Selenops nesophilus Chamberlin, 1924

Crews, Sarah C., 2011, A revision of the spider genus Selenops Latreille, 1819 (Arachnida, Araneae, Selenopidae) in North America, Central America and the Caribbean, ZooKeys 105, pp. 1-182 : 52-54

Selenops nesophilus Chamberlin, 1924 Figs 61 –64198– 199Map 7

Selenops nesophilus Chamberlin 1924: 656, Figs 97-98 (♂, ♀, examined).

Selenops nesophilus Muma 1953: 15, Figs 23-26 (♂, ♀).

Type material.

Holotype male: Isla Tortuga, Baja California Sur, México, 11.V.1921, J.C. Chamberlin (CAS, examined). Paratypes: Female, same data as holotype (CAS, examined).

Other material examined.

MÉXICO: Baja California: 2 miles south El Rosario on Mex Hwy 1 (dirt road) RB-3, 26.XI.1968, P.R. Craig, D. Dailey, 1♂ (CAS); Isla Angel de la Guarda, Palm Cañon, under bark of Palo Verde, 3.V.1921, J.C. Chamberlin, 1♀ (CAS); Isla Raza, 21.IV.1921, J.C. Chamberlin, 1♀, 2 imm. (CAS); 11 mi SW Punta Prieta, 200' 15.IV.1969, S.C. Williams, 1♀ (CAS); Isla Cedros, 2nd Canyon, South of Punta Norte on east side of island, 2.VII.1983, V.F. Lee, 1♂ (CAS). Baja California Sur: between San Juanico and San Ignacio, ~2km East of Ballena, 26°24'00.4"N, 112°35'64.0"W, ~190', 20.III.2004, M.C. Hedin, under rocks, 1♀, 1♂ (EME sel_210, 213); Cuevas Pintas, 25.58.644 N, 111.27.924 W, 31.III.2003, D. Palmer, D. Wood 1 imm. (EME sel_009); Isla Magdalena, Laguna Santa Maria, 9.VII.1983, D.K. Faulkner, D. C. Lightfoot, 1♀ (CAS); West of San Miguel Comondu, road to Villa Insurgentes, ~3 km West of town, 26.01'47.3"N, 111.50'84.8"W, ~590', 18-19.III.2004, M.C. Hedin, under rocks along road, 1p♂ (EME sel_212); Puerto Escondido, 14.VI.1921, E.P. Van Duzee, 1♂ (CAS); 6.8 mi S of San Antonio, 24.VII.1980, Williams, Bentzion, 1♀ (CAS). Sinaloa: 6 miles south of Culiacan, 22.VII.1954, W.J. Gertsch, 1♀ (AMNH). Sonora: Los Algodones, 6 miles west San Carlos Bay, 22-23.III.1980, C.E. Griswold, 1♂ (CAS). UNITED STATES: Arizona: Cochise Co., 2.5 miles southeast of Portal, 26.III.1965, B., C. Durden, several (AMNH); Coconino Co., Grand Canyon National Park, Tapeats Creek Campground, Tapeats Canyon, Tapeats Creek Trail 36°22.826'N, 112°27.744'W, ~2230', 15.VIII. 2004, S. Crews, under rocks, 1 imm. (EME sel_211); Pima Co., Espero Canyon, 1♀ (AMNH); Pima Co., Baboquivari Mountains, Brown Canyon, 9.VI.1952, M. Cazier, W.J. Gertsch, R. Schrammel, several (AMNH); Pima Co., Esperero Co., several (AMNH); Pinal Co., Superior Thompson Southwestern Arboretum, 11.V.1940, 2300', D.C. Lowrie, 1♂ (AMNH). California: San Diego Co., Upper Otay River Valley, 19.V.2003, D. Palmer, 1♀ (EME sel_002); San Diego Co., Jamul, Lyon's Valley, N of Lyon's Pk, 32.71415°N, 116.7677°W, ~674 m, D. Palmer, running on ground, oak woodland, 1♂ (EME sel_837); San Diego Co., Anza Borrego Desert State Park, Carrizo Palm Grove, below railroad tracks, 32.46°N, 116.11°W, ~1800', 23.II.2003, M.C. Hedin, D. Palmer, D. Wood, 2 imm. (EME sel_021, 214).

