Therates vitalisi ida Mandl, 1954

Matalin, Andrey V. & Wiesner, Jürgen, 2023, Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae), Zootaxa 5256 (5), pp. 401-433 : 413-414

Therates vitalisi ida Mandl, 1954

Figs 19–21 , 47 , 71, 72 , 103, 104 , 167 .

Therates fruhstorferi ida Mandl, 1954: 159 (Type locality— China, Kuatun , Fukien).

Therates fruhstorferi ida Mandl, 1954 — Mandl 1955: 335; Cassola 1985: 510; Hua 2002: 5.

Therates fruhstorferi vitalisi W. Horn, 1913 — Wiesner 1988: 40, 1992: 92; Lorenz 1998: 38, 2005: 117; Puchkov & Matalin 2003: 115.

Therates vitalisi W. Horn, 1913 — Wang 2009: 110; Putchkov & Matalin 2017: 245; Wiesner 2020: 95.

Type material. HOLOTYPE, ♂ (by original designation)—“Kuatun (2300 m), 27.40n. Br., 117.40ö. L., J. Klapperich, 25.6.1938 (Fukien)” [printed violet label], “Type Therates, Fruhstorferi, Ida m, Ing. K. Mandl ” [printed and handwritten red label], “Holotypus” [printed red label], “ Therates fruhstorferi , ida Mandl” [typed white label], “ MUSEUM KOENIG, BONN ” [printed orange label] ( ZFMK). PARATYPES: 1♀ —“ Kuatun (2300 m) 27.40n. Br., 117.40ö. L., J. Klapperich, 30.5.1938 (Fukien)“ [printed violet label], “ CoType Therates Fruhstorferi Ida m., Ing. K.Mandl “ [printed and hand-written red label], “ Coll. Ing. K. Mandl ” [printed light blue label]; 1♀ —“KUATUN, FUKIEN, China, 6.8.46, (TSCHUNG SEN.)” [printed and hand-written white label], “ParaType Therates Fruhstorferi , ssp. Ida m., Ing. K.Mandl “ [printed and hand-written red label], “ Coll. Ing. K. Mandl ” [printed light blue label]; 1♀ — “KUATUN, FUKIEN, China, 21.7.46, (TSCHUNG SEN.)” [printed and hand-written white label], “ParaType Therates, Fruhstorferi ssp., Ida m., Ing. K. Mandl ” [printed and hand-written red label], “ Coll. Ing. K. Mandl ” [printed light blue label] (all NHMB) .

Addition material. CHINA: 1♀ —Kuatun, Fukien /Tschung Sen/, 6 Sept. 1946, coll. Klapperich ( JW) .

Reference. CHINA, Zhejiang Prov.: Wuyanlin ( Wang 2009).

Diagnosis. This subspecies is distinguished from the nominotypical subspecies by distinctly transverse labrum in male—LW/LL = 1.14 vs. LW/LL = 1.0–1.08 ( Fig. 71 vs. Fig. 65, 69, 70 ), less serrated antennomeres 9–10, as well as red-brown or brown fore- and middle femora.

Redescription. TL = 11.8 mm in male, 12.3–13.0 mm (mean = 12.78 mm, n = 4) in females.

Head shining blue with violet reflection; orbital plates indistinctly striated in posterior third; frons smooth, slightly elevated, wide semi-circular (see above), practically vertical, slightly expanded forward (see at side), frontal sulci deep and poorly convergent in anterior 2/3, clearly divergent in posterior third; occiput slightly concave in anterior margin. Mandibles in male brown underside, yellow-brown topside with light-brown teeth and apical molar; in females black except yellow-brown basal third of topside. Labial palpi yellow-brownish, except brown apical palpomeres; palpomeres 1 and 2 of maxillary palpi light brown, palpomeres 3 and 4 dark brown. Antennae do not extend posteriorly to the shoulders, scape brown on anterior side and black on posterior side; antennomeres 2–5 black with indistinctly blue tinge and brown apices, antennomeres 9 and 10 in male with serrated anterior lower margins; in female less delated with slightly protruding margins ( Fig. 47 ). Labrum distinctly transverse in male( Fig. 71 ), LW/LL = 1.14, indistinctly transverse in females ( Fig. 72 ), LW/LL = 1.0–1.07 (mean = 1.05, n = 4), blackbrownish with medium-sized or small apical yellow spot separated from central apical teeth.

Pronotum shining blue with light violet tinge; indistinctly transverse, PW/PL = 1.0–1.07 (mean = 1.03, n = 5), apical lobe practically equal wide throughout; thorax shining blue with violet reflection.

Fore and middle femora red-brown or brown with blue-tinged base and knees; hind femora black with blueviolet lustre except pale-yellow base; fore and middle tibiae red-brown; hind tibiae black or brown-black, all tibia with blue tinge; tarsi black with light blue tinge; HTbL/HTaL = 1.21–1.27 (mean = 1.24, n = 5).

Elytra black-blue with purple reflection, practically sub-parallel, EL/EW = 1.98–2.1 (mean = 2.02, n = 5); punctuation in basal half except humeral area deep and regular, then shallow and sparse, apical quarter practically smooth; scutellum black with blue tinge; apical margin indistinctly cut. Elytral pattern presented by small basal portion of humeral lunule, small basal dot poorly visible in some specimens and elongated slightly oblique central dot ( Figs 19, 21 , 103, 104 ).

Aedeagus with short sharply curved blunt apex and distinct lateral carinae ( Fig. 167 ); AL = 2.9, EL/AL = 2.52.

Distribution. CHINA: Fujian, Zhejiang Provinces ( Fig. 184 ).

Figures

FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png.

FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels.

FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales.

FIGURES 65–72. Therates vitalisi ssp. labrum: 65–70—T. vitalisi vitalisi (65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—T. vitalisi ida (China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype.

FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes

FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm.