Raveniola dolosa, Zonstein, 2024

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 29-31

publication ID	https://doi.org/ 10.5852/ejt.2024.967.2699
publication LSID	lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738
DOI	https://doi.org/10.5281/zenodo.14026330
persistent identifier	https://treatment.plazi.org/id/7EC137C9-9E8D-4B2C-8309-358072F8FF08
taxon LSID	lsid:zoobank.org:act:7EC137C9-9E8D-4B2C-8309-358072F8FF08
treatment provided by	Plazi
scientific name	Raveniola dolosa
status	sp. nov.

Treatment

Raveniola dolosa sp. nov.

urn:lsid:zoobank.org:act:7EC137C9-9E8D-4B2C-8309-358072F8FF08

Figs 7 View Figs 1–9 , 41 View Figs 36–44 , 71 View Figs 69–75 , 89 View Figs 82–90 , 116 View Figs 109–117 , 143 View Figs 136–147 , 172 View Figs 172–183 , 208 View Figs 202–210 , 235 View Figs 229–237 , 265 View Figs 265–273 , 295 View Figs 290–309 , 356 View Figs 349–363 , 379–399 View Figs 379–388 View Figs 389–399 , 500–503 View Figs 487–503 , 568–570 View Figs 565–574 , 643–650, 751

Diagnosis

In the structure of the embolus and the spermathecae, Raveniola dolosa sp. nov. resembles R. ignobilis sp. nov.; it can be distinguished from the latter in having the embolus lacking a raised keel and in possessing noticeably shorter spermathecae (vs a clearly keeled embolus and longer branches of the spermathecae in R. ignobilis ; Figs 397–399 View Figs 389–399 , 500–503 View Figs 487–503 cf. Figs 403–408 View Figs 400–408 , 509 View Figs 504–521 ). Additionally, R. dolosa differs from R. ignobilis in having the posterior medium spinnerets strictly reduced in size (vs considerably larger PMS in the latter species; Figs 568–570 View Figs 565–574 cf. Figs 573–574 View Figs 565–574 ).

Etymology

The specific epithet is a Latin adjective ‘ dolosus/-a/-um ’ (= deceptive) referring to a general similarity of this new species to the closest congeners, Raveniola ignobilis sp. nov. and R. sororcula sp. nov.; the gender is feminine.

Material examined

Holotype

TAJIKISTAN • ♂; Peter I Mts , Childara Canyon , Shahobdara Gorge , 4 km NNW of Shahob Village; 38°51′ N, 70°18′ E; 1900 m a.s.l.; 12 Jul. 1988; S. Zonstein leg.; SMNH. GoogleMaps

Paratypes (13 ♀♀)

TAJIKISTAN • 11 ♀♀; same locality as for holotype; 1900–2100 m a.s.l.; 12–13 Jul. 1988; S. Zonstein and S.V. Ovchinnikov leg.; SMNH GoogleMaps • 2 ♀♀; same locality as for holotype, 2.5 km N of Shahob Village ; 38°50′ N, 70°19′ E; 1800 m a.s.l.; 8 Jul. 2019; S. Zonstein leg.; SMNH GoogleMaps .

Description

Male (holotype)

HABITUS. See Fig. 7. View Figs 1–9

MEASUREMENTS. TBL 12.55, CL 5.24, CW 4.97, LL 0.50, LW 0.87, SL 2.51, SW 2.41.

COLOUR. Carapace, chelicerae and leg I dull reddish brown; palps and legs II–IV lighter reddish brown; eye tubercle dark brown; sternum, labium and maxillae light yellowish brown; abdomen tan brown, with dark brown dorsal chevron-like pattern and a few small brown marks on ventral side; book-lungs and spinnerets pale yellowish brown.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 89 View Figs 82–90 . Clypeus and eye group as in Fig. 143 View Figs 136–147 . Eye diameters and interdistances: AME 0.14(0.20), ALE 0.26, PLE 0.16, PME 0.12; AME–AME 0.10(0.05), ALE–AME 0.11(0.08), ALE–PLE 0.10, PLE–PME 0.06, PME–PME 0.37. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 9 promarginal teeth and 3–4 mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in Fig. 208 View Figs 202–210 . Maxillae with 36–39 cuspules each.

LEGS. Tibia and metatarsus I as in Figs 265 View Figs 265–273 , 295 View Figs 290–309 . Scopula: entire and distal on metatarsi I–II; entire on tarsi I–II; narrowly divided by setae on tarsus III; widely divided on tarsus IV. Trichobothria: 2 rows of 8–9 each on tibiae, 12–14 on metatarsi, 10–11 on tarsi, 7–8 on cymbium. PTC I–IV with 9–10 and 7–8 teeth on inner and outer margins, respectively.

