Sigmaboilus peregrinus, Gu, Junjie, Zhao, Yun-Yun & Ren, Dong, 2009

Gu, Junjie, Zhao, Yun-Yun & Ren, Dong, 2009, New fossil Prophalangopsidae (Orthoptera, Hagloidea) from the Middle Jurassic of Inner Mongolia, China, Zootaxa 2004, pp. 16-24 : 19-20

Sigmaboilus peregrinus sp. nov.

( Figs. 3–8 )

Diagnosis: Forewing: ScP long and ending on anterior margin at about the third fifth of the wing; CuA+CuP1a ramified with 4 to 6 terminal branches, first anterior branch of CuA+CuP1a simple, second anterior branch dichotomously ramified; and coloration formed irregular bands.

Description: Forewings: 37.9mm/ 40mm preserved length (respectively left forewing/right forewing, holotype). Long and narrow, base of tegmen constricted. Precostal area between ScA and anterior margin very wide, with ‘radial veinlets’ and regular cross-veins at its base. ScA apparently long, mildly undulate; ScP long and ending on anterior margin at about the third fifth of the wing, branches of ScP numerous and most of them end in stem ScA. Area between ScP and R narrow and with regular cross-veins. R forking into RA and RP at about 1/3 basal tegminal length; RA pectinate with 6-8 terminal branches and RP with 6-7 terminal branches; RP branched beyond first branch of RA, but beyond second branch RA in left forewing of holotype. Area between R and M with oblique and more or less parallel cross-veins before mid-length, dense and network of regular cross-veins covering distal half; M forked before the origin of RP, MA oriented towards RP at its origin, beyond gently arched and oriented towards posterior wing margin; area between MA and MP with dense reticulated cross-veins in distal part, cross-veins at the base of area between CuA+M and CuP sigmoidal. CuP1a twice the length of CuA before their fusion. CuA and CuP1a remaining united as CuA+CuP1a for a short and appreciable distance, first anterior branch of CuA+CuP1a reaching posterior margin distal of the end of ScP; Second anterior branch dichotomously ramified with 2 to 4 branches. The ‘handle’ subparallel to each cross-vein between CuP1b and posterior branch of CuA+CuP1a. CuP1b curved, CuP1b between its base and the ‘handle’ straight, the ‘handle’ subparallel to each crossvein between the ‘handle’ and CuP1a; CuP2 very strong, sharply curved, clearly Z-shaped, cross-veins between CuP2 and CuP1b gently curved but not strong sigmoidal. 1A strong and similar to CuP 2 in shape, convergent to CuP2 at about their midlength; 2A apparently S-shaped. The coloration spread over the whole tegmen and formed irregular bands.

Material: Holotype, male tegmen, CNU-ORT-NN2008018; paratypes, 8, male tegmen: CNU-ORT- NN2008016/19/38/40/41PC/44PC/45/57; 13, female tegmen: CNU-ORT-NN2008017/34/43/47/48PC/51/53- 55/58-61.

Etymology: The specific epithet derives from Latin ‘ peregrinus ’, for its special ‘handle’ vein.

Horizon and locality: Jiulongshan Formation, Middle Jurassic, Daohugou Village, Ningcheng County, Inner Mongolia, China.

Remarks: This species differs from the other species in the following features on the forewing: coloration formed irregular bands and spread over whole tegmen; first anterior branch of CuA+CuP1a simple and second anterior branch dichotomously ramified; and cross-veins between CuP2 and CuP1b gently curved but not sigmoidal.

Figures

FIGURE 3. Sigmaboilus peregrinus sp. nov., A − B. 3, photograph of holotype, A. part: CNU-ORT-NN 2008018 P; B. counterpart: CNU-ORT-NN 2008018 C. C − D. photograph of paratype, C. Ƥ, CNU-ORT-NN 2008034. D. 3, CNU-ORT- NN 2008041 P.

FIGURES 4 – 7. Sigmaboilus peregrinus sp. nov., 4 – 5. line drawing of holotype, 3, CNU-ORT-NN 2008018: 4. left forewing. 5. right forewing. 6 − 7. line drawing of paratype, 6. Ƥ, CNU-ORT-NN 2008034. 7. 3, CNU-ORT-NN 2008041 P.

FIGURE 8. Sigmaboilus peregrinus sp. nov., additional material: 3, A. CNU-ORT-NN 2008016. B. CNU-ORT- NN 2008019. C. CNU-ORT-NN 2008040. D. CNU-ORT-NN 2008045. G. CNU-ORT-NN 2008044 P. Ƥ, E. CNU-ORT- NN 2008043. F. CNU-ORT-NN 2008048 P. H. CNU-ORT-NN 2008061. All scale bars = 5 mm.