Teloconus Grouvelle, 1916: 234

Kirejtshuk, Alexander G. & Kovalev, Alexey V., 2022, Monograph on the Cillaeinae (Coleoptera: Nitidulidae) from the Australian Region with comments on the taxonomy of the subfamily, Zootaxa 5103 (1), pp. 1-133 : 13-15

1. Teloconus Grouvelle, 1916: 234

—Patagonian Region ( Argentina).

The type species of this genus remains without re-examination. The describer mentioned that this genus resembles Gonioryctus with the body 5.0 × as long as wide and glabrous (except for abdomen), rather short head bearing short and convergent antennal grooves, pronotum narrowly bordered at sides and with blunt anterior and posterior angles, elongate striate elytra and conical abdomen with long ultimate segment. The detailed original description of both genus and species ( Teloconus mirificus Grouvelle, 1916 ) seems to make it possible to recognize this species in additional new specimens when they are available. The type species should be deposited in the Museo de la Plata (Facultad de Ciencias Naturales y Museo, Buenos Aires) and Muséum national d’Histoire naturelle (Paris), however, at least in the latter collection no specimen of this species has been found.

Addition to standard measurements (length of body, width in widest part of body, and hight in thorax) in this monograph the length without abdomen (from apex of frons to apices of elytra) which is frequently important when the following abdominal segments can be retracted into the previous segment to varying degree.

2. Key to cillaeine genera and subgenera of Australia and surrounding territories (adults)

1a. Protibia with two strong and widely separated teeth at outer apical angle; meso- and metatibiae with long subapical spines along apical and frequently outer edge;

(1a) Remarks. Body more and less regularly convex; head with eyes located at base, temples scarcely raised or sharply projecting laterally; anterior and posterior angles of pronotum more or less widely rounded; laterosternites V and VI moderately to strongly widened to apex;—apex of male anal sclerites and female pygidium without distinct serration. 2.8–3.4 mm. Figs 36–38 ......................................................................................... Oniphenes gen. nov.

1b. Protibia without two separated teeth at outer apical angle or with teeth gradually increasing before apex, outer apical angle at most rather projecting; meso- and metatibiae usually without long subapical spines along outer edge................... 2

2(1)a. Body markedly wider (slightly more than 2.0 × as long as wide), regularly and evenly convex dorsally and distinctly concave ventrally, subelliptic in outline; scape with long anteroapical process (markedly longer than scape); pronotum with deeply sinuate base; elytra widely explanate at sides and about 0.7 × as long as combined width; elytral epipleura widely and steeply sloping externally; body with strongly widened abdomen behind elytral apices;

2(1)a Remarks. Laterosternites V and VI rather strongly widened at apex; prosternal process medially roof like; pronotal, elytral and abdominal sides densely ciliate; all coxae narrowly separated; tarsomeres 1-3 very widely lobed. 4.9–6.9 mm. Figs 34 , A–I..................................................................................... Onicotis Murray, 1864

2(1)b. Body at most moderately wide (at least 3.0 × as long as wide), not regularly and evenly convex dorsally and somewhat convex ventrally, not subelliptic; scape of usual shape; pronotum with subtruncate somewhat convex base or at most slightly emarginate at posterior pronotal angles; elytra not, narrowly or moderately explanate at sides and about as long as combined width or longer; outlines of antennal grooves somewhat curved; elytral epipleura subhorizontal to somewhat elevated externally; abdomen at most only slightly widened behind elytral apices..... .............................................. 3

3(2)a. Tarsomeres 1–3 subcylindrical and comparatively long with setae on lower surface and tarsomere 1 thicker and usually longer than each of tarsomeres 2–4; anterior part of frons of both sexes not subtruncate (more or less modified); antennal grooves absent or at most represented in depressions along inner edge of eyes which are not distinctly outlined;

