Roeweriinae, Rafael N. Carvalho & Adriano B. Kury, 2018

Rafael N. Carvalho & Adriano B. Kury, 2018, Further dismemberment of Discocyrtus with description of a new Amazonian genus and a new subfamily of Gonyleptidae (Opiliones, Laniatores), European Journal of Taxonomy 393, pp. 1-32 : 10-12

Roeweriinae subfam. nov.

urn:lsid:zoobank.org:act:237C9E13-E34E-44BC-AD97-5B6ACFE915AC

Included genera

Amazochroma gen. nov., Discocyrtanus Roewer, 1929 and Roeweria Mello-Leitão, 1923 (type genus; revalidated by Kury 2003). Discocyrtanus and Roeweria were hitherto in Pachylinae .

Ampycus telifer 0 0 0 - 0 0 0 2 1 1 5 3 1 0 0 1 0 0 1 0 0 1 0 1 2 0 0 0 1 0 1 0 0 0 0 - 0 0 0 1 0 0 0 0 1 2 1 0 1 0 1 0 0 0 0 1 0 - - - 0 1 0 0 0 0

Acanthopachylus aculeatus 0 0 0 - 0 0 1 2 0 0 1 1 0 0 0 0 - 1 0 0 0 1 0 1 1 1 0 1 0 1 1 0 0 0 0 - 0 0 0 0 0 0 0 1 1 1 1 1 0 1 1 0 0 0 0 0 1 0 0 - 1 1 1 1 0 0

Pachylus chilensis 0 0 0 - 0 0 1 1 0 0 1 1 0 1 0 0 - 1 1 0 0 1 0 1 1 0 0 1 0 0 1 0 2 0 0 - 0 0 0 0 0 0 1 1 1 1 1 1 0 1 1 0 0 0 0 0 1 0 0 - 1 1 1 1 0 0

Anoplogynus nasutus 0 0 0 - 2 0 1 2 0 1 4 0 0 1 0 0 - 0 1 0 0 1 0 0 - 0 0 0 0 1 1 0 0 0 0 - 0 0 1 5 0 0 0 1 1 2 0 - 1 1 0 0 0 0 0 0 0 - - - 2 0 - 0 0 0

Gonyleptes horridus 0 0 0 - 1 0 2 0? 1 0 0 1 0 0 0 - 0 1 0 - 1 1 0 - 0 0 1 0 0 1 1 1 0 0 - 0 0 0 1 0 0 0 0 1 2 0 - 1 0 0 0 0 0 2 0 1 1 1 0 3 1 3 1 0 0

Neopachylus imaguirei 0 0 0 - 1 0 1 1 0 1 0 0 1 0 0 0 - 0 1 1 1 0 - 0 - 0 0 0 0 1 0 1 1 0 1 1 1 0 1 0 0 0 0 0 1 0 1 0 1 1 0 0 0 0 1 0 1 1 1 2 4 0 - 2 0 0

Roeweria virescens 1 0 0 - 3 0 3 0 1 1 2 0 1 0 1 0 - 0 2 1 0 1 1 1 0 0 0 1 1 0 0 0 2 1 1 0 1 0 0 1 0 0 0 0 1 2 1 0 0 2 2 1 1 0 1 1 0 - - - 5 1 2 3 0 1

Roeweria bittencourti 1 0 0 - 3 0 3 0 1 1 2 0 1 0 1 0 - 0 2 1 0 1 1 1 0 0? 1 1 0 0 0 2 1 1 0 1 0 0 1 0 0 0 0 1 0 0 - 1 2 0 1 1 0 1 1 0 - - - 5 1 2 3 0 1

Amazochroma carvalhoi gen. et comb. nov. 0 1 1 2 1 0 3 1 1 1 2 2 1 0 0 1 0 0 2 1 0 1 1 1 0 0 1 1 1 0 0 0 2 1 1 0 1 1 1 0 0 0 1 1 0 2 1 0 1 2 0 1 1 0 1 0 0 - - - 5 1 2 3 0 1

Amazochroma pedroi gen. et sp. nov. 0 1 1 2 1 0 3 2 1 1 2 2 1 0 0 1 0 0 1 1 0 1 1 1 0 0 1 1 1 1 0 0 2 1 0 - 0 1 1 0 0 0 1 1 0 2 1 0 1 2 0 1 1 0 1 0 0 - - - 5 1 2 3 0 1

Discocyrtanus goyazius 0 0 1 0 1 0 3 1 1 1 2 0 1 0 0 1 0 0 1 1 0 1 1 1 0 0 1 1 1 0 0 0 1 1 1 0 1 0 1 3 1 0 1 1 1 2 1 0 1 2 0 1 1 0 0 0 0 - - - 5 1 2 3 0 1