SPINATION. All femora with 1–2 basodorsal spines and 3–4 dorsal bristles; metatarsus I and tarsi I–IV aspinose. Palp: femur pd2, rd1; patella pd1; tibia d1, p2, r3, v5; cymbium d4(5)+12–15 spikes. Leg I: femur pd3, rd3; patella p1; tibia p2, pv1, r2, rv2+ 2M. Leg II: femur pd3; patella p2; tibia p3, v7; metatarsus v5(4). Leg III: femur pd3, rd2; patella p2, r1; tibia d2(1), p4, r3, v7; metatarsus d3, p4, r3, v8. Leg IV: femur pd3(2), rd2(1); patella p1, r1; tibia d1, p3, r3, v9(7); metatarsus d3, p4(3), r5(4), v9.

PALP. Tibia, cymbium and copulatory bulb as shown in Fig. 356 View Figs 349–363 . Embolus with moderately long basal portion provided with very low keel and short, hooked apical part ( Figs 397–399 View Figs 389–399 ).

SPINNERETS. See Fig. 568 View Figs 565–574 . PMS: length 0.36, diameter 0.16. PLS: maximal diameter 0.49; length of basal, medial and apical segments 0.75, 0.43, 0.38; total length 1.56; apical segment triangular.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	3.13 (2.79)	1.61 (1.58)	2.49 (1.89)	–	0.83 (1.92)	8.06 (8.18)
Leg I	5.65 (3.70)	2.94 (2.43)	4.32 (2.88)	4.07 (2.21)	2.43 (1.71)	19.41 (12.93)
Leg II	5.26 (3.36)	2.58 (2.16)	3.94 (2.56)	3.93 (2.20)	2.29 (1.79)	18.02 (12.07)
Leg III	4.35 (3.09)	2.07 (1.81)	2.96 (2.04)	4.14 (2.72)	2.14 (1.87)	15.66 (11.53)
Leg IV	5.46 (3.98)	2.46 (2.18)	4.08 (3.31)	5.82 (3.66)	2.78 (2.04)	20.60 (15.17)

Female (paratype)

HABITUS. See Fig. 41. View Figs 36–44

MEASUREMENTS. TBL 15.60, CL 5.26, CW 4.53, LL 0.87, LW 1.11, SL 2.76, SW 2.48.

COLOUR. Similar to that of male, but carapace and legs slightly paler.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 116 View Figs 109–117 . Clypeus and eye group as in Fig. 172 View Figs 172–183 . Eye diameters and interdistances: AME 0.13(0.18), ALE 0.25, PLE 0.18, PME 0.16; AME–AME 0.12(0.07), ALE–AME 0.10(0.07), ALE–PLE 0.09, PLE–PME 0.05, PME–PME 0.35. Cheliceral rastellum absent. Each cheliceral furrow with 9 promarginal teeth and 9–10 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 235 View Figs 229–237 . Maxillae with ca 70 cuspules each.

LEGS. Scopula: distal on metatarsi I–II; entire on palpal tarsus and tarsus I; widely divided by setae on tarsus II; rudimentary on tarsi III–IV. Trichobothria: 2 rows of 11–12 each on tibiae, 16–18 on metatarsi; 13–15 on tarsi; 11 on palpal tarsus. Palpal claw with 5 promarginal teeth. PTC I–II and III–IV with 4–5 and 5–7 teeth on each margin, respectively.

SPINATION. All femora with one basodorsal spine and 3 dorsal bristles; palpal patella, patellae I–II, and tarsi I–IV aspinose. Palp: femur pd1; tibia p1, v7; tarsus v3. Leg I: femur pd1; tibia v6; metatarsus v6. Leg II: femur pd1; tibia p2, v6; metatarsus v6. Leg III: femur pd2, rd2; patella p2, r1; tibia d1, p2, r2(1), v7; metatarsus d3(2), p3, r3, v9(8). Leg IV: femur rd1; patella r1; tibia p2, r2, v7; metatarsus p3, r3, v8(7).

SPERMATHECAE. Each of paired spermathecae V-shaped with relatively low and wide base carrying two densely located, short and weakly diverging branches ( Fig. 500 View Figs 487–503 ).

SPINNERETS. See Figs 569–570 View Figs 565–574 . PMS: length 0.47, diameter 0.15. PLS: maximal diameter 0.57; length of basal, medial and apical segments 0.98, 0.42, 0.43; total length 1.83; apical segment triangular.

Variation

Carapace length varies in females (n =9) from 4.30 to 6.17. Variations in the habitus and structure of the spermathecae as shown in Figs 71 View Figs 69–75 , 501–503 View Figs 487–503 .

Ecology

All spiders were found hiding in soil cavities under stones in riverside woodlands, dominated by Juniperus seravschanica , Juglans regia and Populus sp. ( Figs 643–650 View Figs 643–650 ).

Distribution

Known only from the type locality. See Fig. 751 View Figs 751–760 .