3(2)a Remarks. Exposed abdominal tergites with strong postbasal line reaching apices of tergites and laterosternites rather narrow, usually slightly widened at apex to subparallel-sided or moderately widened; eyes close to anterior part of frons; apices of male anal sclerite and female pygidium with distinct serration; median part of prosternum and prosternal process forming one plane, flat apex of process very wide and with subtruncate to widely rounded posterior edge. 4.8–7.8 mm. Figs 28–30 ............................................................................................ Ithyphenes Murray, 1864

3(2)b. Tarsomeres 1–3 distincly lobed and tarsomere 1 usually not wider and not longer than each of tarsomeres 2–4; anterior part of frons subtruncate (anterior frontal edge of some males of Adocimus (Belonotus) bartenevi sp. nov. with a pair of feeble teeth and a weak emargination between them); antennal grooves well developed at least between genal process and eye........ 4

4(3)a. Eyes located in anterior half of head, whose sides behind eyes not or slightly narrowed; abdominal laterosternites V and VI very narrow and subparallel-sided or distinctly narrowing distally.................................................. 5

4(3)b. Eyes located at midlength or in posterior half of head, whose sides behind eyes distincly narrowed; abdominal laterosternites V and VI wide and distinctly widening posteriorly (except for Adocimus with narrow laterosternites of somewhat different in outline)............................................................................................. 6

5(4)a. Body integument with scarcely conspicuous hairs or almost without them; tergites uncovered by elytra with elongate paramedian depressions; distance between mesocoxae subequal to or smaller than that between procoxae; head with uniform punctures and sculpture interspaces between them on disc of frons, vertex, sides and base; procoxae moderately separated; apex of female pygidium distinctly serrate; antennomeres 2 and 3 comparable in length. 8.6–14.7 mm. Figs 4 , A–F....................................................................................... Allenipeplus Kirejtshuk et Kovalev, 2016

5(4)b. Body integument clearly pubescent; distance between mesocoxae markedly greater than that between procoxae; tergites uncovered by elytra gently convex; head sides and base with clearly weakened punctation and smoothed interspaces between punctures in contrast to disc of frons and vertex; procoxae very narrowly separated; apex of female pygidium feebly serrate; antennomere 2 markedly longer than antennomere 3. 3.5–6.0 mm. Figs 31 , A–J................... Laferollaeus gen. nov.

6(4)a. Abdominal laterosternites V and VI narrow and distinctly narrowing posteriorly or subparallel-sided to weakly widening posteriorly.............................................................. .. 7 ( Adocimus Murray, 1864 s. lato)

6(4)b. Abdominal laterosternites V and VI wide and rather widening posteriorly.. ....................................... 8

7(6)a. Prothoracic lateral carinae absent (at most with line instead of carina more or less reduced); elytral punctation sparse and more or less reduced, elytral longitudinal rows of punctures not arranged in striae or, if punctures absent, they may be represented by incomplete longitudinal lines; scutellar shield subpentagonal; temples not longer than frons before eyes; abdominal ventrite 1 markedly longer than ventrite 2; body more robust with pronotum distinctly transverse; male: tibial apices thickened and with short and rather thick spines; female: only protibial apices thickened and with short and rather thick spines. 4.0–11.0 mm. Figs 1, A–K... ............................................................ subgenus Adocimus Murray, 1864 s. str.

7(6)b. Prothoracic lateral carinae distinct; elytral punctation dense and rather deep, elytral longitudinal rows punctures in distinct striae, at least in distal part; scutellar shield subtriangular; temples longer than frons before eyes; abdominal ventrites 1 and 2 comparable in length; body more slender with pronotum subquadrate or elongate; all tibia of both sexes of usual structure. 3.7- 5.1 mm. Figs 2 , A–J......................................................... subgenus Belonotus subgen. nov.