Diagnosis.

Males can be differentiated from all other species by the embolus, which originates at 6 o'clock, a long process on the MA, and the RTA, which is on a very long stalk and has a very large quadrangular process distally (Figs 61-62). Females can be separated from other species by the posterior margin of the epigynum which is sinuous and heavily sclerotized, with a median septum that is wider anteriorly, and a posterodorsal fold that is sinuous (Figs 63-64).

Description.

Holotype male: Color:carapace orange-brown (recent) pale yellow; sternum (holotype) red-brown (recent) yellow; chelicerae (holotype) brown (recent) yellowish, like carapace, darker toward inner margins; maxillae (holotype) yellow-brown, lightening distally (recent) light yellow, lightening distally; labium (holotype) orange-brown (recent) light yellow-orange; abdomen dorsally (holotype) dark grey-brown (recent) pale yellow to cream with some darker grey flecks, festoon present; ventrally (holotype) grey (recent) light yellow; legs (holotype) orange-brown, no markings evident (recent) pale yellow with darker grey annulations. Cephalothorax:0.91 times longer than broad. Eyes:AER slightly recurved; PER recurved; PME larger than AME, PLE largest, ALE smallest; eye diameters, AME 0.18, ALE 0.10, PME 0.35, PLE 0.55; interdistances AME-PME 0.03, PME-ALE 0.18, ALE-PLE 0.75. PME-PME 1.18. ALE-ALE 2.00; ocular quadrangle AME-AME 0.45, PLE-PLE 2.38; clypeus 0.23 high. Mouthparts:chelicerae with a few stout setae medially and anteriorly; maxillae longer than broad, with tuft of conspicuous setae distally; labium distally rounded. Sternum:1.25 times longer than broad, posteriorly indented. Legs:leg I only slightly shorter than legs II, III and IV; leg formula 2341; scopulae present on all 4 tarsi, as well as tibia and metatarsus of leg I and distally on metatarsi of legs II, III, IV; tarsi I-IV with strong claw tufts; pr claw toothed, rl claw with fewer teeth; spination: leg I, Fm pr 1 –1– 1, d 1 –1– 1, rl 1 –1– 1; Ti pr 1 –0– 1, d 1 –2– 0, rl 1 –0– 1, v 2 –2– 2; Mt pr 1 –1– 0, rl 1 –1– 0, v 2-2; leg II, Fm pr 1 –1– 1, d 1 –1– 1, rl 1 –1– 1; Ti pr 1 –0– 1, rl 1 –0– 1, d 1 –2– 0, v 2 –2–2– 1; Mt pr 1 –1– 0, v 2-2, rl 1 –1– 0; leg III, Fm pr 1 –1– 1, d 1 –1– 1, rl 1 –1– 1; Ti pr 1 –0– 1, d 1 –0– 0, rl 1 –0– 0, v 2 –2–1– 1; Mt pr 1 –1– 0, rl 1 –1– 0, v 2-2; leg IV, Fm pr 1 –1– 1, d 1 –1– 1, rl 1 –1– 1; Ti pr 1 –0– 1, rl 1 –1– 1, d 1 –1– 0, v 2 –2– 2; Mt pr 1 –1– 0, rl 1 –1– 0, v 2-2. Abdomen:without terminal setal tufts. Pedipalp:Fm, spination dorsal 0 –1– 4; cymbium oval in ventral view, slightly angled posterolaterally; conductor large, angular, arising laterally, curved on one side, sinuate on other, tapering distally; embolus long, slender, curved, beginning at 6 o'clock, terminating at 12 o'clock; MA arising at 2 o'clock, directed laterally, with short stout base and long finger-like process; two tibial apophyses, ventral process smaller of the two, blunt and curved, lateral process a stout stalk curving laterally away from cymbium, with large angular blunt process, similar to type of Selenops actophilus , but stalk longer and more curved (Figs 61-62). Dimensions: Total length 8.75. Carapace length 5.13, width 5.65. Ste rnum length 2.50, width 2.00. Abdomen length 3.53, width 2.78. Pedipalp: Fm 1.75, Pt 0.65, Ti 0.75, Ta 1.75, total 4.90. Leg I: Fm 6.75, Pt 2.25, Ti 5.50, Mt 5.25, Ta 2.75, total 22.50. Leg II: Fm 6.00, Pt 2.75, Ti 6.25, Mt 5.60, Ta 2.60, total 23.20. Leg III: Fm 6.75, Pt 2.50, Ti 5.80, Mt 5.75, Ta 2.20, total 23.00. Leg IV: Fm 6.00, Pt 2.00, Ti 6.00, Mt 6.00, Ta 2.75, total 22.75.

