Reicheadella bischoffi ( Meschnigg, 1933 )

Bulirsch, Petr & Guéorguiev, Borislav V., 2008, Taxonomic studies of the Balkan Reicheina (Coleoptera: Carabidae: Clivinini), with a review of the Albanian species, Zootaxa 1679, pp. 39-53 : 42-43

Reicheadella bischoffi ( Meschnigg, 1933)

( Figs 2 , 5 , 8 , 10 , 13 , 16 )

Type material. Lectotype (here designated) ɗ labelled: “Tomor Juli 1932 ”; “ Albania leg. Bischoff 1932”; “ Reicheia Bischoffi , det Ing. Meschnigg, Type ”; “ Lectotypus, Reicheadella bischoffi ( Meschnigg, 1933) , Bulirsch et Guéorguiev des. 2006” ( MNHUB). Paralectotype & labelled: “Tomor Juli 1932 ”; “ Albania leg. Bischoff 1932”; “ Reicheia Bischoffi , det Ing. Meschnigg, Type ”; “ Paralectotypus, Reicheadella bischoffi ( Meschnigg, 1933) , Bulirsch et Guéorguiev des. 2006” ( MNHUB).

Additional material studied. 1 ɗ labelled: “Tomor Juli 1932 ”; “ Albania leg. Bischoff 1932” (cPB); 1 ɗ labelled: “Tomor: Tomor Alban. mer.”; “lg. Winkler Mai 1931 ”; “ Bischoffi Meschn. det. A. Winkler” ( NMW); 1 & labelled “Tomor: Tomor Alban. mer.”; “lg. Winkler Mai 1931 ” ( NMW).

Redescription. Body rusty brown, legs and especially antennae and mouthparts lighter ( Fig. 2 ). Length 2.80–2.95 mm (in lectotype 2.80 mm).

Head. Moderately broad, long, anterior margin of clypeus almost straight, frons with facial furrows moderately deep, broad. Impressions on clypeus oblique, deep; clypeus prolonged apically by rather long, sharp keel. Eyes absent, genae with median part long, slightly vaulted, their hind angles rounded. Vertex with irregularly reticulated microsculpture. Antennomere 2 slightly longer than 3 and 4 combined, antennomeres 6–10 as long as wide. Mandibles rather long, their apical part moderately curved. Terminal maxillary palpomeres moderately long, with apex slightly narrowed, not acicular.

Pronotum. Moderately convex, microsculpture irregular, reticulation just visible. Sides regularly, moderately rounded, not attenuated posteriorly; maximum width at about middle; posterior angles broadly rounded. Reflexed lateral margin entire, extended from rather protruding anterior angles to base as rather fine prebasal groove. Median line narrow, moderately impressed almost towards prebasal groove; anterior transverse impression very superficial, just visible. Episterna just visible from above in very apical part. Lateral channel with 2 lateral setiferous punctures; disc with 3 pairs of submedial discal setae (especially median and posterior pairs aproximate to median line) and without sublateral discal setae. Ratio length: width 0.92–0.96 (in lectotype 0.94); pronotum 1.52–1.59 (in lectotype 1.52) times as broad as head.

Protibia. Apical spine bent outwards in dorsal view, apical spur of almost equal length, slightly curved. Lower marginal teeth very distinct, sharp, upper one smaller, sharp.

Elytra. Convex, disc flattened, outline almost ovale, maximum width at middle. Base distinctly bordered, without distinct granules, moderately sloping; prescutellar setiferous punctures isolated; suture distinctly depressed; humeri rounded, moderately protruding. Lateral channel wide, reflexed lateral margin with 3–6 fine humeral denticles. Striae 1–5 distinct, distinctly punctate, striae 6–7 slightly finer, moderately punctate; all striae, except first, diminish apically, first striae in apical sixth, lateral ones in apical fifth. Striae 1–3 distinct almost up to base, lateral striae slightly to moderately shortened. Intervals in basal part moderately convex. Intervals 3, 5, 7 with rows of setae. Ratio length: width 1.69–1.71 (in lectotype 1.70); ratio elytra: pronotum length 2.36–2.43 (in lectotype 2.43); ratio elytra: pronotum width 1.31–1.35 (in lectotype 1.35).

Venter. Last visible ventral segment in males with slightly reticulated microsculpture in apical half; in females with moderately reticulated microsculpture in apical two thirds.

Male genital armature. Urite as in Fig. 5 ; semicircle, briefly rounded in basal part and widely rounded in apical part, with prominent apical process, distinct subapical vertical lamella and subbasal swelling; sclerites fused. Median lobe as in Figs 8 , 10 ; slightly arcuate in lateral view, apical part wider than basal part, with well­formed hook at tip; both apical and basal orifices concave; internal sac situated in subapical part; copulatory piece with vague inner structures; median lobe in ventral view wide, almost straight, apically with thin and straight lateral tip. Parameres similar in shape and size as in Fig. 13 ; left paramere somewhat smaller, slightly bent near to middle, apex with one short and two long setae situated at equal distances from one another; right paramere elongate, apex with one short and two long setae situated on equal distances from one another.

Female genital armature. Not examined.

Differential diagnosis. Differences between R. smetanai sp. nov., the second known Albanian species, are quoted below and the remaining species were differentiated in Jeannel (1957); Casale et al. (1998) and Giachino & Vailati (2004).

Type locality. The Tomor Mountains in South Albania.

Remark. Lectotype designation is made to stabilize the nomenclature of this species and to give preference to the male syntype illustrated here.

Figures

FIGURE 16. Map of Albania with localities of species. Star: Dalmatoreicheia maderi sp. nov.; black circle: Reicheadella bischoffi; reverse triangle: R. smetanai sp. nov.

FIGURES 1 – 3. Habitus of type species (length in parentheses behind the name). Fig. 1, Dalmatoreicheia maderi sp. nov., (3.20 mm), holotype; Fig. 2, Reicheadella bischoffi, (2.80 mm), lectotype; Fig. 3, Reicheadella smetanai sp. nov., (2.05 mm), holotype.

FIGURES 4 – 6. Ventral view of postabdominal sternum IX (urite). Fig. 4, Dalmatoreicheia maderi sp. nov., holotype; Fig. 5. Reicheadella bischoffi, lectotype; Fig. 6, Reicheadella corcyrea, male labelled “ Gasturi, Korfu 2. IV. (19) 29. Beier ” (NMW). Scale bar = 0.1 mm.

FIGURES 7 – 8. Left lateral view of median lobe of aedeagus. Fig. 7, Dalmatoreicheia maderi sp. nov., holotype; Fig. 8. Reicheadella bischoffi, lectotype. Scale bar = 0.1 mm.

FIGURES 9 – 11. Ventral view of median lobe of aedeagus. Fig. 9, Dalmatoreicheia maderi sp. nov., holotype; Fig. 10, Reicheadella bischoffi, lectotype; Fig. 11, Reicheadella corcyrea, male labelled “ Gasturi, Korfu 2. IV. (19) 29. Beier ” (NMW). Scale bar = 0.1 mm.

FIGURES 12 – 14. Ventral view of parameres. Fig. 12, Dalmatoreicheia maderi sp. nov., holotype; Fig. 13, Reicheadella bischoffi, lectotype; Fig. 14, Reicheadella corcyrea, male labelled “ Gasturi, Korfu 2. IV. (19) 29. Beier ” (NMW). Scale bar = 0.1 mm.