8(6)a. Pronotum with widely rounded anterior and posterior angles; prosternum with anterior edge projecting anteriorly and forming a wide chin piece; male anal sclerite with a deep funnel-shaped invagination in the middle of posterior edge, but without serration;

8(6)a. Remarks. Prosternum and prosternal process forming a unified median plate widening as anteriorly as posteriorly; tarsomeres 1–3 widely lobed; apex of female pygidium without serration. 6.7–8.3 mm. Figs 25 , A–D.......................... ........................................................................ Caledomus Kirejtshuk et Kovalev, 2017

8(6)b. Pronotum at least with distinct tip of posterior angles; prosternum with simple anterior edge (not projecting anteriorly and not forming a chin piece; male anal sclerite without a deep funnel-shaped posterior invagination)......................... 9

9(8)a. Dorsum subconcave (only elytral sides sloping); head, pronotum and dorsal abdominal sclerites densely and uniformly micropunctate or microgranulate; elytra densely and uniformly completely or partly microgranulate; distance between metacoxae subequal with that between mesocoxae and smaller than that between procoxae; all tibiae simple or rather widened in distal half;

9(8)a. Remarks. Pygidium of both sexes longer than wide and widely truncate at apex. 4.0– 6.7 mm. Figs 32–33 ................................................................................................ Matthewsianus gen. nov.

9(8)a. Dorsum convex to subflattened (never concave) and with different sculpture; distance between metacoxae subequal with that between procoxae and smaller than that between mesocoxae; all tibiae narrow or at most gradually widened in distal half (subtriangular)...................................................................................... 10

10(9)a. Labrum unilobed; antennomeres 2–7 about 2.0 × or more as long as wide, and antennal club elongate (at least 2.0 × as long as wide); lateral apical lobes of prosternal process widely rounded; metatibiae at most with one clear external row of spines or setae;

10(9)a Remarks. Pronotum subquadrangular, not arcuately narrowed anteriorly, flattened and with subrectilinear posterior edge; pregenal processes comparatively wide; legs comparatively long; tibiae not subtriangularly dilated but subparallel-sided or rectilinearly narrowing at apex (subtriangularly widened to apex). Figs 23–24. 6.2–15.5 mm............................................................................................... Brittonoma Kirejtshuk et Kovalev, 2016

10(9)b. Labrum bilobed; antennae with shorter flagellomeres (only antennomere 3 sometimes about 2.0 × as long as wide), and regularly oval or subcircular club (not as elongate); lateral apical lobes of prosternal process subangular to narrowly rounded; metatibiae with two rows of spines or setae........................................................................ 11

11(10)a. Elytra about as long as combined width or slightly shorter; pronotum more or less narrowed anteriorly rather than posteriorly, with sides arcuate (not subrectilinear), posterior angles usually with distinct tip, and also with more or less bi-emarginate posterior edge; protibia with evenly small teeth at apex. 3.0–9.0 mm. Figs 5–21 ......... .... Brachypeplus Erichson, 1842

11(10)b. Elytra about 1.5 × as long as combined width; pronotum subquadrangular and not more narrowed anteriorly than posteriorly, with sides more or less subrectilinear), posterior angles usually without tip, and also its posterior edge evenly convex; protibia with teeth gradually increasing at apex. 2.5–7.3 mm. Figs 26–27 ............. Cillaeopeplus Sharp in Sharp & Scott, 1908

Figures

FIGURE 1. Adocimus (Adocimus) bellus, orig. Specimens from Indonesian New Guinea; A, C–E: male, typical form (“Hatam, Doria”); H–J: male, typical form (“…Suma-Uam…”); B: female, unicolorous form (“… Hatam, Beccari, 1875”); K: female, typical form (“… Madang Dist., Finistrerre Mts…”); A: body, male, dorsal (typical form); B: body, female, dorsal (unicolorous form); C-K: typical form—C: head, dorsal; D: head and thorax, ventral; E: lateral side of abdominal segments 5 and 6, dorsal; F: apex of terminal abdominal segment of male with exposed serrate apex of anal sclerite, ventral; G: apex of terminal abdominal segment of female with serrate apex of pygidium, ventral; H: tegmen, ventral; I: penis trunk, dorsal; J: proximal part of inner sac with armature; K: ovipositor, ventral. Scale bars: a = 2.0 mm to Fig. A and 2.3 mm to Fig. B; b = 1.0 mm to Figs C–E; с = 0.5 mm to Figs F–K.

