Syzeuctus vigil ( TOSQUINET , 1900)

Riedel, Matthias, 2021, The genus Syzeuctus FÖRSTER in the Western Palaearctic region and Central Asia (Hymenoptera, Ichneumonidae, Banchinae), with descriptions of 17 new species, Linzer biologische Beiträge 52 (2), pp. 1013-1098 : 1074-1076

Syzeuctus vigil ( TOSQUINET, 1900) ( Figs 50 , 93 , 139)

O r i g i n a l: Lissonota vigil TOSQUINET, 1900: 165-167

H o l o t y p e: (♀) " type; Russia Sarepta 1880 B.; collection Dr. J. Tosquinet; Lissonota vigil TOSQUINET dét. J. TOSQUINET" (Bruxelles), type studied .

P a r a t y p e: (♂) "cotype; Russia Sarepta 1880 B.; collection Dr. J. Tosquinet; Lissonota vigil TOSQUINET dét. J. Tosquinet " (Bruxelles), studied .

S t u d i e d m a t e r i a l: 4♀♀, 4♂♂; Hungary *, Russia, Ukraine *.

D e s c r i p t i o n: ♀: Body length 9-9.5 mm. Antenna with 36 flagellomeres; 1 st flagellomere 3.8-4.0× longer than wide and 0.48-0.5× as long as eye. Temple shortly parallel behind eye, strongly and roundly narrowed apically, dorsally c. 0.4-0.45× as long as eye. OED 1.3-1.7× ocellar diameter. Frons densely punctate, not swollen laterally, moderately impressed medially. Face densely punctate. Clypeus 1.7-2.0× wider than long, with scattered punctures, apical margin rounded. MI 0.5-0.6. Upper mandibular tooth slightly larger than lower one. Genal carina reaching mandibular base.

Mesosoma densely punctate, 1.4× longer than high. Frontoventral edge of pronotum moderately curved, lateral pronotal groove smooth or partly rugose. Speculum partly smooth. Epicnemial carina low, carina on pleural part distinct, transversal part with weak or without carina. Apical transversal carina of propodeum strong and complete. Hind femur 4.8-5.3× longer than wide. Hind metatarsus 0.65-0.67× as long as hind femur. 2 nd

hind tarsomere 1.3-1.4× longer than 3rd one. Hind claw completely pectinate, moderately

bent apically. Areolet stalked; 2 nd recurrent vein in or distal to its middle. Nervulus interstitial. Hairs on basal cell slightly shorter than their intervals. Hind wing with 9

distal hamuli. Metasoma stout. 1st tergite 1.1-1.3× longer than wide, with dense punctures laterally. 2nd

tergite 1.25-1.3× wider than long, with coarse dense punctures. 3 rd tergite 1.25-1.3× wider than long, with dense punctures. Ovipositor 1.0-1.05× as long as fore wing, 1.15- 1.3× longer than metasoma and 2.2-2.3× longer than hind tibia.

Color: Black. Clypeus reddish. Wide frontal orbit andnarrow vertical and outer orbits yellow. Mesosoma black or with ± reddish suffusion (f. rufus AUBERT); collare, sometimes hind edge or upper margin of pronotum, two to four longitudinal stripes on mesoscutum, tegula, sometimes subtegular ridge, usually spots on mesopleuron and metapleuron, side and apex of scutellum, often spot on postscutellum and sometimes small lateral spots on propodeum yellow. Metasoma black; 2 nd and 3 rd tergites often with dark reddish-brown suffusion and narrow yellowish apical bands OR 1 st to 3 rd tergites red (f. rufus AUBERT). Coxae and trochanters black; legs otherwise red or brown; fore and mid femora, tibiae and tarsi with yellowish frontal stripes; hind tibia yellowish basally. Fore wing with subapical brown spot; pterostigma brown, cream-yellow

proximally. ♂: Body length 6.5-9.5 mm. Antenna with 34-37 flagellomeres; 1st flagellomere 3.4-3.7×

longer than wide and 0.45-0.5× as long as eye; preapical flagellomere 1.2× longer than wide. Temple parallel behind eye and moderately narrowed apically, dorsally c. 0.55- 0.7× as long as eye. OED 1.3-1.6× and OOD 2.5× ocellar diameter. Frons with rather dense punctures and distinct median impression, ± swollen laterally. Clypeus 1.8-2.0× wider than long, with scattered punctures, apical margin rounded. MI 0.55-0.65. Upper mandibular tooth slightly larger than lower one. Genal carina reaching mandibular base.

Lateral pronotal groove smooth ventrally. Speculum usually with scattered punctures. Epicnemial carina low, low carina on pleural part present, carina on transversal part absent or weak. Apical transversal carina of propodeum complete. Hind femur 4.3-5.3× longer than wide. Hind metatarsus 0.65-0.75× as long as hind femur. 2 nd hind tarsomere 1.2-1.25× longer than 3 rd one. Hind claw with short teeth, moderately curved apically. Areolet stalked; 2 nd recurrent vein in or distal to its middle. Nervulus interstitial or slightly postfurcal by 1× its width. Hind wing with 8 distal hamuli.

Metasoma stout; 1 st tergite 1.2-1.4× longer than wide, with rather dense or dense coarse punctures laterally. 2 nd tergite 1.05-1.3× wider than long, with coarse dense punctures. 3 rd tergite 1.0-1.3× wider than long, with dense but sometimes superficial punctures. Spiracles of 1 st to 3 rd tergites ± protruding

Color: Black. Clypeus reddish-yellow in apical half. Base of mandible, entire orbits, sometimes malar space and sometimes transverse stripe on occiput yellow. Mesosoma black; collare and upper margin of pronotum, two paramedian stripes on mesoscutum, tegula, two spots on mesopleuron and spot on metapleuron, side and apex of scutellum and postscutellum yellow). Metasoma black, sometimes 2 nd and 3 rd tergites often dark reddish-brown; in paratype 5 th to 7 th tergites with yellow apical bands, but sometimes all tergites with yellow apical bands; 1 st tergite often with yellow basal spots. Coxae and trochanters black, usually with yellow stripes; fore and mid legs reddish or yellow, femora sometimes with caudal brown stripes; hind femur reddish or brown, sometimes yellowish dorsally, with apical yellow spot; hind tibia yellowish, often brownish internally and/or apically; hind tarsomeres yellow, with brownish tips. Fore wing (without or) with subapical brown spot; pterostigma brown, cream-yellow proximally.

Hosts unknown.

Figures

Figs 49-52: Basal tergites of ♀♀ from dorsal: (49) Syzeuctus tigris SEYRIG, (50) S. vigil (TOSQUINET) HT, (51) S. ibericus nov.sp. HT, (52) S. kazakhstanicus nov.sp. PT. Figs 53-55: Basal tergites of ♂♂: (53) Syzeuctus flavator nov.sp. HT, (54) S. flagellator nov.sp., (55) S. dusmeti SEYRIG with enlarged tergal spiracles, from lateral. Fig. 56: Hind tarsomere of Syzeuctus crassitarsis TELENGA ♀.

Figs 93-95: Faces of ♀♀ from frontal: (93) Syzeuctus vigil (TOSQUINET) HT, (94) S. ibericus nov.sp. HT, (95) S. kazakhstanicus nov.sp. PT. Figs 96-97: Faces of ♂♂ from frontal: (96) Syzeuctus flavator nov.sp. HT, (97) S. flagellator nov.sp. PT. Figs 98-99: Propodeum (98) Syzeuctus kazakhstanicus nov.sp. ♂, (99) S. petiolaris (GRAVENHORST) ♂.