Salcedia baroensis, Balkenohl, 2020

Balkenohl, Michael, 2020, A genus in disguise. Revision of the genus Salcedia Fairmaire, 1899 with descriptions of nine new species (Coleoptera, Carabidae, Scaritinae, Salcediini), ZooKeys 901, pp. 1-81 : 1

publication ID	https://dx.doi.org/10.3897/zookeys.901.39432
publication LSID	lsid:zoobank.org:pub:843F6065-D00E-48E6-9A26-9CEB044BF102
persistent identifier	https://treatment.plazi.org/id/D305BE20-5EE4-5307-92CB-742FBE62222F
treatment provided by	ZooKeys by Pensoft
scientific name	Salcedia baroensis
status	sp. nov.

Treatment

Salcedia baroensis View in CoL sp. nov. Figs 2 View Figure 2 , 6b View Figure 6 , 19 View Figures 14–19 , 37 View Figures 36–43 , 50 View Figures 44–51 , 63 View Figures 58–63 , 78 View Figures 77–83 , 84 View Figure 84

Type material.

Holotype: ♂, with labels and data: white, black printed, with black cross stripe "Coll. Mus. Tervuren Ethiopie: Ilubabor prov Pokwo, Baro riv 25.VIII 72" / white, black printed "Coll. Mus. Tervuren Ethiopie R.O.S.Clarke" (MRACT). Paratypes: 2 ♂♂, 4 ♀♀, 57 specs., same data as holotype (MRACT, CBB); 1 spec., white, black printed and handwritten with black ink "Illubabor Prov.: POKWO besides Bari Riv. VIII.72" / "Coll. Mus. Tervuren Ethiopie R.O.S. Clarke" (MRACT); 7 specs., white, black printed and handwritten with black ink "Illubabor Prov.: alt. 1800 m POKWO VIII.72" / "Coll. Mus. Tervuren Ethiopie R.O.S. Clarke" (MRACT); 1 ♂, 2 ♀♀, 16 specs., white, black printed "Illubabor Prov.: Gambela 16/19.X.1972" / "Coll. Mus. Tervuren Ethiopie R.O.S. Clarke" (MRACT, CBB); 2 specs., same data but "1 km W of Gambela 17XI.1972" (MRACT); 1 spec., same data but handwritten with black ink “21–23.V.1972” (MRACT).

Diagnosis.

A large sized species, with sub-elongate outline of the elytra with maximum width behind middle and the pronotum with a rudiment of the outer lateral carina at the posterior third. The pseudohumerus is rectangular and shows a laterally projecting tooth. The antennomeres are oblong-elongate. Distinguished most clearly from the similar species S. nigeriensis by the lateral margin of the pronotum with eleven tubercles and the oblique base. In addition, the elytra diverge slightly posteriorly. Moreover, it also differs from S. utetea sp. nov. in the elongate antennomeres, and shorter elytra.

Description.

Measurements in Table 1 View Table 1 .

Colour and surface: Piceous, shiny; top of carinae on head, pronotum and elytra as well as margins of pronotum and elytra opaque, covered with pale grey pili; legs and mandibles fuscous, antennae and palpi leoninous.

Head: Four-fifths of the pronotum width. Outline semi-circular. Clypeus wide, distinctly convex anteriorly, fused with clypeal wings, separated from supra-antennal plates by obtuse notches, with raised pentagon-shaped field at middle, separated from frons by broad flattened shiny transverse furrow. Frons with two raised paramedian carinae, joining anteriorly V-like into a broad slightly elevated keel, with two small glossy teeth bilaterally anterior to central keel, with two short diverging carinae paralaterally at base; frons separated from supraorbital plates by flattened broad furrows; with conspicuously deep and broad pit at front-eye level; supra-antennal and supraorbital plates margined, margin of supraorbital plate distinctly raised, carina-like, supra-antennal plates slightly vaulted. Basal border with narrow emargination at middle, sub-rectangular laterally (angle around 107°). Eyes large, convex, genae slightly convex, both of them just visible from above, with longovoid shape and straight anterior and posterior margin in lateral view. Antenna with segments five to ten oblong-elongate (L/W 1.41), densely pubescent, segments two to four scarcely pubescent, scapus with longitudinal reticulation. Labrum completely covered by clypeus, convex anteriorly. Mandible moderately short, wide, regularly arcuate, more distinct at apex. Mentum small, sub-elongate, isodiametrically reticulated with obtuse tooth at middle, epilobes wide, projecting and angled anteriorly, margined laterally, surface with indistinct punctures, shiny.