FIGURE 2. Adocimus (Belonotus) bartenevi sp. nov., orig. Holotype, male from QLD (“… 11 km W of Bald Hill, Mcllwaith Range…”); A: body, dorsal; B: head and thorax, ventral; C: head, dorsal; D: drawing of anterior part of frons, dorsal; E: lateral side of abdominal segments 5 and 6, dorsal; F: male anal sclerite and spiculum gastrale, ventral; G: tegmen, ventral; H: penis trunk, dorsal; I: proximal part of inner sac with armature. Scale bars: a = 1.0 mm to Fig. A; b = 1.0 mm to figs B–C and E–F; с = 0.25 mm to Figs G–I.

FIGURE 4. Species of the genus Allenipeplus, after Kirejtshuk & Kovalev 2016.A–C: holotype of Allenipeplus philippinensis, female from Philippines (ZIN: “E Luzon, Sierra Madre, Isabela, xi.2012, I. Lumawig”); A: body of holotype, female, dorsal; B: head, ventral; C: thorax, ventral; D–F: holotype of Allenipeplus vitellinus, female from Papua New Guinea (NHML: “… Torecella M...” = Torricelli Mountains); D: body of holotype, female, dorsal; E: head, ventral; F: thorax, ventral. Scale bars: a = 2.0 mm to Figs A and D; b = 2.0 mm to Figs B and E; с = 2.0 mm to Figs C and F.

FIGURE 5. Brachypeplus apicalis, orig. A–E, G, I, K: specimens from QLD (“… Cape Tribulation, Pilgrim Sands…”); A: body, dorsal; B: head and thorax, ventral; C: male anal sclerite and spiculum gastrale, ventral; E: tegmen, ventral; G: penis trunk, dorsal; I: proximal part of inner sac with armature; K: ovipositor, ventral; D, F, H, J: specimen from Indonesian New Guinea (“Halmahera NW, 7 km N Jallolo….”); D: male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; H: penis trunk, dorsal. Scale bars: a = 1.0 mm to Fig. A; b = 1.0 mm to Fig. B; с = 0.25 mm to Figs C–K.

FIGURE 6. Brachypeplus barronensis, orig. Specimens from NSW (“North Sydney, Narrabeen”). A: body, male dorsal; B: male anal sclerite and spiculum gastrale, ventral; C: tegmen, ventral; D: penis trunk, dorsal; E: proximal part of inner sac with armature; F: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 0.25 mm to Figs B–F.

FIGURE 7. Brachypeplus basalis, orig. A–D, F, G, I—specimens from TAS (“…Mt Field NP, SE end Lake Fenton…”); A: body, male dorsal; B: shoulder part of elytron, dorsal; C: male anal sclerite and spiculum gastrale, ventral; D: tegmen, ventral; F: penis trunk, dorsal; H: proximal part of inner sac with armature; I: ovipositor, ventral; E, G: specimen from New Zealand (“…Lewis Pass Nat. Res., 13.2 km S…”); E: tegmen, ventral; G: penis trunk, dorsal. Scale bars: a = 1.0 mm to Fig. A; b = 0.5 mm to fig. B; с = 0.25 mm to Figs C–I.

FIGURE 8. Brachypeplus binotatus, orig. A-B: specimen from NSW (“… Lilyvale…”); A: body, dorsal; B: shoulder part of elytron, dorsal; C, E, G, I: specimen from NSW (“… Lansdowne S.F…”); C: male anal sclerite and spiculum gastrale, ventral; E: tegmen, ventral; G: penis trunk, dorsal; I: proximal part of inner sac with armature; D, F, H, J: specimen from QLD (“… env. Ingham …”); D: male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; H: penis trunk, dorsal; J: proximal part of inner sac with armature; L: ovipositor, ventral; specimen from ACT (“… Tidbinbilla Nat.Pk…”). Scale bars: a = 1.0 mm to Fig. A; b = 0.5 mm to Fig. B; с = 0.25 mm to Figs C–L.

