Sagola walkerae Park and Carlton

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 59

Sagola walkerae Park and Carlton , new species

( Figs. 31d, 31h–j , 32 )

Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ ( NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND NN Beebys Knob 1220m 5 Feb 1978 A.K.Walker ”, “Moss and rotten wood 78/76”, “ N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand”, “ HOLOTYPE Sagola walkerae Park and Carlton 2013 ” . Paratypes (n = 56: 35 males; 21 females). NEW ZEALAND: Nelson : 3♂♂ 7♀♀ ( FMNH), Kahurangi NP, Mt. Arthur tr., below Mt. Arthur Hut , 1200m, 28 XI 2005, Nothofagus forest, FMHD#2005-048 , berl., dead wood, D. Clarke, ANMT site 1157 ; 1♂ 2♀♀ ( FMNH), Kahurangi NP, Mt. Arthur tr., below Mt. Arthur Hut , 1200m, 28 XI 2005, Nothofagus forest, FMHD#2005-049 , berl., leaf litter, A. Solodovnikov, D. Clarke, ANMT site 1157 ; 6♂♂ 3♀♀ ( FMNH), Kahurangi NP, Arthur Range, above Flora Saddle , 1000m, 41″11.351′S, 172″44.456′E, 28 XI 2005, Nothofagus - dominant forest, pyr.-fogging mossy old logs, A. Newton, M. Thayer, ANMT site 1156 ; 4♂♂ 1♀ ( FMNH), Kahurangi NP, Arthur Range, above Flora Saddle , 1000m, 41″11.351′S, 172″44.456′E, 28 XI 2005, Nothofagus -dominant forest, berl., leaf & log litter, A. Newton, M. Thayer, ANMT site 1156 ; 4♂♂ 2♀♀ ( FMNH), Arthur Range, w side Flora Saddle, 950m, 1 I 1985, Nothofagus mossy forest, A. Newton, M. Thayer 725, pyrethrin fogging fungusy Nothofagus log; Buller: 2♂♂ 1♀ (1♂, slide-mounted; FMNH), Nelson Lks. NP, n slope Mt. Robert, Speargrass tr., 880m, 21 XII 1984, Nothofagus forest, A. Newton, M. Thayer 704, Pyrethrin fogging fungusy Nothofagus logs ; 6♂♂ 4♀♀ ( FMNH & DSC), Nelson Lks . NP, n slope Mt. Robert, Pinchgut tr., 1290m, 18–26 XII 1984, Notho. sol. elfin forest, A. Newton, M. Thayer 716, Pyrethrin fogging fungusy Nothofagus logs ; 5♂♂ ( FMNH & DSC), Nelson Lks. NP, n slope Mt. Robert, Pinchgut tr., 950m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 707, window trap ; 1♂ ( FMNH), Nelson Lks. NP, Lake Rotoiti, St. Arnaud tr., 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT & window trap ; 1♂ ( DSC), Lewis Pass Nat. Res., 0.6km s Lewis Pass , 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, berl., leaf & log litter, forest floor ( DSC) ; 1♂, Nelson Lks. NP , Robert Ridge, nr. Julius Summit, ca. 1700m, 41″52′S 172″46.5′E, 1 XII 2005, FMHD#2005-063 , berl., debris at Bases of tussocks, A. Solodovnikov ; 1♂ ( NZAC), Paparoa Ra., 1036, Mt. Dewar , 12 XII 1969, J.C. Watt, moss 69/260 ; 1♀ ( NZAC), Reefton , 12 IV 1977, J.A. Wightman, pit trap cutover pine .

Etymology. This species is named for the collector of the holotype, Annette K. Walker.

Diagnosis. This species can be distinguished from the other species of this species-group by the following combination of characters: body length 2.3–2.5 mm; ventral surface of male head transversely convex with setose gular region; eye large and prominent, approximately one-half length of temple; antennomeres 3–10 subquadrate; shape of genitalia unique to species.

Description. Length 2.3–2.5 mm. Body reddish brown, antennae, elytra, legs, and maxillary palpi paler ( Fig. 31d ). Head: Bluntly triangular, widest across eyes ( Fig. 31j ), ventral surface of male head transversely convex with setose gular region. Antennomere 1 approximately 2 times longer than wide, 2 longer than wide, 3–10 subquadrate ( Fig. 31i ). Frontal rostrum prominent, covering anterior frontal fovea. Frontal sulcus deep, reaching hind point of eye from end of frontal rostrum. Anterior frontal fovea oval, posterior frontal fovea elongate. Eye large and prominent, approximately one-half length of temple. Thorax: Prosternum longer than wide, widest at one-third length. Elytra rectangular ( Fig. 31d ). Meso- and metathorax trapezoidal, longer than wide. Abdomen: Male tergite IV with pair of transverse patches of microtrichia, less developed in female. Abdominal ventrites III–V with very small basolateral fovea. Aedeagus: Median lobe divided, major lobe as wide as minor ( Fig. 31h ). Phallobase of median lobe symmetrical and rounded ( Fig. 31h ). Parameres symmetrical, broad with setae apically ( Fig. 31h ).

Distribution. Buller, Nelson ( Fig. 32 : stars).

Habitat. Most specimens of this species were collected at high elevations (> 670 m) by fogging, with window and flight intercept traps, or by sifting leaf litter in Nothofagus mossy forests.

Figures

Fig. 31. Sagola species in the rustica species-group. Habitus, dorsal views: a) S. rustica, b) S. duplicata, c) S. marlboroughensis, d) S. walkerae. Aedeagi, dorsal views: e) S. rustica, f) S. duplicata, g) S. marlboroughensis, h) S. walkerae. S. walkerae, male: i) Antenna, j) Dorsal surface of head. Scale bars a–d = 1 mm, e–j = 0.1 mm.

Fig. 32. Known collection localities of New Zealand Sagola species. S. rustica: black circle; S. duplicata: triangles; S. marlboroughensis: black square; S. walkerae: stars.