Leptotyphlus tuniseus, Krasa, 1941

Herman, Lee H., 2001, Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the End of the Second Millennium IV. Staphylinine Group (Part 1) Euaesthetinae, Leptotyphlinae, Megalopsidiinae, Oxyporinae, Pseudopsinae, Solieriinae, Steninae, Bulletin of the American Museum of Natural History 2440 (265), pp. 1807-2440 : 1807-2440

tuniseus Krása, 1941 , see: mendicus Erichson, 1840 .

turbulentus Bondroit, 1912a: 417 ( Stenus ; Type locality: Athos , Macédoine).

— L. Benick, 1917c: 60 ( Stenus ; notes).

— L. Benick, 1929a: 70 ( Stenus ; subgenus Parastenus ; characters in key).

— Scheerpeltz, 1933: 1198 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1972s: 270 ( Stenus ; notes).

— Puthz, 1979j: 326 ( Stenus ; Turkey).

— H. Franz, 1984: 59 ( Stenus ; Macedonia).

— H. Franz, 1987: 69 ( Stenus ; Cyprus).

— Assing and Wunderle, 2001: 36 ( Stenus ; Cyprus).

- DISTRIBUTION: Macedonia, Turkey, Cyprus.

turcicus Bernhauer, 1912 k: 260 ( Stenus ; Type locality: Berg Athos in Macedonien; Thessalien: Volo; Kleinasien : Alhem Dagh ).

— L. Benick, 1929a: 63 ( Stenus ; subgenus Parastenus ; characters in key).

— Scheerpeltz, 1933: 1198 ( Stenus ; subgenus Hemistenus ; catalog).

— Puthz, 1972s: 276 ( Stenus ; subspecies of cribratus ; notes).

— Puthz, 1981i: 701 ( Stenus ; note; valid species).

— Ryvkin, 1990: 211 ( Stenus ; cited as valid species; notes; Turkey).

- DISTRIBUTION: Macedonia, Greece, Turkey.

turgidicollis Puthz, 1986 k: 954 ( Stenus ; subgenus Nestus ; Type locality: Pakistan: Swat: au-dessus de Miandam, 2300 m).

- DISTRIBUTION: Pakistan.

turk Puthz, 1972 l: 178 ( Stenus ; [Note: This species is the “ cordatoides A” reported and characterized in Puthz, 1972s: 280]; Type locality: no type locality cited; [Note: Several localities are listed for Turkestan, Iran, and the Caucasus in the 1972s report]).

— Ryvkin, 1990: 213 ( Stenus ; notes; Georgia; Azerbaijan; Iran).

— Betz, 1996: 25 ( Stenus ; labial adhesion capture apparatus).

- DISTRIBUTION: Turkestan, Georgia, Azerbaijan, Iran, Turkmenistan, Uzbekistan. turkestanicus Bernhauer and Schubert, 1911, see: bernhauerianus Jakobson, 1909 .

turneri Bernhauer, 1934 , see: uluguruensis Bernhauer, 1915 .

turneri Cameron, 1950 , see: kenyanensis Puthz, 1965 .

turnerianus Cameron, 1950a: 182 ( Stenus ; subgenus Nestus ; Type locality: Kenya Colony: Londiani, 7000 feet).

— Puthz, 1971: 145 ( Stenus ; characters; Kenya; Tanzania; Congo).

- DISTRIBUTION: Kenya, Tanzania, Congo.

tylocephalus Kraatz, 1857 , see: guynemeri Jacquelin du Val, 1850 .

tythus Schaufuss, 1882 , see: melanopus Marsham, 1802 .

uenoi Naomi and Nomura, 1990: 46 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Nagano Pref.: Odona, Neba-mura).

- DISTRIBUTION: Japan.

ukereweanus Bernhauer, 1937, see: pilus Schubert, 1911 .

ulcerosus L. Benick, 1929: 95 ( Stenus ; subgenus Hemistenus ; Type locality: Binaluan, Palawan).

— Scheerpeltz, 1933: 1198 ( Stenus ; subgenus Hemistenus ; catalog).

- DISTRIBUTION: Philippines.

uluguruensis Bernhauer, 1915d: 111 ( Stenus ; subgenus Hypostenus ; Type locality: Deutsch-Ostafrika: Süd-Uluguru).

— Scheerpeltz, 1933: 1184 ( Stenus ; subgenus Hypostenus ; cited as ulugurensis; catalog).

— Cameron, 1950g: 22 ( Stenus ; subgenus Hypostenus ; Belgian Congo).