FIGURE 9. Brachypeplus blandus, orig. A, D–G: male from ACT (“Black Mt., Canberra, ACT, Nov. 19’, 1981…”); A: body, dorsal; D: male anal sclerite and spiculum gastrale, ventral; E: tegmen, ventral; F: penis trunk, dorsal; G: proximal part of inner sac with armature; B-C, H: female from SA (“Port Lincoln, Blackburn”); B: body, dorsal; C: shoulder part of elytron, dorsal; H: ovipositor, ventral. Scale bars: a = 1.0 mm to Figs A and B; b = 0.5 mm to Fig. C; с = 0.25 mm to Figs D–H.

FIGURE 10. Brachypeplus dorsalis, orig. A–C, E, G: paralectotype, male from Sumatra (“… Siboga …” = Subolga); A: body, dorsal; B: male anal sclerite and spiculum gastrale, ventral; C: tegmen, ventral; E: penis trunk, dorsal; G: proximal part of inner sac with armature; D, F, H, J: specimens from Taiwan (“Lanyu Island, Taitung County, Yung Hsin Farm…”): D: tegmen, ventral; F: penis, dorsal; H: proximal part of inner sac with armature; I: ovipositor, ventral. Scale bars: a = 1.0 mm to Figs A; b = 0.25 mm to Figs B–I.

FIGURE 11. Type specimens of the genus Brachypeplus. A–C: B. apicalis; A: lectotype of B. apicalis, male from Maluku Islands (“… Mysol” = Misool), orig.; B: lectotype of B. caudalis, female from Maluku Islands (“Bac, 103” (Murray, 1864: 304: “Batchian”) = Bacan), orig.; C: paralectotype of B. fimbriatus, male from Maluku Islands (“Ternate, X.1875…”), pictured by R. Poggi; D: holotype of B. decoratus, female from Philippines (“Luzon, Benguet…”), orig.; A–D: body, dorsal. Scale bar: 1.0 mm to Figs A and B, ~1.1 mm to Fig. C, 1.4 mm to Fig. D.

FIGURE 12. Brachypeplus instriatus sp. nov., orig. A–E: male from ACT (“Canberra… apricots”); A: body, dorsal; B: head, dorsal; C: head and thorax, ventral; D: shoulder place of elytron, dorsal; E: abdominal apex, ventrolateral; F-I: male from? QLD (“in sawdust Caboolture…”); F: male anal sclerite and spiculum gastrale, ventral; G: tegmen, ventral; H: penis trunk, dorsal; I: proximal part of inner sac with armature; J: female from SA (“Port Lincoln, Blackburn”); ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 1.0 mm to Figs B, C, E; с = 0.5 mm to Fig. D; d = 0.25 mm to Figs F–J.

FIGURE 13. Brachypeplus affl. instriatus sp. nov., orig. Specimens from Australia captured in New Zealand. A: adult male, dorsal; B: same, lateral; C: anterior part of mature larva, anterolateral; D: body of mature larva, dorsal. Scale bar: 1.0 mm to Figs A–B and D; 0.25 mm to Fig. C.

FIGURE 14. Brachypeplus kemblensis, orig. A, D: male from NSW (“The Island Bellinger…”); A: body, dorsal; D: shoulder part of elytron, dorsal; B, E: paratype of B. pudicus, male from Azores; B: body, dorsal; E: shoulder part of elytron, dorsal; C, F: male from QLD (“…Mt. Glorious…”); C: body, dorsal; F: shoulder part of elytron, dorsal. Scale bars: a = 1.0 mm to Figs A–C; b = 0.5 mm to Figs D–F.

