Thryptocerina Jeannel, 1949, 1916

Will, Kipling & Gueorguiev, Borislav, 2021, Phylogenetic systematics of the genera of Thryptocerina Jeannel, 1949 and new species from New Caledonia (Coleoptera, Carabidae, Oodini), ZooKeys 1044, pp. 375-425 : 375

Thryptocerina Jeannel, 1949

Thryptocerini Jeannel, 1949: 775, 829, 841. Type genus: Thryptocerus Chaudoir, 1878: 74.

Diagnosis.

Male protarsomeres 2-4 eccentrically attached to the preceding tarsomere (with basal axis of the former affixed on lateral 1/2-1/3 of the latter); squamose setae only on apical 1/3 or 1/2 of ventral surface of male protarsomere 1 (except in Adelopomorpha species that have narrow, symmetrically attached male protarsomeres 2-4 lacking any such setae, a derived loss of the two characteristics). Pronotum without setiferous punctures near the hind angles. Parascutellar seta present, puncture to accommodate it rather large, foveate. Abdominal ventrites 1-5 without ambulatory setae (present as a reversal on ventrites 3-5 only in Coptocarpus erwini ).

Recognition among the Carabidae of New Caledonia. All oodines are distinguished by having the elytral intervals 7 and 8 fused posteriorly, forming a ridge over a deeply impressed stria 8 that extends to the suture, and having the metepisternum laterally coadunate with the epipleuron. All are broadly oval, very convex (Figs 18-21 , 41 , 42 ), black with the head and legs often reddish brown or rufous, have the flight wings reduced, and the metasternum and metepisternum very short. The general form is similar to some smaller species of Cerabilia Laporte ( Abacetini ) that are found in rainforest leaf litter with oodines in New Caledonia. Cerabilia species are readily separated by the presence of two supraorbital setae, presence of marginal pronotal setae, the lack of an elytral plica, and the male protarsomeres ventrally biseriate squamulose, compared to oodine species with one or no supraorbital and no pronotal setae, a large elytral plica, and the male protarsomere squamulose with the setae forming broad pads or being entirely unmodified.

Figures

Figures 18 - 21. Dorsal habitus of 18 C. erwini sp. nov. 19 holotype of Coptocarpus cyllodinus, with labels 20 C. amieuensis sp. nov. 21 C. lescheni sp. nov. Scale bars: 1.0 mm.

Figures 41, 42. Dorsal habitus of 41 Adelopomorpha tuberculata 42 holotype of A. glabra Heller, with labels. Images of A. glabra by O. Jaeger, reproduced with permission from MTKD.