Thaumatosmylus Krüger, 1913a

Winterton, Shaun L., Martins, Caleb Califre, Makarkin, Vladimir, Ardila-Camacho, Adrian & Wang, Yongjie, 2019, Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581 (1), pp. 1-99 : 65-66

Thaumatosmylus Krüger, 1913a

( Fig. 43–44 )

(= Glenosmylus Krüger, 1913a: 91 . Type species. Glenosmylus elegans Krüger, 1913a: 91 ) syn. n.

Type species. Osmylus diaphanus Gerstaecker, 1894: 168 (by original designation).

Diagnosis. Ocelli present; wings broad, ovoid to sub-triangular with expanded posterior margin at mid-length of wings; wings hyaline with sparse to dense patterning especially in FW; FW costal area relatively broad along entire length; subcostal veinlets simple and densely arranged; stem of RP short, RP with 11-15 branches, slightly sinuous distally; crossveins numerous and semi-regularly arranged in basal half of wings, more regular distally with outer gradate series distinct from other crossveins while inner gradate series (when present) is often not distinct from basal crossveins; end-twigging regular and close to wing margin in radial and medial fields of both wings, endtwigging less extensive in cubital field; M vein forked before of the origin of t RP1; FW with 2–4 m-cu crossveins before fork of M; FW lacking embossed bullae, and fenestrate areas absent in infuscate areas of wing; male genitalia with ectoproct with one or two pairs of rounded lobes; gonarcus and entoprocessus narrow and joined by extensive membrane area; female spermatheca shape variable, often with only a few lobes.

Comments. Thaumatosmylus is distinguished by the presence of 2–4 m-cu crossveins before the fork of M in the forewing. Other characteristics include the lack of fenestrate spots in wing markings and lack of embossed bullae. Species from continental regions have relatively narrower wings, while those found on islands (e.g., Malaysia and Indonesia) tend to have broader wings. Glenosmylus elegans Krüger is moved to this genus based on the crossveins in the M-Cu area while G. kruegeri is transferred to Thyridosmylus (see below). Spilosmylus fraternus (Banks) and S. conspersus (Walker) are transferred from Spilosmylus since they have more than one forewing m-cu crossvein before the fork of MA-MP and lack an embossed spot along the posterior forewing margin.

Figures

FIGURE 43. Spilosmylinae. Thaumatosmylus delicatus Banks.

FIGURE 44. Spilosmylinae. Thaumatosmylus hainanus Yang (photo: Shaun L. Winterton).