Rhynchosauroides kuletae, Ptaszyński & Niedźwiedzki, 2004

Ptaszyński, Tadeusz & Niedźwiedzki, Grzegorz, 2004, Late Permian vertebrate tracks from the Tumlin Sandstone, Holy Cross Mountains, Poland, Acta Palaeontologica Polonica 49 (2), pp. 289-320 : 297-300

publication ID

https://doi.org/ 10.5281/zenodo.13510407

persistent identifier

https://treatment.plazi.org/id/03F387F5-1639-7C34-FFBE-FBDB8B926107

treatment provided by

Felipe

scientific name

Rhynchosauroides kuletae
status

 

Rhynchosauroides kuletae ichnosp. nov.

Figs. 9A–C View Fig , 10A–D View Fig , 11A, B View Fig , 12A–D View Fig .

Rhynchosauroides cf. pallinii Conti et al., 1977 ; Ptaszyński 2000b: fig. 2, trackway described in this paper as Muz. PIG OS−220/26: 1–5.

Holotype. Set of left pedal and manual imprints Muz. PIG OS−220/27: 11, 12 ( Figs. 9A View Fig 1 View Fig , 10 A View Fig 1 View Fig , 11A View Fig 1 View Fig , 12A View Fig 1 View Fig ).

Type locality: Tumlin Gród quarry, 11 km NW of Kielce, Holy Cross Mts., Poland.

Other localities: Sosnowica Hill quarry, 8 km N of Kielce, Holy Cross Mts., Poland.

Type horizon: Tumlin beds, Lower Buntsandstein, uppermost Permian. Derivation of the name: In honour of Maria Kuleta, researcher and expert of the Holy Cross Mountains Buntsandstein lithostratigraphy and sedimentary environment.

Material.—Tumlin Gród quarry: Muz. PIG OS−220/26: 1–5 trackway consist of two sets and the next consecutive manual imprint; Muz. PIG OS−220/27: 1–14 seven sets of imprints ( Figs. 9A View Fig 1 View Fig , 10A View Fig 1 View Fig , A 2 View Fig , 11A View Fig 1 View Fig , 12A View Fig 1 View Fig ), among them the fifth, left set of pedal and manual imprints (Muz. PIG OS−220/27: 11, 12 is designated as the holotype); Muz. PIG OS−220/27: 15–31 nine consecutive sets of imprints ( Figs. 9A View Fig 2 View Fig , 10A View Fig 3 View Fig , A 4 View Fig , 11A View Fig 2 View Fig , 12A View Fig 3 View Fig ); Muz. PIG OS−220/27: 119–136 nine consecutive sets of imprints; Muz. PIG OS−220/28: 1–11 six consecutive sets of imprints, the second represented only by manual imprint ( Figs. 9B View Fig , 10C View Fig , 11B View Fig , 12B View Fig ); Muz. PIG OS−220/29: 1 isolated left pedal imprint; Muz. PIG OS−220/73: 1–26 eleven consecutive sets of pedal and manual imprints ( Figs. 9C View Fig , 12C View Fig 2 View Fig ); Muz. PIG OS−220/73: 27–31 three consecutive sets of pedal and manual imprints; Muz. PIG OS−220/73: 88, 89 set of right pedal and manual imprints, the largest specimen accounted to this ichnospecies ( Figs. 10B View Fig , 12C View Fig 1 View Fig ); Muz. PIG OS−220/73: 33–38 three consecutive sets of imprints; Muz. PIG OS−220/73: 39–41 right pedal imprint and the next (left) set of imprints; Muz. PIG OS−220/27: 76, 77 isolated set of right pedal and manual imprints ( Fig. 12A View Fig 2 View Fig ); Muz. PIG OS−220/27: 84, 85 isolated set of imprints; Muz. PIG OS−220/27: 78, 79 isolated set of imprints. Sosnowica Hill quarry: Muz. PIG OS−220/1 set of right pedal and manual imprints; Muz. PIG OS−220/6: 1, 2 set of left pedal and manual imprints ( Figs. 10D View Fig , 12D View Fig ).

Diagnosis.—Large Rhynchosauroides with a total pedal length of about 60 mm. Pedal pace angulation ranges from about 100° up to 111–116°. Both pedal and manual digit V is relatively long and distinct. The pedal axis diverges from the midline at –4°, while manual one is inclined to the midline at about +23°. Divarication of manual digits I– V about 90°. The stride/pedal length ratio attains 7.2.

Description.—Trackway. Trackway features typical of the ichnogenus Rhynchosauroides . The relative position of manus and pes in the same set varies in known trackways. Pedal imprints may distinctly overstep manual ones (Muz. PIG OS−220/27: 15–31); both may be on the same line (Muz. PIG OS−220/27: 1–14, Figs. 9A View Fig 1 View Fig , 11A View Fig 1 View Fig ) or manual imprints may be situated before pedal ones (Muz. PIG OS−220/28: 1–11, Figs. 9B View Fig , 11B View Fig ); Muz. PIG OS−220/73: 1–26, Fig. 9C View Fig ). Pedal axes diverge slightly from the midline at about –4° on average; the average inclination to the midline of manual axes is +23°. The length of stride ranges from 340 mm up to 447 mm. In all known trackways pedal and manual pace angulation attain 100–116° and 117–153°, respectively. The stride/pedal length ratio is high, up to 7.2.

Pes.— The length of pedal imprints is about 60 mm. The largest known specimen shows a pes length of about 65 mm (Muz. PIG OS−220/73: 88, 89, Figs. 10B View Fig , 12C View Fig 1 View Fig ). There are specimens from Sosnowica Hill quarry (Muz. PIG OS−220/1: 1, 2 and Muz. PIG OS−220/6: 1, 2; Figs. 10D View Fig , 12D View Fig ) with pedal imprints only 40 mm long. The digit group I– IV is relatively short and wide, with typical digit length ratios of Rhynchosauroides , I< II < III < IV: 23, 29, 32, and 34 mm in the holotype. Divarications of digits II – IV and I– IV are 22 and 33° on average, respectively. The digit V, distinctly visible in most of known specimens ( Figs. 10A, B, D View Fig , 12A, B, C View Fig 1 View Fig , D), as long as digit III or IV, diverges from digit I at about 64°. Deep imprints show its whole shape, as like as the whole pedal imprint; in some shallower imprints only a proximal, rounded pad and the distal part of the digit, with a distinct, big claw mark is visible ( Figs. 10A View Fig 1 View Fig , A 2 View Fig , A 4 View Fig , 12A View Fig 1 –A View Fig 3 View Fig ) .

Manus.—Manual imprints are relatively large ( Figs. 10A View Fig 1 View Fig , B, C, 12A View Fig 1 View Fig , A 2 View Fig , B, C 1 View Fig ). Digits I– IV increase in length from I to IV: 14, 21, 24, 28 mm in the holotype. The digit group I– IV is 32.4 mm long by 34.1 mm wide, on average. Digit V is as long as digit III, diverge from digit I at 75–90°; this value does not exceed the right angle. All digits have claw marks. Discussion.— Rhynchosauroides kuletae ichnosp. nov. shows similarity to Rhynchosauroides pallinii Conti et al., 1977 , and is also of the same size. Rhynchosauroides kuletae ichnosp. nov. however differs from the latter as it has higher pace angulation and a relatively bigger digit V, both in pedal and manual imprints. Its manual digit V diverges from the digit IV at smaller angle, being perpendicular not to the digit IV as in R. pallinii , but to digit I.

Ichnogenus Palmichnus Schmidt, 1959 Remarks.—Medium−sized Rhynchosauroidae representatives. The manual pace width is close to the pedal one. Pace angulation about 90° or more. Manual imprints situated before those of pes or on the same line. The pedal digit V is as long as digit III. In both pedal and manual imprints digits diverge at high angles; that of digits I–V attains 90° in pedal imprints and much more in those of manus. Stride/pedal length ratio is relatively low, and does not exceed 5. Except for Palmichnus tambachensis Haubold, 1970 considered (Haubold 1998) as synonym of Varanopus microdactylus , there are only two known ichnospecies representing this ichnogenus: Palmichnus renisus Schmidt, 1959 and Palmichnus lacertoides ichnosp. nov.

V

Royal British Columbia Museum - Herbarium

Kingdom

Animalia

Phylum

Chordata

Genus

Rhynchosauroides

Loc

Rhynchosauroides kuletae

Ptaszyński, Tadeusz & Niedźwiedzki, Grzegorz 2004
2004
Loc

Palmichnus lacertoides

Ptaszyński & Niedźwiedzki 2004
2004
Loc

Rhynchosauroides cf. pallinii

Conti 1977
1977
Loc

Palmichnus tambachensis

Haubold 1970
1970
Loc

Rhynchosauroidae

Haubold 1966
1966
Loc

Palmichnus

Schmidt 1959
1959
Loc

Palmichnus renisus

Schmidt 1959
1959
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