Sinoennea fartoidea ( Theobald, 1870 ) Páll-Gergely & Hunyadi & Grego & Sajan & Tripathy & Chen, 2020

Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Sajan, Sheikh, Tripathy, Basudev & Chen, Zhe-Yu, 2020, A review of the Diapheridae (Gastropoda: Eupulmonata: Streptaxoidea), with special emphasis on India and Myanmar, Raffles Bulletin of Zoology 68, pp. 682-718 : 704-706

Sinoennea fartoidea ( Theobald, 1870) , new combination

( Figs. 1K, L , 15 , 22P )

Pupa fartoidea Theobald, 1870: 400 .

Ennea (Huttonella) fartoides (sic) — Nevill, 1878: 7.

Pupa (Ennea) fartoides (sic) — Hanley & Theobald, 1876: 40, pl. 100, fig. 5.

Ennea (Indoennea) fartoidea — Kobelt, 1904b: 278, pl. 20, fig. 21.

Ennea fartoidea — W.T. Blanford & Godwin-Austen, 1908: 18.

Gulella fartoidea — Richardson, 1988: 81.

Type locality. “ Shan States ” .

Material examined. 1 shell, probable syntype ( NZSI M.33964/9), Shan States, coll. W. Theobald; 2 shells in separate vials (probably syntypes) ( NHMUK 1888.12.04.491– 2), Upper Salwin valley .

Remarks. Shell shape elongate ovoid with cylindrical sides and domed dorsal side. Upper whorls and last quarter whorl regularly ribbed, remaining ca. 2 last whorls with weak growth lines. Aperture subrectangular. Parietal callus distinct, not “smeared” onto preceding whorl, does not reach the middle of the preceding whorl. Apertural dentition 3-fold. Parietal lamella strong, elevated, straight, its inner end is not visible in oblique view to the aperture. Sinulus teardrop-shaped, opens slightly laterally, relatively small, and not conspicuously isolated due to the relatively weak palatal tooth. Palatal wall with two denticles. Upper one strong but blunt, situated on the peristome edge, lower one lower in position and deeper, very weak, only slightly indicated. No structure that would be homologous with the columellar lamella of other Sinoennea species is visible in oblique view to the aperture. The slight thickening on the columellar edge is homologous with the inner columellar edge of other Sinoennea species , and probably not homologous with the columellar lamella of other Sinoennea species (see Fig. 1B, K ). Peristome strongly thickened and expanded, not reflected, relatively weak around the sinulus. Basal swelling very slight, situated very close to peristome. There is a slight depression just behind the peristome on the outer side (corresponds with the upper palatal tooth). Inner (umbilical) side very short, without depression. Umbilical development not visible. Periumbilical keel moderate, but relatively clearly visible from umbilical view, especially due to the slight furrow on the outer side.

Most shell characters of the general shell shape and the apertural barriers agree with other Sinoennea species , therefore this species is moved to that genus. The absence of the columellar lamella ( Fig. 1K, L ) is surprising, and was not found in any other Sinoennea species.

Figures

Fig. 1. Nomenclature of the apertural barriers. A–H, Sinoennea; I, J, Pupennea, new genus; K, L, Sinoennea fartoidea; M, N, Platylennea, new genus; O, Rowsonia, new genus; P, Gulella (Huttonella). Abbreviations: BK, basal keel; BS, basal swelling; BT, basal tooth; CD, columellar denticle; CL, columellar lamella; CTd, depression corresponding with the columellar lamella; IBD, inner basal denticle; ICE, inner columellar edge; LBD, lower basal denticle; LPF, lower palatal fold; LPT, lower palatal tooth; LPTd, depression corresponding with the lower palatal tooth; MPF, middle palatal fold; OBD, outer basal denticle; PL, parietal lamella; PT, palatal tooth; S, sinulus; UBD, upper basal denticle; UPF, upper palatal fold; UPT, upper palatal tooth.

Fig. 15. Syntype of Sinoennea fartoidea (Theobald, 1870) (NZSI M.33964/9).

Fig. 22. Apertures of Streptaxidae (A) and Diapheridae (B–P). A, Gulella (Huttonella) bicolor (Hutton, 1834); B, Rowsonia beddomei (W.T. Blanford, 1881); C, Platylennea subcostulata (W.T. Blanford, 1881); D, Pupennea planguncula (Benson, 1863); E, Diaphera polita, Páll-Gergely new species; F, Diaphera cylindrelloidea (Stoliczka, 1871); G, Bruggennea cf. bongi (Dance, 1970), H, Parasinoennea ovulum (Bavay & Dautzenberg, 1912); I, Sinoennea otostoma Páll-Gergely, A. Reischütz & Maassen in Páll-Gergely, Reischütz, Maassen, Grego & Hunyadi, 2020; J, Sinoennea infantilis Páll-Gergely & Grego in Páll-Gergely et al., 2020; K, Sinoennea variabilis Páll-Gergely & Grego in Páll-Gergely et al., 2020; L, Sinoennea sutchariti Páll-Gergely & Hunyadi in Páll-Gergely et al., 2020; M, Sinoennea montawana Páll-Gergely & Hunyadi, new species; N, Sinoennea angustistoma Páll-Gergely, A. Reischütz & Maassen in Páll-Gergely et al., 2020, O, Sinoennea woodthorpei (Peile, 1929); P, Sinoennea fartoidea (Theobald, 1870). Not to scale.