Discocyrtanus oliverioi 0 0 1 0 1 0 3 1 1 1 2 0 1 0 0 1 0 0 1 1 0 1 1 1 0 0? 1 1 0 0 0 1 1 1 0 1 0 1 3 1 0 1 1 1 2 1 0 1 2 0 1 1 0 0 0 0 - - - 5 1 2 3 0 1

Discocyrtus crenulatus 0 0 0 - 1 1 3 2 0 2 0 0 0 1 0 0 - 0 1 1 1 1 1 1 0 1 1 1 1 0 0 0 1 1 1 1 1 0 0 2 0 0 0 1 1 1 1 0 1 1 0 0 0 1 2 0 1 1 1 1 5 1 0 1 1 0

Discocyrtus testudineus 0 0 0 - 1 1 3 2 0 1 2 0 0 1 0 0 - 0 1 1 1 1 1 1 0 1 1 1 1 0 1 0 1 1 1 1 1 0 0 2 0 0 0 1 1 1 1 1 1 1 0 0 0 1 2 0 1 1 1 1 5 1 0 1 1 0

Discocyrtoides nigricans 2 0 1 1 1 0 3 0 0 1 0 0 1 0 1 0 - 0 2 1 1 1 1 1 0 1 1 1 0 0 0 0 1 1 1 1 1 0 0 4 0 1 0 0 1 0 0 - 1 1 0 0 0 1 2 0 1 1 1 1 5 1 0 1 1 0

Mitobates triangulus 2 0 0 - 4 0 3 0 2 1 3 0 0 0 0? 1 0 2 1 1 1 1 1 0 1 1 1 0 0 0 0 - - 0 - 1 0 0 4 0 1 0 0 1 0 0 - 1 3 0 0 0 1 2 0 1 1 1 0 5 1 1 1 1 0

Diagnosis

Dry marks either around the tubercles or restricted to their tips, but in both cases all around dorsal scutum (only in Amazochroma gen. nov. and Discocyrtanus , absent in Roeweria ; contrasting with Discocyrtus and some Mitobatinae [e.g., in Ischotherus Kury, 1991, Mitobates Sundevall, 1833 Neoancistrotus Mello-Leitão, 1927 , Promitobates Roewer, 1913 , Ruschia Mello-Leitão, 1940 ] where it is absent and with other Mitobatinae which possess dry marks arranged in broad stripes, often cruciform [e.g., in Discocyrtoides Mello-Leitão, 1923 , Longiperna Roewer, 1929 ). Ocularium convex, with a median depression (contrasting with the convex form without depression in Discocyrtus and Discocyrtoides or rectangular form in Mitobates ). Mesotergum divided into 4 areas (same as in Discocyrtus testudineus Holmberg, 1876 and Mitobatinae ). Area I with a pair of conspicuous paramedian tubercles (same as D. testudineus , absent in D. crenulatus Roewer, 1913 and Discocyrtoides ). Retrodorsal distal apophysis of Cx IV of males with only a main tubercle (absent in Amazochroma pedroi gen. et sp. nov., contrasting the form with a geminated tip present in Discocyrtus and some Mitobatinae ). Distal part of the VP of penis forming a trapezium, with an apical minor base (contrasting with a trapezium with an apical major base occurring in Discocyrtus and Discocyrtoides or the M-form of Mitobates ). Macrosetae C1–C3 of VP elongate and slender (contrasting with the short and thick form found in Discocyrtus and Mitobatinae ). Macrosetae C1–C3 insertion with C 3 set widely apart from the other C, forming a diastema (present in Amazochroma gen. nov. and Roeweria , contrasting with all macrosetae C placed distally with subequal intervals between them as found in Discocyrtanus and with macrosetae C placed distally, united as in Discocyrtus and Mitobatinae ). Ventral of glans process extremely reduced or absent (contrasting with the presence of a stem and a flabellum in Discocyrtus and Mitobatinae ). Stylus sigmoid, strongly bent to dorsal proximally and overall curved to ventral more distally (present in Amazochroma gen. nov. and Discocyrtanus , contrasting with the C-shape, overall arched to dorsal found in Roeweria and the substraight shape as in Discocyrtus and Mitobatinae ). Angle of basal curvature of stylus around 90 degrees (in opposition to form with less than 90 degrees in Discocyrtus and Discocyrtoides or more than 90 degrees in Mitobates ). Apical winglets of stylus (present in Amazochroma gen. nov., Discocyrtanus and Roeweria , absent in Discocyrtus and Mitobatinae ).