Sarika pumicata (Morelet, 1875)

Pholyotha, Arthit, Sutcharit, Chirasak, Tongkerd, Piyoros & Panha, Somsak, 2020, Integrative taxonomic revision of the land snail genus Sarika Godwin-Austen, 1907 in Thailand, with descriptions of nine new species (Eupulmonata, Ariophantidae), ZooKeys 976, pp. 1-100 : 1

Sarika pumicata (Morelet, 1875) Fig. 50C, D

Helix pumicata Morelet, 1875: 248, pl. 12, fig. 2. Type locality: "Ajuthia, Siam" [Phra Nakhon Si Ayutthaya Province, Thailand]; Breure et al. 2018: 399, figs 901, 902.

Nanina (Macrochlamys) pumicata : Tryon 1886: 89, pl. 29, figs 40-42; Fischer and Dautzenberg 1904: 395.

Ariophanta (Xesta) pumicata : Fischer 1891: 20.

Sarika pumicata : Godwin-Austen 1907: 181, pl. 116, fig. 1; Sutcharit and Panha 2008: 96.

Macrochlamys pumicata : Panha 1996: 34; Hemmen and Hemmen 2001: 44.

Type material.

Syntype NHMUK 1893.2.4.980-981 (two shells; Fig. 50C, D ) from Siam [Thailand].

Other material examined.

Siam: NHMUK 1901.1.6.246 (one broken shell, specimen dissected by Godwin-Austen 1907) ex. Daly collection.

Diagnosis.

Shell depressed to conoid-depressed, large size (shell width up to 26.0 mm, shell height up to 16.0 mm) and rather thin. Shell surface smooth, slightly shining above and more shining below; shell colour brownish. Whorls 7, increasing regularly; body whorl large and obtusely angulated. Spire high-conical; suture impressed. Aperture crescent-shaped and obliquely opened. Peristome simple. Columellar margin simple. Umbilicus narrowly opened (Fig. 50C, D ).

Remarks.

Godwin-Austen (1907: 181, pl. 116, Fig. 1 ) examined the rehydrated specimens from Thailand and placed this species in the genus Sarika . We have examined the specimen dissected (only shell remained) by Godwin-Austen (NHMUK 1901.1.6.246) and noticed that the shell morphology slightly differs from the syntypes. The dissected specimen has a rounded body whorl and lower spire, whereas the syntypes have obtusely angulated body whorl and higher spire. Unfortunately, we could not find any specimens identical to the syntypes during this survey. Additional specimens with precise collection locality and anatomical studies are necessary to confirm the taxonomic position of this species.

Figures

Figure 1. Maximum likelihood tree showing the relationships among species of Sarika based on the mitochondrial COI gene sequences. Numbers by the nodes are the ML bootstrap values (left) and Bayesian posterior probabilities (right); shown only for the nodes supported by ML or BI (≥ 70 % and ≥ 0.95). Each clade colour refers to a genus.

Figure 50. Shells of Sarika spp. A, B Sarika gratesi sp. nov. A holotype CUMZ 7275 and B paratype CUMZ 7276 C, D S. pumicata syntype NHMUK 1893.2.4.980 - 981 E S. ochtogyra holotype SMF 227103 / 1 F S. benoiti modified from Crosse and Fischer (1863) G S. rex modified from Preston (1909).