Raveniola karategensis, Zonstein, 2024

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 38-39

publication ID	https://doi.org/ 10.5852/ejt.2024.967.2699
publication LSID	lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738
persistent identifier	https://treatment.plazi.org/id/631DA123-F0BE-4C2B-A58E-F073BB22E75B
taxon LSID	lsid:zoobank.org:act:631DA123-F0BE-4C2B-A58E-F073BB22E75B
treatment provided by	Plazi
scientific name	Raveniola karategensis
status	sp. nov.

Treatment

Raveniola karategensis sp. nov.

urn:lsid:zoobank.org:act:631DA123-F0BE-4C2B-A58E-F073BB22E75B

Figs 44 View Figs 36–44 , 119 View Figs 118–126 , 176 View Figs 172–183 , 238 View Figs 238–246 , 316–317 View Figs 310–318 , 510–512 View Figs 504–521 , 576 View Figs 575–583 , 659–666, 675, 678–679, 753

Diagnosis

The new species shares with Raveniola afghana sp. nov., R. alajensis sp. nov. and R. hirta sp. nov. the presence of the modified long hairs on the female tibia and metatarsus IV. Females of R. karategensis sp. nov. can be distinguished from those of R. afghana and R. hirta by having a fuzzily ornamented (vs uniformly coloured) abdomen (see Fig. 44 View Figs 36–44 cf. Figs 37, 42 View Figs 36–44 ), and from females of R. alajensis in possessing a less pronounced abdominal pattern and the distinctly configured spermathecae with wider bases, broadly spaced stalks and clearly wider inner branches ( Figs 44 View Figs 36–44 , 510–512 View Figs 504–521 cf. Figs 38 View Figs 36–44 , 491–493 View Figs 487–503 ).

Etymology

The specific epithet is a toponym referring to the range of this species, confined to the historical area Karategin (also Karategen), an eastern province of the Bukhara Emirate in the 19 th century, which included the Karategin Mts and Peter I Mts.

Material examined

Holotype

TAJIKISTAN • ♀; Peter I Mts (southern slope), Harvikush Canyon , 35 km ENE of Tavildara Village; 38°52′ N, 70°49′ E; 1900–2200 m a.s.l.; 10 Jul. 2019; S. Zonstein and A. Hakimov leg.; SMNH. GoogleMaps

Paratypes (10 ♀♀) TAJIKISTAN • 4 ♀♀; same collection data as for preceding; SMNH • 3 ♀♀; same collection data as for preceding; 9 Jul. 1978; V. I. Ovcharenko leg.; SMNH • 3 ♀♀; same collection data as for preceding; ZISP .

Description

Female (holotype)

HABITUS. See Fig. 44. View Figs 36–44

MEASUREMENTS. TBL 20.95, CL 7.46, CW 6.10, LL 0.59, LW 1.37, SL 3.88, SW 3.45.

COLOUR. Carapace dull reddish brown with clypeus and eye tubercle even darker brown, eyes encircled with partially fused wide blackish brown rings; chelicerae dark cherry red brown; sternum, labium, maxillae, epigastrum, book-lungs and spinnerets light yellowish brown; palps and legs light to medium brownish orange; abdomen medium brown, dorsally with several small light brownish spots and diffuse dark brown chevron-like pattern.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 119 View Figs 118–126 . Clypeus and eye group as in Fig. 176 View Figs 172–183 . Eye diameters and interdistances: AME 0.16(0.20), ALE 0.28, PLE 0.28, PME 0.19; AME–AME 0.21(0.17), ALE–AME 0.20(0.18), ALE–PLE 0.22, PLE–PME 0.09, PME–PME 0.55. Weak rastellum composed of 30–40 slightly thickened spikes on anterior cheliceral edge. Each cheliceral furrow with 9 promarginal teeth and 6–7 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 238 View Figs 238–246 . Maxillae with 26–29 cuspules each.

LEGS. Tibia and metatarsus IV densely covered with long modified hairs, as shown in Fig. 316 View Figs 310–318 . Scopula: entire and distal on metatarsi I–II; entire on palpal tarsus and tarsi I–II; sparse and widely divided by setae on tarsi III–IV. Trichobothria: 2 rows of 8–9 each on tibiae, 15–17 on metatarsi, 17–20 on tarsi. Palpal claw with 4 promarginal teeth. PTC I–IV with 6–7 teeth on each margin.

SPINATION. All femora with one basodorsal spine and 3–4 median and/or apical bristles; tarsi I–IV aspinose. Palp: femur pd1; patella p1; tibia p1(0), v6; tarsus v2. Leg I: femur pd2; patella p1; tibia p1(0), v4; metatarsus v4. Leg II: femur pd3; patella p1; tibia p2, v6; metatarsus v7(6). Leg III: femur pd4(3), rd3; patella p3, r2(1); tibia d1, p2, r3, v7; metatarsus p2, r3, v7. Leg IV: femur rd3(2); patella r1; tibia r3, v7; metatarsus p2, r4, v9.

SPERMATHECAE. Each of paired spermathecae U-shaped with a relatively low and wide base carrying two more or less broadly spaced and unevenly shaped branches: a longer and wider trapezoidal inner branch and a more slender, shorter and club-like outer one ( Fig. 510 View Figs 504–521 ).

SPINNERETS. See Fig. 576 View Figs 575–583 . PMS: length 0.89, diameter 0.36. PLS: length of basal, medial and apical segments 1.55, 0.73, 0.42; total length 2.70; apical segment triangular.

Male

Unknown.

Variation

Carapace length in paratype females (n=7) varies from 6.71 to 7.88. All the examined females are habitually very similar to each other. Variations in the structure of the eye group, tibia and metatarsus IV, and the spermathecae as shown in Figs 177 View Figs 172–183 , 317 View Figs 310–318 , 511–512 View Figs 504–521 .

Ecology

The spiders were found in the midland zone at an altitude of 1900–2000 m a.s.l. inhabiting open woodland dominated by Juniperus seravschanica . Females of Raveniola karategensis sp. nov. live in open burrows of 30–40 cm depth, provided with a weakly silk-lined entrance rim, walls and living chamber ( Figs 659–666 View Figs 659–666 , 675, 678–679 View Figs 675–682 ).

Distribution

Tajikistan: Peter I Mts. See Fig. 753 View Figs 751–760 .

SMNH	Department of Paleozoology, Swedish Museum of Natural History
V	Royal British Columbia Museum - Herbarium
ZISP	Zoological Institute, Russian Academy of Sciences