— Puthz, 1967i: 236 ( Stenus ).

— Puthz, 1971: 273 ( Stenus ; subgenus Hypostenus ; characters; Tanzania; Kenya; Zaire).

— Puthz, 1976b: 341 ( Stenus ; Tanzania).

- DISTRIBUTION: Sierra Leone, Guinea, Kenya, Tanzania, Congo, South Africa. lugubris L. Benick, 1923: 249 ( Stenus ; Type locality: Amani, Deutsch-Ostafrika).

— Scheerpeltz, 1933: 1179 ( Stenus ; subgenus Hypostenus ; catalog).

— Puthz, 1965: 543 ( Stenus ; subgenus Hypostenus ; characters).

— Puthz, 1967i: 236 ( Stenus ; synonym of uluguruensis ).

— Puthz, 1971: 273 ( Stenus ; synonym of uluguruensis ).

gramineus L. Benick, 1932: 241 ( Stenus ; Type locality: Pietermaritzburg).

— Scheerpeltz, 1933: 1178 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1933: 302 ( Stenus ; characters in key).

— Puthz, 1967i: 236 ( Stenus ; synonym of uluguruensis ).

— Puthz, 1967k: 9 ( Stenus ; synonym of uluguruensis ).

— Puthz, 1971: 273 ( Stenus ; synonym of uluguruensis ).

turneri Bernhauer, 1934 h: 499 ( Stenus ; subgenus Hypostenus ; Type locality: Zululand: Eshowe; Natal: Kloof).

— Puthz, 1965: 555 ( Stenus ; subgenus Hypostenus ; characters; South Africa).

— Puthz, 1967i: 236 ( Stenus ; synonym of uluguruensis ).

— Puthz, 1967k: 9 ( Stenus ; synonym of uluguruensis ).

— Puthz, 1971: 273 ( Stenus ; synonym of uluguruensis ).

anchisteus Puthz, 1971: 274 ( Stenus ; subgenus Hypostenus ; subspecies of uluguruensis ; Type locality: Sierra Leone: Monts Loma, ravin S.E. du Camp II, 1600 m. Also cited from Guinea).

umbifrons Puthz, 1967 j: 35 ( Stenus ; subgenus Hypostenus ; Type locality: Ruanda: Ruhengeri, riv. Penge, 1800–1825 m).

— Puthz, 1971: 262 ( Stenus ; subgenus Hypostenus ; characters; Rwanda).

- DISTRIBUTION: Rwanda.

umbratilis Casey, 1884: 156 ( Areus ; Type locality: Lake Quesnel , British Columbia).

— Bernhauer and Schubert, 1911: 170 ( Stenus ; subgenus Tesnus ; catalog).

— Van Dyke, 1924: 17 ( Stenus ; synonym of fraternus ).

— Sanderson, 1946: 428 ( Stenus ; characters; Alberta; Manitoba; Northwest Territory; Quebec). — Sanderson, 1957: 250 ( Stenus ; subgenus Hemistenus ; characters; British Columbia).

— Szujecki, 1961: 50 ( Stenus ; subgenus Hemistenus ; characters; Poland).

— Horion, 1963: 377 ( Stenus ; subgenus Hemistenus ; Russia; Finland; Sweden; Norway; Britain; Denmark; Germany).

— Lohse, 1964: 123 ( Stenus ; subgenus Hemistenus ; characters; central Europe).

— Puthz, 1967l: 50 ( Stenus ; notes).

— Puthz, 1971v: 105 ( Stenus ; notes).

— Puthz, 1972d: 88 ( Stenus ; subgenus “ Hemistenus ”; characters).

— Puthz, 1973o: 129 ( Stenus ; Britain).

— Pope, 1977: 26 ( Stenus ; Britain).

— Anderson, 1984: 249 ( Stenus ; collecting notes; Ireland).

— Lucht, 1987: 97 ( Stenus ; checklist; Germany; Poland; Denmark).

— Lohse and Lucht, 1989: 145 ( Stenus ; notes).

— Campbell and Davies, 1991: 109 ( Stenus ; USA; Canada).

— Hromádka, 1993: 45 ( Stenus ; subgenus Hemistenus ; Slovakia).

— Hodge and Jones, 1995: 35 ( Stenus ; characters; Britain).

— Assing and Puthz, 1998: 131 ( Stenus ; Slovakia).

— Burakowski, Mroczkowski, and Stefańska, 2000: 45 ( Stenus ; subgenus Hemistenus ; catalog; Poland).