FIGURE 15. Brachypeplus kemblensis, orig. A, D, G, K: male from NSW (“The Island Bellinger…”); A: male anal sclerite and spiculum gastrale, ventral; D: tegmen, ventral; G: penis trunk, dorsal; K: proximal part of inner sac with armature; B, E, H, I, L, N: males and female from QLD (“…Mt. Glorious…”); B: male anal sclerite and spiculum gastrale, ventral; E: tegmen, ventral; H and I: penis trunk, dorsal; L: proximal part of inner sac with armature; N: ovipositor, ventral; C, F, J, M, O: paratypes of B. pudicus, male from Azores; C: male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; J: penis trunk, dorsal; M: proximal part of inner sac with armature; O: ovipositor, ventral. Scale bar: = 0.25 mm to Figs A-O.

FIGURE 16. Brachypeplus macleayi, orig. A-E: male from ACT (“…waxmoth culture”); A: body, dorsal; B: shoulder place of elytron, dorsal; C: male anal sclerite and spiculum gastrale, ventral; D: tegmen, ventral; E: penis trunk, dorsal; F: female from NSW (“… Kioloa SF, 15 km NE Batemans Bay…”); ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 0.5 mm to Fig. B; с = 0.25 mm to Figs C–F.

FIGURE 17. Brachypeplus makarovi sp. nov., orig. Holotype, male and paratype, female from QLD (“Milli Scrub…”); A: body, dorsal; B: head, dorsal; C: head, ventral; D: thorax, ventral; E: male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; G: penis trunk, dorsal; H: proximal part of inner sac with armature; I: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 0.5 mm to Figs B–D; с = 0.25 mm to Figs E–I.

FIGURE 18. Brachypeplus nypicola sp. nov., orig. Holotype, male and paratype, female from QLD (“…Claudie River …”); A: body, dorsal; B: head, dorsal; C: head, ventral; D: thorax, ventral; E: male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; G: penis trunk, dorsal; H: proximal part of inner sac with armature; I: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 0.5 mm to Figs B–D; с = 0.25 mm to Figs E–I.

FIGURE 19. Brachypeplus planus, orig. A, B, C: male from TAS (“…Mt. Field West…”); A: body, dorsal; B: shoulder part of elytron, dorsal; C: head and thorax, ventral; D-G: male from Tasmania (“…Mt. Field N. P…”); D: male anal sclerite and spiculum gastrale, ventral; E: tegmen, ventral; F: penis trunk, dorsal; G: proximal part of inner sac with armature; H: female from Tasmania (“…Lower Bardon R…”); ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 1.0 mm to Fig. B; с = 0.5 mm to Fig. C; d = 0.25 mm to Figs D–H.

FIGURE 20. Type specimens of synonyms of Brachypeplus binotatus, pictured by K. Matsumoto.A: lectotype of B. binotatus, male from VIC (Murray, 1864: 290: “Victoria”); B: body of lectotype of B. cowlei, male from QLD (Blackburn, 1902: 304: “Cairns”); C: body of lectotype of B. kobelei, male from QLD (Blackburn, 1902: 305: “North Queensland”); A–C: body, dorsal. Scale bar = ~ 1.3 mm for Figs A and C; = ~1.0 mm for Fig. B.

FIGURE 21. Type specimens of the genus Brachypeplus, A–B, E: pictured by K. Matsumoto; C-D, F: orig. A: lectotype of B. blandus, male from VIC (“Port Lincoln…”); B: body of lectotype of B. kemblensis, male from QLD (“M. Kembl.”); C–D: paratypes of B. pudicus, male and female from Açores (“… Furnas …”); E: body of lectotype of B. wattsensis, male from VIC (Blackburn, 1902: 304: “Watts River district”); paralectotype of B. wattsensis, female from the same series; A–E: body, dorsal; F: ovipositor, ventral. Scale bars = ~ 1.3 mm for Figs A, C and D; = ~1.0 mm for Fig. B; =~ 1.1 mm for Fig. E; =~ 0.25 mm for Fig. F.