Pronotum (Fig. 37 View Figures 36–43 ): Outline rectangular, transverse, a third wider than long. Lateral margin slightly and regularly convex, maximum width at middle. Lateral margin distinctly crenulated, with eleven distinct tubercles, tubercle anterior to basal angle larger, with two notches at posterior angles. Base oblique laterally, with declining flat ridge at middle. Disc flattened in lateral view, with two distinctly raised paramedian carinae slightly concavely depressed in lateral view, running parallel to median line and diverging posteriorly, with median line medium sized, ending in broad pits anteriorly and posteriorly; with two additional shorter carinae bilaterally at base, joining with the paramedian carinae and forming tooth-like tubercle at base pointing posteriorly, without anterior extension, without inner lateral carina, with very short and partly indistinct outer lateral carinae. All carinae sub-crenulate. Lateral margin broadly wing-like bent up, with six large and deep transverse pits, the basal ones somewhat flattened. Space between carinae and pits smooth at middle.

Elytron (Fig. 50 View Figures 44–51 ): Flattened and slightly depressed in anterior half (lateral view), convex in frontal view. Sub-elongate, slightly convex laterally and diverging posteriorly, maximum width slightly behind middle, indistinctly narrowed posterior pseudohumerus. Pseudohumerus rectangular, with laterally projecting tooth. Apex rounded, retracted, with small but acute tooth at suture. Disc with interneur six crenulated; interneur one with traces of carina basally, interneur three slightly carinate in whole length, interneur two running up to apex as slightly convex line, moderately raised, reaching apex; interneur four running in parallel to lateral carina, reaching base, distinctly shortened apically. Interneur five and six with two rows of serial pits, the latter ones partly merging transversally.

Hind wings: Fully developed.

Lower surface (Figs 2 View Figure 2 , 6b View Figure 6 ): Antennal channel of pronotum with isodiametric reticulation. Pseudoepipleura with two rows of pits, indistinctly hollowed out basally, lateral margin of elytron sub-crenulate. Metepisternum elongate, with broad longitudinal groove. Metasternum and abdominal sternite one with numerous irregularly situated larger and smaller pits, sternites two to four smooth, with band of small punctures at middle, sternite one and two with longitudinal reticulation, all other with isodiametric reticulation laterally. Sternum four to six slightly sulcate. Sternum six two irregular transverse rows of punctures, with obtuse transverse sinuate carina apically.

Legs: Profemora with surface indistinctly reticulated. Protibia with short, robust, moderately curved terminal spine, laterally with four teeth of decreasing size, dorsally and ventrally with two carinae. First tarsomeres distinctly elongated, as long as tarsomeres two to four together.

External sexual dimorphism: Not observed.

Male genitalia (Fig. 63 View Figures 58–63 ): Median lobe large, in dorsal view moderately arcuate from base up to apical fifth, in lateral view nearly straight, with fine pili laterally in middle third, apex thickened, convex, club-like, in cross section bilaterally with concavity at apex. Oroficium nearly half as long as median lobe. Endophallus with three groups of fine microtrichia, laterally with few larger ones in between. Dorsal paramere distinctly sinuate, with one minute seta at apex, with distinct apophyses, ventral one like a convex spatula, both parameres slightly distorted.

Female genitalia (Fig. 78 View Figures 77–83 ): Coxostylus slender, distinctly broadened to base, distinctly curved, acute at apex, with fine carina dorsally in apical third, at end of basal third with one larger and six slender nematiform setae laterally, SSO with one microtrichium.

Variation: The slight carina on interneur three of the elytron shows variation in length, width and height, but is developed in all specimens. The indistinct thin carina at the base of interneur one varies much more, and is visible in approximately half of the specimens. At sternum six the number and position of punctures varies. In any case they form two irregular transverse bands of punctures. However, no differences between sexes were observed.

Etymology.

The name refers to the River Baro in the west of Ethiopia where the species has been found.

Distribution

(Fig. 84 View Figure 84 ). Known from several localities at the River Baro, Illubabor province, in Ethiopia.