Taraxacum goloskokovii Schischkin (1964: 748)

Kirschner, Jan & Štěpánek, Jan, 2017, A revision of Taraxacum sect. Atrata, a dandelion group centred in the Middle Asia, and the problem of Taraxacum brevirostre, Phytotaxa 305 (4), pp. 225-261 : 241

7. Taraxacum goloskokovii Schischkin (1964: 748) .

Type: — KAZAKHSTAN. Dzhungarskiy Alatau , bassein r. Koksu, lednikovye istoki r. Korzhun, po melkozemlistym mestam drevnikh moren vblizi sovremennykh [the Dzhungar Alatau, Koksu River basin, glacier sources of Korzhun River, shallow soil sites of old moraines in the vicinity of the current ones], 3300 m, 24 Aug 1948, V. Goloskokov ( LE, no. det. 6526, holotype; isotypes: LE, no. det. 17899, 17900) .

Description: —Plants 5–8 (–10) cm tall. Petiole pinkish green, narrow, long, ± unwinged; leaves mid-green, linear to linear-oblanceolate, 5–8 × (0.3–) 0.5–0.7 (–1.2) cm, glabrous, undivided, margin entire or with irregular short linear lateral teeth (1–3 on each side) or pinnatilobed; with 2 or 3 pairs of linear-triangular, to 5 mm long, usually subrecurved, entire lateral lobes; interlobes narrow, ± entire; terminal lobe linear-triangular, often elongated, base sagittate. Scapes brownish green, ± equalling leaves, glabrous. Capitulum ca. 1.5 cm wide. Involucre 6–7 mm wide, base ± rounded. Outer phyllaries 10–15, deep greyish green, not imbricate, narrowly lanceolate, outermost ones 5–6 (–7) × 1.3–2 mm and ca. 1/2 as long as inner ones, ± loosely appressed, with blackish middle line, the rest of the middle part from light greyish green to deep grey, border paler, narrow, sometimes indistinct, with blurred transition, usually ± sharply delimited and 0.1–0.3 mm wide, margin glabrous, flat to blackish callose below obtuse apex; inner phyllaries 1–1.2 cm, flat to callose below apex. Ligules yellow; outer ligules subtubular to canaliculate, outside striped ± black to pinkish black. Stigmas deep (pale brownish) yellow. Anthers polliniferous; pollen grains irregular in size. Achene light stramineous-brown with slightly greyish hue, ca. 4.7–5.4 × 0.8–0.9 mm; body ± smooth or apically with a few minute tubercles, cone not discernible; beak 4.9–5.3 mm, ± thick (0.25–0.3 mm). Pappus ± white, ca. 5.5 mm.—Agamospermous.— Fig. 9C , 10D, E , 15D .

Distribution: —The geographical range of T. goloskokovii occupies the Dzhungar Alatau and the Zailiyskiy Alatau Range ( Fig. 14 , only specimens seen have been mapped). The literature record from the W. Xinjiang ( Zhai 1999) also may be considered as quite reliable.

Specimens examined: — KAZAKHSTAN: The Dzhungar Alatau, Koksu River basin, left banks of middle stream of the Kazan River, in the vicinity of old moraines, 30 Aug 1948, V. Goloskokov (LE, no. det. 17901, 17902).—The Dzhungar Alatau, Koksu River basin, sources of Tenteksai River, 3200 m, 15 Aug 1948, V. Goloskokov (LE, no. det. 17903).—The Zailiyskiy Alatau Range (Khrebet Ileyskiy Alatau), Almaty region, catchment area of Malaya Almatinka (Kishi Almaty), 0.5–0.8 km N to NNE of the Institute of Geography (and „CSKA“) field station („Stantsia Tuyuksu“), north of the Lednik Molodyozhnyj Glacier, a part of the Tuyuksu Glacier complex, loc. 31, 43° 04‘ 28.8“ N, 77° 04‘ 26.5“ E, ca. 3370 m, 6 Aug 2009, L. Kirschnerová & J. Kirschner KAZ09-93 (PRA, no. det. 28206).— Ibidem, L. Kirschnerová & J. Kirschner 3099 (PRA, no. det. 28131).— Ibidem, L. Kirschnerová & J. Kirschner KAZ09-95 (PRA, no. det. 28209).— Ibidem, L. Kirschnerová & J. Kirschner KAZ09-105 (PRA, no. det. 28208).— Ibidem, L. Kirschnerová & J. Kirschner KAZ09-105, cultivated from roots, collected 2010 (PRA, no. det. 28207).

Less certain identification:—The Dzhungar Alatau, upper Sarkand River, upper stream of Kara-saryk River, vicinity of the pass near the left tributary of Kara-saryk, 13 Aug 1939, A. Pyataeva 284 (TASH, no. det. 15851, 15852).

Figures

FIGURE 9. A, Taraxacum corvinum from Dzhusaly-Kezen Pass; B, T. atrochlorinum; C, T. goloskokovii, both from the Tuyuksu Glacier complex.

FIGURE 10. Achenes of the members of T. sect. Atrata: A, T. alpigenum (PRA, no. det. 28132); B, T. atrochlorinum (PRA, #72); C, T. corvinum (PRA, #114/1); D, T. goloskokovii (PRA, #105/2); E, T. goloskokovii (LE, no. det. 6526); F, T. lilacinum (PRA, no. det. 28204). Scale bar = 1 mm.

FIGURE 14. The distribution range of T. goloskokovii.

FIGURE 15. Selected herbarium specimen details.A, T. dzhungaricola; B, T. pseudoatratum; C, T. subglaciale; D, T. goloskokovii; E, F, T. sinotianschanicum. All specimens are preserved at LE.