Thysananthus gottschei (J.B.Jack & Steph.) Stephani (1912a: 787)

Sukkharak, Phiangphak, 2015, A systematic monograph of the genus Thysananthus (Lejeuneaceae, Marchantiophyta), Phytotaxa 193 (1), pp. 448-450 : 448-450

8. Thysananthus gottschei (J.B.Jack & Steph.) Stephani (1912a: 787) . Thysanolejeunea gottschei Jack & Stephani (1892: 20) . Type: PHILIPPINES. Luzon: 1870, Wallis s.n. (holotype G!).

Thysananthus ( Lejeuneaceae , Marchantiophyta) Phytotaxa 193 (1) © 2015 Magnolia Press • 31

Thysananthus borneensis Stephani (1912a: 786) . Type: MALAYSIA. Sarawak: without date, Pearson s.n. (holotype G!); Icon. Steph. nr. 10168.

Thysananthus reversus Stephani (1912a: 789) . Thysanolejeunea reversa Stephani (1896: 139) . Type: PHILIPPINES. Zamboanga del Norte: Dapitan, Micholitz 44 (holotype G!).

Thysananthus rigidus Stephani (1912a: 790) . Type: PAPUA NEW GUINEA. New Ireland [“Nova Hibernia”]: 1893, Micholitz s.n. (holotype G!); Icon. Steph. nr. 10207.

Plants dioicous, with projecting growth, turning upwards and becoming ascending to erect, reddish brown in herbarium specimens, up to 2.6 cm long × 1.6–3 mm wide. Stems rather rigid; ventral merophyte 8–10 cell rows wide; stem in cross section orbicular, 289–320 µm high × 187–240 µm wide, 10–16 cell layers high, composed of 34–44 epidermal cells surrounding 82–121 medullary cells, epidermal cells as large as medullary cells. Leaves strongly imbricate, when dry strongly convolute, when moist strongly convex and ventrad, apical part turned to the ventral side, recurved; dorsal lobe asymmetrically ovate to broadly ovate, 0.7–1.9 × 1.1–1.7 mm, apex mucronate-apiculate to acute, margin entire, dorsal base auriculate, auricle 65–165 × 75–200 µm, ventral margin incurved 2/3 × leaf length; cells elongate-hexagonal with acute ends, vitta absent, marginal cells 7–15 × 4–17 µm, median cells 18–42 × 5–15 µm, basal cells 25–75 × 15–25 µm, trigones cordate, often coalesced, intermediate thickenings 0–2 per cell; oil bodies unknown. Lobules rectangular, 0.3–0.6 × 0.1–0.2 mm, 1/3 × lobe length; appendage on surface of lobule base not developed; keel without appendage or with appendages on one side where leaves and underleaves are free and opposite to adnate ones, appendage always curved towards the stems; lobule apex oblique, free margin continuing into the ventral lobe margin, apex with 1–2 triangular teeth, the first tooth consisting of 3–8 cells, being 2–3 cells wide at base and ending in a row of 1–2(–3) cells, the second tooth of one cell only, often absent. Underleaves imbricate, slightly squarrose, broadly oblong-orbicular to obovate, 0.9–2.1 × 0.6–1 mm, 3–4 × stem width, apex truncate to rounded, plane to recurved, margins entire, bases cuneate, underleaf bases adnate with leaves on one side, on left-hand side for right branches and right-hand side for left branches; cells 12–37 × 7–10 µm. Androecia terminal-intercalary on lateral branches, bracts, and bracteoles in 3–4 pairs, bracts hypostatic, 0.9–1 × 0.7–0.8 mm, apex acute, margins entire; antheridia 2 per bract. Gynoecia with one lejeuneoid innovation forming a monochasial pattern; lobe broadly ovate, 2.1–2.6 × 1.6–1.8 mm, apex acute to apiculate, margin entire; lobules broadly ovate, 1/2–2/3 × lobe length, apex apiculate to obscurely bifid; margins with triangular teeth, the teeth consisting of 5–8 cells, being 2–3 cells wide at base and ending in a row of 1–2 cells; bracteoles spathulate, 1.7–2.4 × 0.7–2.3 mm, apex slightly emarginate, 1/2 × bracteole length with laciniate teeth, the teeth consisting of 3–10 cells, being 2–3 cells wide at base and ending in a row of 1–2 cells, margins recurved. Perianths oblong-cylindrical, 1.7–2.5 × 0.8–1 mm long, keels in upper 1/3 with numerous laciniate teeth, the teeth consisting of 4–15 cells, being 2–3 cells wide at base and ending in a row of 2–6 cells; beak 25–50 µm (3–4 cells) in length or absent.