Scamandra sanana, Constant & Rbins, 1758

Constant, Jérôme & Rbins, Fulgoroidea. Introduction The study of unidentified material in the accessions of, 1758, Four new species of the Oriental lanternfly genus Scamandra Stål, 1863 from Sulawesi and neighbouring islands with taxonomic notes on the genus (Hemiptera: Fulgoromorpha: Fulgoridae), Belgian Journal of Entomology 50, pp. 1-21 : 1-21

publication ID	https://doi.org/ 10.5281/zenodo.13272198
persistent identifier	https://treatment.plazi.org/id/03EF8795-283D-4F08-F37F-DE6DFB2FFCF9
treatment provided by	Felipe
scientific name	Scamandra sanana
status	sp. nov.

Treatment

Scamandra sanana View in CoL sp. nov.

urn:lsid:zoobank.org:act:AF88E03C-3565-4564-AAEF-F7896B5FD5D4

Figs 1 View Fig , 3 View Fig , 9–11. View Fig View Fig View Fig

ETYMOLOGY. The species epithet is the name of the island from which the type material originates. It is used in apposition.

MATERIAL EXAMINED.

TYPE MATERIAL. INDONESIA, Sanana Island : Holotype ♂: [Coll. I.R.Sc.N.B., Indonesia, Sanana Isl., i. 2012, local collectors, purchased from A. Müller, I.G.: 32.861] ( RBINS). Paratypes: 5 ♀♀: same data as holotype ( RBINS) .

ADDITIONAL MATERIAL. INDONESIA, Sanana Island : 2 ♂♂, 3 ♀♀: Sanana Isl., I.2012 ( TP) .

DIAGNOSIS. (1) disc of frons with large black-brown area surpassing carinae laterally ( Fig. 9 D View Fig ); (2) tegmina mostly dark green or olivaceous green on corium; (3) nodal line of tegmina C-shaped in middle ( Fig. 9 A View Fig ); (4) membrane of tegmina representing about half of LTg; (5) hind wings with anal and sutural zones white ( Fig. 9 A View Fig ); (6) femora brown to black-brown ( Fig. 9 B View Fig ).

DESCRIPTION.

Measurements and ratios. LT: ♂ (n = 1): 27.7 mm; ♀: (n = 5): 33.4 mm (31.6–34.6); LTg/BTg: ♂: 2.11; ♀: 2.25; BV/LV: 3.4; BF/LF: 1.0; BT/LP + LM: 1.2; LM/LP: 1.9; LMe/LCo: 0.97.

18 Head: ( Figs 9 B, D, F View Fig ; 10 B, D, F View Fig ) yellow-brown with disc of frons black-brown and clypeus brown; black brown area of frons extending laterally well beyond carinae; antennae yellow-brown cephalic process reaching posterior margin of vertex ( Figs 9 B View Fig ; 10 B View Fig ); vertex short, with disc wrinkled, and lateral and posterior margins carinate ( Figs 9 B View Fig ; 10 B View Fig ); frons coriaceous with short hairs and 2 longitudinal carinae ( Figs 9 D View Fig ; 10 D View Fig ); labium brown, surpassing metatrochanters ( Figs 9 C View Fig ; 10 C View Fig ); pedicel of antennae kidney-shaped with flagellum inserted dorsolaterally.

Thorax: ( Figs 9 B, D, F View Fig ; 10 B, D, F View Fig ) prothorax yellow-brown with lateral lobes of pronotum largely black-brown anteroventrally; lateral carinae and area between carinae, yellow-brown; wrinkled, with short hairs and 2 impressed points on disc of pronotum; median carina obsolete, barely distinct ( Figs 9 B View Fig ; 10 B View Fig ). Mesonotum yellow-brown, slightly darker than pronotum, with large black-brown marking on disc, reaching anterior margin; apex of scutellum very slightly projecting dorsally.

Tegmina: ( Figs 9 A, C View Fig ; 10 A, C View Fig ) yellow basally, with a broad yellow transverse band before middle; yellow zones marked with white wax on ventral side; nodal line strongly bisinuate and with C-shaped narrow darker marking in middle; nodal line followed by very narrow, whitish line extending on all breadth; membrane brown. Corium showing two types of colouration: (1) in green phase ( Fig. 10 View Fig ): corium dark green with veins green-blue and costal area green-blue with dark green marking; (2) in olivaceous phase ( Fig. 9 View Fig ): corium olivaceous with veins greenish, with irregular darker or reddish markings and costal area paler. Costal margin straight on most length; posterior margin broadly rounded; sutural margin slightly sinuate after nodal line.

Hind wings: ( Figs 9 A, C View Fig ; 10 A, C View Fig ) basicostal angle yellowish white; anal area and broad band along sutural margin white; membrane yellow-brown. Large coloured marking on corium: (1) in green phase ( Fig. 10 View Fig ): dark vinaceous with azureous blue cross-veins; reticulum of cross-veins more dense towards costal margin; (2) in olivaceous phase ( Fig. 9 View Fig ) vinaceous red with bluish white cross-veins; reticulum of cross-veins more blue towards costal margin.

Legs: ( Figs 9 A, C View Fig ; 10 A, C View Fig ) anterior and middle legs entirely brown to black-brown, with femora darker; posterior legs brown with femora darker distally; profemora slightly inflated apically; metatibiae with 4–5 lateral spines, basal one well developed, and 7 apical spines.

Abdomen: ( Figs 9 A, C View Fig ; 10 A, C View Fig ) yellowish brown with terminalia black-brown.

19 Male genitalia: ( Fig. 11 View Fig ) pygofer 3.5 times higher than long in middle in lateral view, with posterior margin slightly sinuate on dorsal ¼ ( Fig. 11 A View Fig ). Anal tube short, transverse, about 1.3 times broader than long in dorsal view, and slightly less than two times broader at apex than at base ( Fig. 11 C View Fig ); sides curved, subparallel on distal half in dorsal view ( Fig. 11 C View Fig ); apical margin pointed in middle dorsally, strongly projecting downwards ventrally ( Fig. 11 A, C View Fig ); in lateral view, apical margin deeply emarginate with ventral portion surpassing dorsal one ( Fig. 11 A View Fig ). Gonostyli 1.67 times longer than high in lateral view, subtriangular with apical angle rounded and dorsal angle roundly right obtuse ( Fig. 11 A View Fig ); laterodorsal tooth of gonostyli projecting lateroventrally, with apex acute and incurved in posteroventral view ( Fig. 11 A, B View Fig ).

DISTRIBUTION. Sanana Island near Sulawesi ( Fig. 3 View Fig ) .

NOTE. Scamandra sanana sp. nov. is rather similar to S. diana Distant, 1892 but differs from the latter (1) the black area of frons broader and largely surpassing carinae (narrower and very slightly surpassing the longitudinal carinae in S. diana ); (2) the apical margin of tegmina rounded at sutural angle (in oblique S. diana ); (3) the membrane of tegmina representing nearly half of LTg (membrane representing only less than 1/3 of LTg in S. diana ).