Quercus palmeri (Engelm.) Engelm., Bot.

Bartholomew, Bruce & Almeda, Frank, 2023, Nomenclator botanicus of Fagaceae in Latin America, Phytotaxa 580 (1), pp. 1921-1935 : 1921-1935

Quercus palmeri (Engelm.) Engelm., Bot. California 2: 97 (1880).

Quercus chrysolepis Liebm. subsp. palmeri Engelm., Trans. Acad. Sci. St. Louis 3: 393. (1877). Type :— U.S.A., California: E. