Reicheadella smetanai, Bulirsch, Petr & Guéorguiev, Borislav V., 2008

Bulirsch, Petr & Guéorguiev, Borislav V., 2008, Taxonomic studies of the Balkan Reicheina (Coleoptera: Carabidae: Clivinini), with a review of the Albanian species, Zootaxa 1679, pp. 39-53 : 45-47

Reicheadella smetanai sp. nov.

( Figs 3 , 15 , 16 )

Type material. Holotype & labeled: “ Albania mer. Liqueni i Butrintit, 1.V.1958; A. Smetana leg.”; “ Holotype, Reicheadella smetanai sp. n. Bulirsch et Guéorguiev des., 2006” ( NMP).

Description. Body yellow­brown, legs, antennae and mouthparts rusty yellow, front legs slightly darker ( Fig. 3 ). Length 2.05 mm.

Head. Moderately broad, long, anterior margin of clypeus almost straight, frons with facial furrows shortened, shallow, broad. Impressions on clypeus oblique, broad, shallow. Eyes absent, genae with median part short, regularly vaulted, their hind angles rounded. Vertex with slightly reticulated microsculpture. Antennae length 0.69 mm; antennomere 2 slightly longer than 3 and 4 combined, antennomeres 6–10 as long as wide. Mandibles rather long, their apical part slightly curved. Terminal maxillary palpomeres moderately long, with apex slightly narrowed, not acicular.

Pronotum. Moderately convex, microsculpture regular, but reticulation hardly visible. Sides regularly, moderately rounded, slightly attenuated posteriorly; maximum width before middle; posterior angles moderately rounded. Reflexed lateral margin entire, extended from blunt, rather protruding anterior angles to base as fine prebasal groove. Median line narrow, distinctly impressed almost towards prebasal groove; anterior transverse impression shallow, slightly deeper laterally. Episterna just visible from above in apical part. Reflexed lateral margin with 2 pairs of standard lateral setiferous punctures; disc with 2 pairs of submedial setae; the first very distinct pair in middle of fore third and the second, much finer pair in posterior third, close to median line. Ratio length: width 1.00; pronotum 1.42 times as broad as head.

Legs. Protibia with apical spine bent outwards in dorsal view, apical spur of almost equal length, slightly curved. Lower marginal tooth large, prominent, upper one distinct, sharp. Hind tarsi length 0.23 mm; first tarsomere not elongated.

Elytra. Convex, disc slightly flattened, outline almost oval, maximum width at middle. Base slightly bordered, slightly sloping, without distinct granules; prescutellar setiferous punctures isolated; suture not depressed; humeri rounded, not protruding. Lateral channel very wide, reflexed lateral margin with several (about 30) denticles; in basal two thirds denticles very dense, sharp, apically much finer, but recognizable up to apex with slightly protruded suture. Striae coarsely punctate; first striae deeper than striae 6–7; all striae strongly diminish apically, first striae in apical fifth, lateral ones in apical third; apically with only traces of fine punctures. Striae 1–5 distinct up to base, striae 6 and especially 7 shortened. Intervals 1–5 in basal part convex, lateral ones flattened. Intervals 3, 5, 7 with rows of setae. Ratio length: width 1.77; ratio elytra: pronotum length 2.34; ratio elytra: pronotum width 1.32.

Venter. Last visible ventral segment in female with moderately reticulated microsculpture in apical two thirds.

Aedeagus. Unknown.

Female genital armature. Ovipositor as in Fig. 15 ; apical and basal stylomeres of both styli tightly fused; dorsal ensiform setae varied in number; two dorsolateral ensiform setae long and thick; nematiform not visible (probably absent); valvifer long and thin.

Differential diagnosis. R. smetanai sp. nov. belongs to the genus Reicheadella ; it can be easily distinguished from R. bischoffi , the only other known Albanian species, as well as from the Greek species of this genus, by the smaller size of the body, by only 2 pairs of the discal submedial setae and by several elytral lateral denticles.

Etymology. The species is named in honour of Aleš Smetana, Ottawa, Canada, collector of the holotype. Type locality. South Albania: Liqueni i Butrintit, more precise data not available.

Figures

FIGURE 15. Ventral view of styli. Reicheadella smetanai sp. nov., holotype. Scale bar = 0.1 mm.

FIGURE 16. Map of Albania with localities of species. Star: Dalmatoreicheia maderi sp. nov.; black circle: Reicheadella bischoffi; reverse triangle: R. smetanai sp. nov.

FIGURES 1 – 3. Habitus of type species (length in parentheses behind the name). Fig. 1, Dalmatoreicheia maderi sp. nov., (3.20 mm), holotype; Fig. 2, Reicheadella bischoffi, (2.80 mm), lectotype; Fig. 3, Reicheadella smetanai sp. nov., (2.05 mm), holotype.