FIGURE 23. Brittonoma mandibulare, comb. nov., orig. B–C, E–H: paratype, male from QLD (ANIC: “6 m. up Mt. Lewis Rd., N Q, 27.12.72, A. Walford-Huggins”, “from palm”); A, D, I: female “Mt. Lewis Road, via Julatten…”; A: body, dorsal: B: head, dorsal; C: head and thorax, ventral; D: lateral side of abdominal segments 5 and 6, dorsal; E: male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; G: penis trunk, dorsal; H: proximal part of inner sac with armature; I: ovipositor, ventral. Scale bars: a = 2.0 mm to Fig. A; b = 1.0 mm to Figs B–D; с = 0.25 mm to Figs E–I.

FIGURE 24. Brittonoma pygidiatum, comb. nov., orig. Holotype, male from QLD (ANIC: “Mt. Lewis Rd., N. Q, 3000’, 30 Oct. 1966, E. Britton”); A: body, dorsal: B: head, dorsal; C: head and thorax, ventral; D: male anal sclerite and spiculum gastrale, ventral; E: tegmen, ventral; F: penis trunk, dorsal; G: proximal part of inner sac with armature. Scale bars: a = 2.0 mm to Fig. A; b = 1.0 mm to Figs B–D; с = 0.25 mm to Figs E–I.

FIGURE 25. Caledomus extraordinarius, after Kirejtshuk & Kovalev 2017. A, C–D: holotype, male from New Caledonia (NHML: “… Tinchialit, 2,020 ft, 21.ix–3.x.1949, L.E. Cheesman, B.M. 1950-1”); B: paratype, female from the same series; A: body, dorsal; B: body, dorsal; C: head, ventral; D: thorax, ventral. Scale bars: a = 2.0 mm to Fig. A; b = 1.0 mm to Figs B–D.

FIGURE 26. Cillaeopeplus rectinotus sp. nov., orig. A–I: holotype, male from QLD (“… 20 km SW Mosmann…”) and J: paratype, female from the same series; A: body, dorsal; B: head, dorsal; C: head, ventral; D: thorax, ventral; E: lateral side of abdominal segments 5 and 6, dorsal; F: male anal sclerite and spiculum gastrale, ventral; G: tegmen, ventral; H: penis trunk, dorsal; I: proximal part of inner sac with armature; J: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 1.0 mm to Figs B–E; с = 0.25 mm to Figs F–J.

FIGURE 27. Cillaeopeplus temporalibus sp. nov., orig. A–E: holotype, female from QLD (“… 20 km SW Mosmann…”); A: body, dorsal; B: head, dorsal; C: head, ventral; D: thorax, ventral; E: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 1.0 mm to Figs B–C; с = 0.25 mm to Fig. E.

FIGURE 28. Ithyphenes australiensis sp. nov., orig. A–D, F–H: holotype, male from NSW (“3 km N of Landsowne …”); E, I: paratype from QLD (“… Cape Tribulation…”); A: body, dorsal; B: head, dorsal; C: head, ventral; D: thorax, ventral; E: terminal abdominal segment, ventral; F: male anal sclerite, ventral; G: tegmen, ventral; H: penis trunk, dorsal; I: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 0.5 mm to Figs B–E; с = 0.25 mm to Figs F–I.

FIGURE 29. Ithyphenes marinae sp. nov., orig. A–G, I–K: holotype, male from QLD (“… Iron Range…”); H, J: paratype, male from QLD (“… 9 km ENE of Mt. Tozer…”); A: body, dorsal; B: head, dorsal; C: head, ventral; D: thorax, ventral; E: terminal abdominal segment, ventral; F: male anal sclerite and spiculum gastrale, ventral; G–H: tegmen, ventral; I–J: penis trunk, dorsal; K: proximal part of inner sac with armature. Scale bars: a = 1.0 mm to Fig. A; b = 1.0 mm to Figs B–E; с = 0.25 mm to Figs F–K.

FIGURE 30. Ithyphenes rectifrons sp. nov., orig. Holotype, male from MSW (“… Umumgar S.F.…”); A: body, dorsal; B: head, dorsal; C: head and prothorax, ventral; D: terminal abdominal segment, ventral; E male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; G: penis trunk, dorsal; H: proximal part of inner sac with armature. Scale bars: a = 1.0 mm to Fig. A; b = 0.5 mm to Figs B–D; с = 0.25 mm to Figs E–H.

