Salix rehderiana C.K. Schneider (1916: 66)

He, Li, 2018, Taxonomy and nomenclature of Salix baileyi, S. rehderiana, and S. disperma, Phytotaxa 349 (1), pp. 54-60 : 58

Salix rehderiana C.K. Schneider (1916: 66) .

Salix dolia var. rehderiana (C.K. Schneider) N. Chao & G.T. Gong in Zhao et al. (2012: 5). Type:— China. Sichuan: Baoxing County, 2000 m, 24 April 1912, Wilson 1403 (holotype A-00031203, collected from “cuttings cultivated… in the Arnold Arboretum” ( Schneider 1916), digital images examined, ♀). —For image of the holotype, see Fig. 2B .

= Salix dolia C.K. Schneider (1916: 65) . — Salix rehderiana var. dolia (C.K. Schneider) N. Chao in Chou et al. (1984: 324). Type:— China. Sichuan: Wenchuan County, Niutou Mountain, 3300 m, 20 June 1908, Wilson 2133 (holotype A-00031164; isotypes GH-00106781, BM-000958030, E-00301588, US-00105109; digital images examined; ♀ only). (Image of A-00031164 is available at https:// s3.amazonaws.com/huhwebimages/417BAF8D4BDB46A/type/full/31164.jpg).

Shrubs or small trees. Branchlets brown or dull brown, glabrous or sparsely pubescent. Buds yellowish brown, ovoidoblong. Stipules semiovate-elliptic, 7–8 mm, margin glandular dentate, apex long acuminate; petiole 0.2–0.5 cm, pubescent; leaf blade lanceolate to oblanceolate, 4.7–9.2 (–11) × 1–1.7 (–2.5) cm, abaxially greenish, sparsely short-silky or glabrous, adaxially dark green, pubescent, glabrescent, base cuneate to rounded, margin entire or glandular crenate, revolute, apex obtuse, acute, or shortly acuminate. Catkins just before leaves emerge, densely flowered; male catkin ellipsoid to shortly terete, 1.6–2.7 × 0.6–1 cm; peduncle 0.2–0.5 cm, with 0–2 leaflets; female catkin terete, 1.7–3.9 × 0.6–1 cm, 3.5–8.8 × 0.9–1.2 cm in fruit, rachis clearly visible; peduncle 0.2–0.7 cm, with 0–3 leaflets; floral bract brown or bicolor, oblong, elliptic, or lanceolate, villous. Male flower: adaxial nectary narrowly oblong; stamens 2; filaments distinct or connate at base, glabrous or puberulent at base; anthers golden yellow or purple. Female flower: adaxial nectary narrowly oblong, 0.5–0.7 mm, equal or longer than stipe; stipe 0.3–0.6 mm; ovary narrowly conical, 4–6 mm, downy or subglabrous, ovary beak gradually tapering to style; styles ca. 1.3 mm; stigmas 2 (–4). Capsules brownish, pilose or glabrous.

Phenology: Flowering mid-March to early June, and fruiting in May and June.

Habitat: Mountain slopes, thickets, streamsides; 920–4250 m a.s.l.

Distribution: Western China (Gansu, Ningxia, Qinghai, Shaanxi, Sichuan, Xizang, Yunnan).

Note: —Chao (in Chou et al. 1984) considered Salix dolia similar to S. rehderiana , and treated it as a variety of S. rehderiana . Soon after this study, Chao & Zhou (1985) accepted S. dolia and reduced S. rehderiana to its synonym without explanation. Li (1993) supported a view that S. rehderiana and S. dolia represent the same species but considered S. rehderiana as the accepted name. Subsequently, Chao & Gong (in Zhao et al. 2012) treated S. rehderiana as a variety, S. dolia var. rehderiana .

The author did not find relevant differences between the type specimens of Salix rehderiana and S. dolia . Hence, the author followed Chao & Zhou (1985) and Li (1993) in considering them conspecific. However, S. rehderiana and S. dolia were published in the same volume by Schneider (1916), and therefore have equal nomenclatural priority. Therefore, under the Art. 11.5 of the ICN (Melbourne Code, McNeill et al. 2012), the correct name must be S. rehderiana because S. dolia was treated as a variety of S. rehderiana ( S. rehderiana var. dolia ) by Chao (in Chou et al. 1984) who first united these two taxa.

Representative specimens examined:— CHINA. Gansu: Luqu County, Langmu, 3364 m, 25 June 2012, L. He & X.X. Mao PH20120618-11 ( BJFC). Qinghai: Le Du, Yaocaotai, Xigou, 3033 m, 21 July 2016, L. He & J. Zhao HL 48 ( FJFC). Sichuan: Baoxing County , 2000 m, November 1908, Wilson 1403 (A, ster.) ; Barkam County, Zhegu Mountain , 4031 m, 15 June 2012, L. He & X.X. Mao PH 20120615-11, PH 20120615-12 ( BJFC) ; Tianquan County, 1720 m, 2 April 1953, X.L. Jiang 33700 ( IBSC, PE, SCFI) ; ibidem, 1650 m, 10 April 1953, X.L. Jiang 33803 ( IBSC, PE, SCFI) ; ibidem, 2200 m, 29 April 1953, X.L. Jiang 34042 ( IBK, IBSC, PE) ; Wenchuan County, 3300 m, 20 June 1908, Wilson 2133 a (A, GH, BM, E, US, ♂ only). Xizang: Mainling County, Pai Town , 3600 m, 21 July 1983, Z.C. Ni, S.Z. Cheng & B.S. Li 5712 (PE) ; Mêdog County, Beibeng , 2400 m, 2 May 1983, S.Z. Cheng & B.S. Li 4456 (PE) ; Mêdog County, Hanmi to Pai Town , 3200–4000 m, 4 July 1983, S.Z. Cheng & B.S. Li 5351 (PE). Yunnan: Lijiang , Yulong Snow Mountain , 3100 m, 25 April 1973, Z.F. Fang & S.D. Zhao 525 ( IFP, PE) .

Figures

FIGURE 2. Type specimens of Salix baileyi, S. rehderiana, S. disperma and S. leveilleana.A. holotype of S. baileyi (L.H. Bailey s.n., BH- 000136934, it is used by the permission of L.H. Bailey Hortorium, Sect. of Plant Biology, SIPS, Cornell University (all rights reserved).). B. holotype of S. rehderiana (Wilson 1403, A-00031203, available at https://s3.amazonaws.com/huhwebimages/3ABADA455AC24BD/ type/full/31203.jpg); C. lectotype of S. disperma (N. Wallich 3700 A, BM-000833494, available at http://data.nhm.ac.uk/object/9c1ad515- c912-4141-bbc8-41a741b6e501). D. holotype of S. leveilleana (E.E. Maire s.n., E-00301599, available at http://data.rbge.org.uk/herb/ E00301599).