Acanthoderes (Scythropopsis) pupillata Bates, 1880

Santos-Silva, Antonio & Nascimento, Francisco E. De L., 2018, Synonymies, transferences, redescriptions and new species in Lamiinae (Coleoptera, Cerambycidae), Zootaxa 4462 (2), pp. 192-210 : 201-202

Acanthoderes (Scythropopsis) pupillata Bates, 1880 , comb. nov.

( Figs 20–24 )

Acanthoderes pupillatus Bates, 1880: 60 .

Acanthoderes pupillata ; Lameere, 1883: 62 (cat.); Blackwelder, 1946: 611 (checklist).

Acanthoderes (Psapharochrus) pupillata ; Aurivillius, 1923: 387 (cat.); Gilmour, 1965: 614 (cat.); Monné & Giesbert, 1994: 231 (checklist); Monné, 1994b: 67 (cat.); Martínez, 2000: 94 (distr.).

Psapharochrus pupillatus ; Monné, 2005: 211 (cat.); 2018: 293 (cat.); Bezark, 2018: 242 (checklist).

Description. Female ( Figs 20–23 ). Integument mostly black; antennomeres III–VIII with dark reddish-brown ring anteriorly, gradually smaller toward VIII; femora dark brown, almost black; tibiae dark reddish-brown, almost brown, on anterior 2/3; tarsomere III gradually reddish brown toward apex; tarsomere V yellowish-brown except black claws.

Head. Frons densely micropunctate; with greenish-brown pubescence obscuring integument (somewhat yellowish-green depending on angle of light source); with a few long, erect dark setae laterally. Vertex with slightly elevated gibbosity between antennal tubercles and upper eye lobes; densely micropunctate; with greenish-brown pubescence obscuring integument (somewhat yellowish-green depending on angle of light source) except glabrous tip of gibbosity. Area behind upper eye lobes with dense greenish-brown pubescence greenish-brown obscuring integument. Area behind connection of eye lobes densely micropunctate toward connection (this area forming moderately wide, glabrous reentrance toward tumid area behind lower eye lobe), finely, abundantly punctate toward prothorax; with narrow greenish-brown pubescent band close to connection, glabrous on remaining surface. Area behind lower eye lobes tumid, densely micropunctate on large area close to eye, finely, moderately abundant punctate close to prothorax, gradually finer, sparser toward ventral side; with greenish-brown pubescence on tumid area, except on glabrous reentrance, glabrous on remaining surface. Genae densely micropunctate; with greenishbrown pubescence partially obscuring integument, except on glabrous apex. Antennal tubercles with sculpturing and pubescence as on frons, except smooth, glabrous apex. Median groove distinct from clypeus to prothoracic margin. Postclypeus widely emarginate centrally; with greenish-brown pubescence not obscuring integument on wide central area close to frons, glabrous laterally; with dense, bristly yellowish-brown pubescence close to anteclypeus; with long, erect dark setae on wide central area. Labrum coplanar with anteclypeus at posterior 2/3, inclined at anterior third; posterior 2/3 with greenish-brown pubescence partially obscuring integument, and erect setae close to inclined area (setae darkened basally, yellowish toward apex); anterior third nearly glabrous, with fringe of golden setae at anterior margin. Gulamentum with greenish-brown pubescence not obscuring integument on anterior third. Lower eye lobes 0.5 times genal length; connection between eye lobes formed by narrow carina, without ommatidia; distance between upper eye lobes 0.72 times length of scape; in frontal view, distance between lower eye lobes slightly larger than length of scape. Antennae 1.2 times elytral length, nearly reaching posterior fifth of elytra. Scape widened on posterior half (slightly narrowed from middle of posterior half); with yellowishbrown pubescence partially obscuring integument; with long, erect yellowish setae ventrally, more abundant on basal third. Pedicel with yellowish-brown pubescence partially obscuring integument; with long, erect yellowish setae ventrally. Antennomeres III–IV with pale-yellow pubescence on anterior half, gradually sparser, slightly conspicuous toward apex; apex with short, thick, dark setae on ventral and inner side; with long, erect, sparse yellowish setae ventrally on III, nearly absent on IV. Antennomeres V–VIII with sparse pale-yellow pubescence on anterior third, sparser, slightly conspicuous toward apex; remaining antennomeres with minute, slightly conspicuous brownish pubescence. Antennal formula based on antennomere III (only one female measured): scape = 0.85; pedicel = 0.21; IV = 0.73; V = 0.46; VI = 0.40; VII = 0.36; VIII = 0.28; IX = 0.22; X = 0.20; XI = 0.20.

