Spinaxytes palmata Srisonchai, Enghoff & Panha

Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai & Panha, Somsak, 2018, A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae), ZooKeys 797, pp. 19-69 : 19

publication ID

https://dx.doi.org/10.3897/zookeys.797.29510

publication LSID

lsid:zoobank.org:pub:A5403588-F43D-40FF-81EE-2009E49823B4

persistent identifier

https://treatment.plazi.org/id/2A4B8447-3443-44D7-B463-0A6A2AB840C9

taxon LSID

lsid:zoobank.org:act:2A4B8447-3443-44D7-B463-0A6A2AB840C9

treatment provided by

ZooKeys by Pensoft

scientific name

Spinaxytes palmata Srisonchai, Enghoff & Panha
status

sp. n.

Spinaxytes palmata Srisonchai, Enghoff & Panha View in CoL sp. n. Figs 2; 4N, O; 5F; 21, 22, 23

Material examined.

Holotype. ♂, THAILAND, Surat Thani Province, Phanom District, Wat Tham Wararam, 8°53'07"N, 98°40'01"E, ca. 51 m a.s.l., 6 Aug. 2016, ASRU members leg. (CUMZ-pxDGT00215). Paratypes. 23 ♂♂, 22 ♀♀, same data as for holotype (CUMZ-pxDGT00216); 1 ♂, 1 ♀, same data as for holotype (ZMUC00040254); 1 ♂, 1 ♀, same data as for holotype (ZMUM); 1 ♂, 1 ♀, same data as for holotype (NHMW9425); 1 ♂, 1 ♀, same data as for holotype (NHMUK). Further specimens, not paratypes. THAILAND: 1 ♂, 2 ♀♀, Surat Thani Province, Phanom District, Wat Tham Wararam, 8°53'07"N, 98°40'01"E, ca. 51 m a.s.l., 5 Aug. 2014, ASRU members leg. (CUMZ).

Etymology.

The species name is a Latin adjective, referring to the tip of lamina medialis which is somewhat hand-shaped.

Diagnosis.

Male femora without modification. Similar in this respect to S. efefi sp. n., S. hasta sp. n., S. sutchariti sp. n. and S. tortioverpa sp. n., but differs from them by having: anterior part of sternal lobe between male coxae 4 bifurcate, fork-like; tip of lamina medialis expanded, hand-shaped.

Description.

SIZE. Length 26-30 mm (male), 27-32 mm (female); width of midbody metazona 1.9-2.2 mm (male), 2.0-2.4 mm (female). Width of collum = 2 = 3 = 4 <head <5-16, thereafter body gradually tapering towards telson.

Colour (Figure 21 A–C). Specimens in life with body black; paraterga orange; head, antennae (except whitish distal part of antennomeres 7 and 8), collum, prozona, metaterga (except white spines in posterior row) and surface below paraterga black; sterna and legs brown; epiproct pale brown; a few basal podomeres whitish brown.

Antennae (Figure 22M). Reaching to body ring 8 (male) and 6 or 7 (female) when stretched dorsally.

Collum (Figure 22A). With three transverse rows of setiferous tubercles/cones, 4+4 in anterior row, 1(0)+1(0) in intermediate row and 2(1)+2(1) in posterior row; with one conspicuous setiferous notch at lateral margin; paraterga spiniform, long, tip sharp, elevated at ca. 15°-20° (male) 10°-15° (female), directed almost laterad.

Tegument. Very shining; collum coarsely microgranulate; metaterga and surface below paraterga smooth.

Metaterga (Figure 22A, C, E). With two transverse rows of setiferous cones and setiferous spines; metaterga 2-19 with 2+2 cones in anterior row and 2+2 spines in posterior row; lateral cones/spines of posterior row bigger and longer than mesal ones, gradually reduced in size and length on the following rings.

Paraterga (Figure 22 A–E, H). Very long; directed dorsolaterad on body rings 2-17, elevated at ca. 50°-60° (male) 45°-60° (female), directed increasingly caudad on body rings 18 and 19. Ozopore visible in dorsolateral view.

Telson (Figure 22E, H, I). Epiproct quite short; tip subtruncate; lateral setiferous tubercles conspicuous; apical tubercles inconspicuous. Hypoproct subtrapeziform (in some specimens subtriangular); caudal margin round (in some specimens angular), with inconspicuous setiferous tubercles.

Sterna (Figs 4N; 22F, G). Sternal lobe between male coxae 4 with two parts; anterior part bifurcate, tuning-fork-like, long, tips sharp, in situ directed ventroanteriad; posterior margin of anterior part bearing 1 pore; posterior part swollen, short.

Legs (Figs 4O, 22J). Male femora without modification.

Gonopods (Figs 5F, 23). Coxa shorter than femur. Prefemoral part ca. half as long as femur. Femur not enlarged distally. Postfemoral part broad. Mesal sulcus and lateral sulcus wide. Solenophore a bit bigger than postfemoral part: lamina lateralis small, oval, tip round: lamina medialis long; basally enlarged and slightly attenuated near the tip; tip fringed, hand-shaped; tip curving down, in situ resting close to solenomere. Solenomere curved and twisted, compressed in transverse section, tip directed posteriad.

Distribution and habitat

(Figure 21D). S. palmata sp. n. is known only from the type locality. We regard this species as endemic for the Thai fauna. The new species can be found in syntopy with Desmoxytes corythosaurus Srisonchai, Enghoff & Panha, 2018, crawling on humid rock walls.

Remarks.

There are variations in the hypoproct: subtrapeziform in some specimens, subtriangular in the others; caudal margin in some individuals round, angular in the others.

Kingdom

Animalia

Phylum

Arthropoda

Class

Diplopoda

Order

Polydesmida

Family

Paradoxosomatidae

SubFamily

Paradoxosomatinae

Tribe

Orthomorphini

Genus

Spinaxytes