- DISTRIBUTION: Canada, USA, Europe.

pseudopubescens A. Strand, 1940: 180 ( Stenus ; subgenus Hemistenus ; Type locality:

numerous localities in Norway, Sweden, Denmark, and Germany).

— L. Benick, 1941e: 136 ( Stenus ; characters; variation).

— Renkonen, 1941: 105 ( Stenus ; characters; notes).

— Tottenham, 1948: 188 ( Stenus ; subgenus Hemistenus ; characters; Britain).

— Steel, 1953c: 198 ( Stenus ; synonym of umbratilis ).

— Szujecki, 1961: 9 ( Stenus ; synonym of umbratilis ).

— Lohse, 1964: 123 ( Stenus ; synonym of umbratilis ).

— Puthz, 1967l: 50 ( Stenus ; synonym of umbratilis ).

— Puthz, 1972d: 90 ( Stenus ; synonym of umbratilis ).

— Pope, 1977: 26 ( Stenus ; synonym of umbratilis ).

umbricus Baudi, 1870: 395 ( Stenus ; Type locality: Italia media; [ Note : See lectotype designation by Puthz, 1970p]. Originally cited from Umbriae Trasimeni lacus).

— Fauvel, 1873: 94 [= 1873a: 252] ( Stenus ; characters; Italy).

— Ganglbauer, 1895: 572 ( Stenus ; subgenus Nestus ; characters; Italy).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— L. Benick, 1929a: 51 ( Stenus ; subgenus Nestus ; characters in key).

— Puthz, 1965f: 26 ( Stenus ; characters; note).

— Puthz, 1970p: 171 ( Stenus ; lectotype designation: Italia media; notes) .

— Puthz, 1971v: 96 ( Stenus ; notes).

— Bordoni, 1975c: 181 ( Stenus ; collecting notes; Italy).

— Lucht, 1987: 96 ( Stenus ; checklist; Czechoslovakia; Austria).

— Puthz and Zanetti, 1995: 19 ( Stenus ; Italy).

- DISTRIBUTION: Italy.

umbrosus L. Benick, 1928: 20 ( Stenus ; Type locality: Ost-Afrika, Nandi Reservat ).

— Scheerpeltz, 1933: 1167 ( Stenus ; subgenus Stenus ; catalog).

— Puthz, 1971: 103 ( Stenus ; characters; Kenya; Uganda).

- DISTRIBUTION: Uganda, Kenya, Rwanda, Congo.

capriciosus Tottenham, 1956: 233 ( Stenus ; Type locality: Ruanda: terr. Nyanza , Gitarama, 1850 m).

— Puthz, 1971: 107 ( Stenus ; subspecies of umbrosus ; characters; Rwanda).

nyamuragiraensis Puthz, 1967 j: 20 ( Stenus ; subgenus Stenus ; subspecies of gedyei ; Type locality: Nyasheke, volc. Nyamuragira, 1820 m).

— Puthz, 1971: 105 ( Stenus ; subspecies of umbrosus ; characters; Congo).

londianiensis Puthz, 1967 j: 21 ( Stenus ; subgenus Stenus ; subspecies of gedyei ; Type locality: Londiani, K. C.).

— Puthz, 1971: 106 ( Stenus ; subspecies of umbrosus ; characters; Kenya).

luberoensis Puthz, 1971: 107 ( Stenus ; subspecies of umbrosus ; Type locality: Congo: Kivu: Territoire Lubero, Mont Bugera, 2100 m).

itombweensis Puthz, 1971: 109 ( Stenus ; subspecies of umbrosus ; Type locality: Congo: Kivu: Territoire Mwenga, S.O. Itombwe, Luiko, 1900 m).

unagi Hromádka, 1979d: 105 ( Stenus ; subgenus Parastenus ; Type locality: Japan: Pref. Osaka: Mt. Iwawaki).

— Naomi, 1988h: 68 ( Stenus ; characters; Japan).

— Puthz, 1993a: 145 ( Stenus ; subgenus Hypostenus ; characters; Japan).

- DISTRIBUTION: Japan.

ookami Hromádka, 1979d: 106 ( Stenus ; subgenus Parastenus ; Type locality: Japan: Pref. Osaka: Mt. Iwawaki).

— Naomi, 1988h: 68 ( Stenus ; characters; Japan).