FIGURE 31. Laferollaeus angustissimus gen. et sp. nov., orig. A–I: holotype, male and J: paratype, female from QLD (“… Cape Tribulation area…”); A: body, dorsal; B: head, dorsal; C: head, ventral; D: thorax, ventral; E: lateral side of abdominal segments 5 and 6, dorsal; F: male anal sclerite, ventral; G: tegmen, ventral; H: penis trunk, dorsal; I: proximal part of inner sac with armature; J: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 0.5 mm to Figs B–E; с = 0.25 mm to Figs F–J.

FIGURE 32. Matthewsianus oliffi comb. nov., A–C: lectotype of Brachypeplus insignis (now Matthewsianus) from WA (after ALA 2021) (“…Cue”) and D: paralectotype of B. insignis from WA (“…Cue”), female, orig.; A: body, dorsal; B: tegmen, ventral; C: penis trunk, dorsal; D: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A and 0.5 mm to Fig. D; b = 0.5 mm to Figs B and C.

FIGURE 33. Matthewsianus polinae gen. et sp. nov., orig. A–E: holotype, female from QLD; A: body, dorsal; B: head, dorsal; C: head and thorax, ventral; D: lateral side of abdominal segments 5 and 6, dorsal; E: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 1.0 mm to Figs B–C; с = 0.25 mm to Figs E.

FIGURE 34. Onicotisauritus, adult, orig.A-H:male from NSW (“5852…”) andI: female from? (“Native bee nest, Froggatt…”); A: body, dorsal; B: head, dorsal; C: head and thorax, ventral; D: lateral side of abdominal segments 5 and 6, dorsal; thorax, ventral; E: male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; G: penis trunk, dorsal; H: proximal part of inner sac with armature; I: ovipositor, ventral. Scale bars: a = 1.0 mm to Fig. A; b = 1.0 mm to Figs B–D; с = 0.5 mm to Figs E–I.

FIGURE 36. Oniphenes bicoloratus gen. et sp. nov., orig. A–H: holotype, male and I: paratype, female from QLD (“… 3 km ENE of Mt. Tozer …”); A: body, dorsal; B: head, dorsal; C: head and thorax, ventral; D: lateral side of abdominal segments 5 and 6, dorsal; E: male anal sclerite, ventral; F: tegmen, ventral; G: penis trunk, dorsal; H: proximal part of inner sac with armature; I: ovipositor, ventral. Scale bars: a = 0.5 mm to Fig. A; b = 0.5 mm to Figs B–D; с = 0.25 mm to Figs E–I.

FIGURE 37. Oniphenes lobanovi gen. et sp. nov., orig. A–D: paratype, female from QLD (“… Cape Tribulation area, 21–28 Mar. 1984 …”), E–H: holotype, male from QLD (“Cairns”) and I: paratype, female from QLD (“… Cape Tribulation area, 1–11 May 1992 …”); A: body, dorsal; B: head, dorsal; C: head and thorax, ventral; D: lateral side of abdominal segments 5 and 6, dorsal; E: male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; G: penis trunk, dorsal; H: proximal part of inner sac with armature; I: ovipositor, ventral. Scale bars: a = 0.5 mm to Fig. A; b = 0.5 mm to Figs B–D; с = 0.25 mm to Figs E–I.

FIGURE 38. Oniphenes subunicolor gen. et sp. nov., orig. A–H: holotype, male from QLD (“… Cape Tribulation area, 1–11 May 1992 …”) and I: paratype, female from QLD (“Bellenden Ker Range …”); A: body, dorsal; B: head, dorsal; C: head and thorax, ventral; D: lateral side of abdominal segments 5 and 6, dorsal; E: male anal sclerite and spiculum gastrale, ventral; F: tegmen, ventral; G: penis trunk, dorsal; H: ovipositor, ventral. Scale bars: a = 0.5 mm to Fig. A; b = 0.5 mm to Figs B–D; с = 0.25 mm to Figs E–I.