Paratype female: Color:carapace (paratype) red-brown (recent) yellow-tan, brown line extending from center of cephalic area medially thorough entire cephalothorax, dark spots laterally; sternum (paratype) brown, darker around border (recent) yellow, darker around border; chelicerae (paratype) red-brown (recent) light brown medially, darker laterally; maxillae (paratype) brown (recent) yellow-orange, both lightening distally; labium (paratype) brown (recent) orange-brown, lightening distally; abdomen dorsally (paratype) grey-brown (recent) grey-brown, lighter medially, dark festoon present; ventrally cream colored; legs (type) orange-brown, annulations barely visible (recent) pale yellow-brown with dark annulations. Carapace:0.92 times longer than broad. Eyes:AER slightly recurved; PER recurved; PME larger than AME, PLE largest, ALE smallest; eye diameters, AME 0.15, ALE 0.08, PME 0.45, PLE 0.65; interdistances AME-PME 0.03, PME-ALE 0.15, ALE-PLE 0.70. PME-PME 1.35. ALE-ALE 2.35; ocular quadrangle AME-AME 0.53, PLE-PLE 2.68; clypeus 0.15 high. Mouthparts:chelicerae with a few stout setae medially and anteriorly; maxillae longer than broad, with tuft of conspicuous setae distally. Sternum:1.20 times longer than broad, posteriorly indented. Legs:leg formula 4321; scopulae present on all 4 tarsi and metatarsi and distally on tibiae on legs I and II; tarsi I-IV with strong claw tufts; pr claw per foot slightly toothed; spination: leg I, Fm pr 1 –1– 0, d 1 –1– 1, rl 1 –1– 1; Ti v 2 –2– 2; Mt v 2-2; II, Fm pr 1 –0– 0, d 1 –1– 1, rl 1 –1– 1; Ti v 2 –2– 2; Mt v 2-2; III, Fm pr 1 –1– 0, d 1 –1– 1, rl 1 –1– 1; Ti v 2 –2– 0; Mt v 2-2. Abdomen:without terminal setal tufts. Pedipalp:claw with 9 teeth. Epigyne:median septum present, wider anteriorly, rounded to pointed terminally, posterior margin of plate sinuous, with lateral protrusions, genital openings located at lateral margins of septum; internally, cylindrical ducts directed anteriorly and anterolaterally, fertilization ducts located laterally, directed anterolaterally, posterodorsal fold present, not connected medially, heavily sclerotized, not covering internal ducts (Figs 63-64). Dimensions: Total length 11.50. Carapace length 5.68, width 6.20. Sternum length 3.00, width 2.50. Abdomen length 5.83, width 3.73. Pedipalp: Fm 1.75, Pt 0.75, Ti 1.00, Ta 2.00, total 5.50. Leg I: Fm 6.00, Pt 2.50, Ti 5.00, Mt 4.00, Ta 1.75, total 13.25. Leg II: Missing. Leg III: Fm 6.50, Pt 2.50, Ti 5.50, Mt 5.00, Ta 2.00, total 21.50. Leg IV: Missing.

Natural History.

This species has been found under rocks (Figs 198-199) as well as running on the ground in oak woodlands.

Distribution.

Known from southwestern North America, from Baja California Sur and Sinaloa to Arizona (Map 7).