— Puthz, 1993a: 145 ( Stenus ; synonym of unagi ).

uncifer Puthz, 1967 , see: convergens L. Benick, 1933 .

undulatus Wollaston, 1854: 599 ( Stenus ; Type locality: Madeira: Cruzinhas at about 5000 ft.; [Note: See lectotype designation by Puthz, 1966g]).

— Wollaston, 1857: 197 ( Stenus ; habitat; Madeira Islands).

— Fauvel, 1902b: 74 ( Stenus ; Madeira).

— Bernhauer and Schubert, 1911: 168 ( Stenus ; catalog).

— Smetana, 1963b: 33 ( Stenus ; subgenus Stenus ; Madeira).

— Puthz, 1966g: 130 ( Stenus ; lectotype designation; notes; characters).

- DISTRIBUTION: Madeira.

uneme Naomi, 1989d: 8 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Ehime Pref.: Omogokei).

— Puthz, 1993a: 164 ( Stenus ; subgenus Parastenus ; notes; Japan).

- DISTRIBUTION: Japan.

unguinosus Ryvkin, 1992 , see: deliculus Ryvkin, 1992 .

unguliventris Puthz, 1998: 388 ( Stenus ; subgenus Parastenus ; Type locality: Vietnam:

Tam Dao. Also cited from India).

- DISTRIBUTION: Vietnam, India.

unicolor Stephens, 1833 , see: nigritulus Gyllenhal, 1827 .

unicolor Erichson, 1840 , see: brunnipes Stephens, 1833 .

unicus Bernhauer, 1915 , see: abdominalis Fauvel, 1895 .

uniformis Fauvel, 1904e: 312 ( Stenus ; Type locality: Madagascar: Fort Dauphin ; [Note: See lectotype designation by Puthz, 1967i]. Also cited from Madagascar: Diego-Suarez; vallee du Fanjahira: Isaka; Suberbieville).

— Bernhauer and Schubert, 1911: 178 ( Stenus ; subgenus Hypostenus ; catalog).

— Scheerpeltz, 1961g: 235 ( Stenus ; subgenus Hypostenus ; Madagascar).

— Puthz, 1967i: 245 ( Stenus ; characters; lectotype designation: [ Note : Locality not cited but in Puthz, 1972: 9, Fort Dauphin is cited as the locality for the lectotype]; Madagascar) .

— Puthz, 1972: 9 ( Stenus ; characters; Madagascar).

— Puthz, 1992a: 15 ( Stenus ; Madagascar).

- DISTRIBUTION: Madagascar.

unzenmontis Naomi and Puthz, 1996: 156 ( Stenus ; Type locality: Japan: Nagasaki Pref.:

Mt. Unzen).

- DISTRIBUTION: Japan.

upembaensis Puthz, 1971: 282 , 283 ( Stenus ; subgenus Hypostenus ; Type locality: Congo: Parc National de l’Upemba, Gorges de la Pelenge).

— Puthz, 1972a: 6 ( Stenus ; subgenus Hypostenus ; [Note: Cited as a new species but the 1971 citation preceded this publication]; characters; Zaire).

- DISTRIBUTION: Congo.

usambarae Bernhauer, 1915 , see: jovino Eichelbaum, 1913 .

ussuriensis Ryvkin, 1987c: 150 ( Stenus ; subgenus Stenus ; Type locality: USSR: Far East, Maritime Prov.: Ussurijsk Distr., nr. Kamenushka village).

- DISTRIBUTION: Russia.

utan Naomi, 1998: 388 ( Stenus ; subgenus Hypostenus ; Type locality: Japan: Kanagawa Pref.: Kamiyama, Hakone).

- DISTRIBUTION: Japan.

utenis Casey, 1884 , see: pinguis Casey, 1884 .

utzungweanus Bernhauer, 1915d: 110 ( Stenus ; Type locality: Deutsch-Ostafrika: Utzungwe-Berge , 1500–1600 m).

— Scheerpeltz, 1933: 1184 ( Stenus ; subgenus Hypostenus ; catalog).

— L. Benick, 1951: 202 ( Stenus ; characters).

— Puthz, 1965: 493 ( Stenus ; subgenus Hypostenus ; characters; Tanzania; Zaire; Somalia).

— Puthz, 1967i: 193 ( Stenus ; characters in key; Kenya; Congo; Tanzania) .

— Puthz, 1971: 210 ( Stenus ; subgenus Hypostenus ; Tanzania; Somalia; Kenya; Zaire; Rwanda). — Puthz, 1972a: 4 ( Stenus ; characters; Zaire).