29.2 mm, MD: 9.6 mm, Konopiská clay pit at Rohožník, Slovakia, middle Miocene (Badenian), Natural History Museum of Slovak National Museum, Bratislava, Slovakia, illustrated in Biskupič (2020, figs 2A–B), holotype of Vexillum svagrovskyi Biskupič, 2020 , figs 19A 1 –A 2; SNM-PM Z 40045, SL: 40.2 mm, MD: 12.5 mm, paratype of Vexillum svagrovskyi Biskupič, 2020 , illustrated in Biskupič (2020, figs 3C–D), figs 19B 1 –B 2; SNM-PM Z 40046, SL: 30.3 mm, MD: 10.4 mm, paratype of Vexillum svagrovskyi Biskupič, 2020 , illustrated in Biskupič (2020, figs 3E–F), figs 19C 1 –C 2; SNM-PM Z 40053, illustrated in Biskupič (2020, figs 2G–H), figs 19D 1 –D 2. 7 specimens, SNM-PM Z 40041–Z 40047, all from Konopiská clay pit at Rohožník, Slovakia, middle Miocene (Badenian), Natural History Museum of Slovak National Museum, Bratislava, Slovakia, paratypes of Vexillum svagrovskyi Biskupič, 2020 .

Revised description. Shell large, moderately slender fusiform, solid, with weakly gradate spire. Protoconch unknown. Teleoconch of 11 whorls; spire whorls subcylindrical, with impressed suture. Axial sculpture of early teleoconch whorls of densely set opisthocline axial ribs, separated by narrower interspaces. Axial ribs weakening on seventh to ninth teleoconch whorl, becoming slightly more widely spaced and blurred on penultimate and last whorls. Spiral cords most prominent on early teleoconch whorls in axial interspaces, separated by narrow grooves. Broad subsutural band formed by deep spiral groove cutting axial ribs. Spiral band solid, or split into three to five densely spaced spiral cords. All spiral sculpture becoming subobsolete on penultimate and last whorls, rarely subsutural band persisting as weak spiral cord. Last whorl moderately convex, slowly contracting at base, covered by weakly sigmoidal growth lines. Several broad, flat spiral cords with delicate axial threads on weak fasciole. Aperture narrow with indistinct anal sinus; columellar callus indistinct, poorly delimited. Columella with four to five folds, weakening rapidly abapically. Outer lip thin without lirae or denticles. Siphonal canal moderately long and wide, straight, with shallow siphonal notch. [Note that this description is based on the holotype and paratypes of Vexillum svagrovskyi Biskupič, 2020 , except for the fifth columellar fold, which is only known from the syntypes of V. neudorfense ].

Shell measurements and ratios. SL: 28.5–40.2 mm, MD: 9.6–12.5 mm; AA: 30–34°, SL/MD: 3.0–3.2, AL/ AW: 4.3–5.1, AH/S: 1.6–2.2.

Discussion. This species is characterised by the presence of a distinct spiral groove a short distance below the suture that bisects the ribs, resulting in a broad subsutural band.

The syntypes of Tosapusia neudorfensis seem to be lost, but the illustration in Schaffer (1889) is rather precise. Based on that illustration, it is most likely that Vexillum svagrovskyi Biskupič, 2020 is a junior synonym of T. neudorfensis . Both taxa are only known from the upper Badenian Studienka Formation in Slovakia. Both species are identical in shape and sculpture and the only difference is the weak fifth columellar fold of Tosapusia neudorfensis . We assume that the presence of a fifth columellar fold is rather a matter of intraspecific variability.

Palaeoenvironment. Only known from the upper Badenian Studienka Formation, which represents middle to outer neritic depositional environments ( Biskupič 2020).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Devínska Nová Ves, Rohožník–Konopiská ( Slovakia) ( Schaffer 1898; Biskupič 2020).