Thaumatodryinus miocenicus Olmi, 1995

Martins, Andre L. & Melo, Gabriel A. R., 2020, Revision of the fossil species of Thaumatodryinus Perkins from Dominican amber, with a new combination and description of a new species (Hymenoptera, Dryinidae), Journal of Hymenoptera Research 79, pp. 77-88 : 77

† Thaumatodryinus miocenicus Olmi, 1995

Figure 2

Thaumatodryinus miocenicus Olmi, 1995: 263. Female holotype. Dominican Republic: amber from unknown mine, probably El Valle mine (AMNH, examined).

Extended diagnosis.

Female holotype, 5.6 mm (see fig. 1-2 in Olmi (1995) and fig. 140 in Olmi and Virla (2014)). Thaumatodryinus miocenicus is easily recognized by the following combination of characters: black head, except antenna brown to dark brown; mesosoma mostly black; legs and metasoma apparently brown. Antenna with dense and short setae. Head slightly convex, granulated; upper frons and vertex with pilosity; occiput slightly excavated and densely hairy; frontal line present but apparently incomplete in holotype (complete in paratype Do-4066-M); mid portion of clypeus with anterior margin convex. Mesoscutum reticulate rugose; notauli incomplete, reaching approximately half the length of mesoscutum; mesoscutellum reticulate rugose, hairy and shorter than mesoscutum; anterior margin of mesoscutellum with a foveate narrow groove; metanotum reticulate rugose, shorter than scutellum. Propodeum apparently reticulate rugose and short, distinctly shorter than high in lateral view; fore wing overlapped in holotype and color not visible, but in paratypes, fore wing with one dark transverse band beneath pterostigma. Ocellar ratio: POL= 7; OL= 3; OOL= 10; OPL= 2; TL= 3. Antennomeres in following proportions: 14:7:22:24:34:20:19 (flagellomere 8-10 not visible). Eye somewhat elongate, about 1.3 × longer than wide; maximum head width about 1.7 × distance between inner margins of tegulae. Pronotum shorter than head (16:37). Mesoscutum longer than pronotum (20:16). Fore wing with 2r-rs shorter than 3Rs&4Rs. Fore leg segments in following proportions: 30 (coxa): 15 (trochanter): 40 (femur): 43 (tibia): 22 (1th tarsomere): 3 (2nd tarsomere): 7 (3rd tarsomere): 22 (4th tarsomere): 36 (5th tarsomere); 5th tarsomere longer than enlarged claw and with two rows of at least 30 lamellae; distal apex with group of approximately three long lamellae; enlarged claw with two subapical teeth and one row of 29 lamellae. Propodeum short, distinctly shorter than high in lateral view, with dorsal surface shorter than posterior surface (45:50).

Remarks.

While in the original description Olmi (1995) mentioned that the mandibles are "not distinctly visible", Olmi and Virla (2004) stated that the mandibles are “tridentate”. We could not clearly see the condition in the type material but consider more likely that T. miocenicus has quadridentate mandibles as in all other Thaumatodryinus . When the mandibles are closed, it is quite hard to observe the tooth closer to the clypeus, giving the impression that the mandibles are tridentate.

Examined material.

Female holotype, in amber from the Dominican Republic: amber from an unknown mine (probably El Valle mine) DR-99-146 HT (AMNH). Two paratype females: same locality label as holotype, DR-99-147 PT and DR-99-148 PT (AMNH) .

Figures

Figure 2. † Thaumatodryinus miocenicus Olmi, 1995, female holotype. A habitus, dorsal view B habitus, ventral view C habitus, lateral view D habitus, dorsal view E head and mesosoma, ventral view F habitus, lateral view. Scale bars: 1 mm (A-C), 1 mm (D, F), 1 mm (E).