Thorax. Prothorax 2.0 times wider than long (including lateral tubercles); lateral tubercles placed at middle, conical, large, with blunt apex. Pronotum whit large conical tubercle with blunt apex on each side, from about anterior fifth to distal quarter (apex directed outward); with wide longitudinal gibbosity centrally, from near anterior area to near posterior area, narrowed along middle length; with row of coarse punctures posteriorly on each side of central gibbosity, with coarse, sparse punctures on remaining surface, except on tubercles and central gibbosity; with yellowish-white (more whitish depending on angle of light source) pubescence partially obscuring integument between lateral tubercles and laterally (including on lateral tubercles of prothorax), greenish-brown on narrow anterior and posterior areas and between areas with yellowish-white pubescence, and two spots on each side with dark-brown pubescence (one on anterior surface of tubercle, another between posterior margin and tubercle); tips of tubercles and central gibbosity glabrous. Sides of prothorax with yellowish-white pubescence on wide central area, partially obscuring integument; anterior and posterior areas with greenish-brown pubescence. Prosternum with greenish-yellow pubescence not obscuring integument, slightly denser laterally. Posterior area of prosternal process slightly wider than procoxal cavity, forming distinct tubercle on each side of posterior curvature. Ventral side of meso- and metathorax with greenish-yellow pubescence partially obscuring integument, except glabrous anterocentral area of mesoventrite. Mesoventral process about 1.3 times as wide as mesocoxal cavity; with distinct tubercle on each side of anterior area. Scutellum with wide, longitudinal dark-brown pubescent band centrally, with pale-yellow pubescence laterally. Elytra. With distinct carina from anterior area between humerus and scutellum to outer apex, more elevated, granulated at anterior fifth (anterior area projected forward); humerus projected; with small, sparse granules between carina and lateral curvature, and near suture; apex truncate, with outer angle acutely projected; with greenish-brown pubescence obscuring integument, interspersed with irregular areas with white pubescence, except: nearly glabrous along carina; two subcircular dark-brown pubescent spots on anterior third (one on each side of carina); two subcircular dark-brown pubescent spots dorsally just after middle, both spots placed between carina and lateral curvature; small, irregular macula with dark-brown pubescence near apex; two subcircular dark-brown pubescent spots on inclined area, one before middle, another after middle; with a few moderately short erect brownish setae on basal third. Legs. Femora pedunculate-clavate, with peduncle of profemora shorter than in meso- and metafemora; with greenish-yellow pubescence partially obscuring integument. Protibia distinctly widened toward apex; with short tooth ventrally at about middle; with greenishyellow pubescence on anterior 2/3 except greenish-brown macula dorsally near middle, brownish, less conspicuous and interspersed with long, erect black setae on posterior third. Meso- and metatibiae with greenish-yellow pubescence, except two wide rings with greenish-brown pubescence, one before middle, another after middle (the latter distinctly darkened laterally and dorsally on mesotibiae); posterior area with long, erect dark setae dorsally. Tarsomeres I with dense, black setae (longer dorsally) interspersed with grayish setae on dorsal base (this area wider on meso- and metatarsomeres); tarsomeres II with dense black setae except yellowish-brown setae on apex of ventral side; tarsomeres III with dense black setae dorsally on about basal 2/3, with yellowish-brown setae on remaining surface (distinctly denser ventrally); tarsomeres V with yellowish-brown setae not obscuring integument.

Abdomen. Sides of ventrites with greenish-yellow pubescence, gradually sparser toward center, especially sparser on anterocentral area of ventrite V. Ventrite V finely, slightly sulcate centrally from base to near apex; posterior third depressed on each side of central area; posterior margin nearly rounded.

Dimensions (mm), 3 females. Total length, 12.70–21.40; prothoracic length, 2.50–4.15; posterior prothoracic width, 3.25–5.85; anterior prothoracic width, 3.30–5.30; widest prothoracic width, 4.90–8.10; humeral width, 5.30–9.60; elytral length, 8.80–14.30.

Material examined. VENEZUELA, 1 female, no date indicated, D. Poggi col. ( MZSP) . Distrito Capital: Caracas (Berg Avila) , 1 female, no date indicated, P.C. Vogl col. ( MZSP) . 1 female, Van Lansberg (no other data) ( MZSP) .

Remarks. The type series includes males and females. Unfortunately, we do not examine males of this species. See remarks in Acanthoderes (Scythropopsis) abstersa .

Figures

FIGURES 20–24. Acanthoderes (Scythropopsis) pupillata Bates, 1880 comb. nov. 20–23, female: 20, Dorsal habitus; 21, Ventral habitus; 22, Head, frontal view; 23, Lateral habitus. 24, Syntype female, dorsal habitus.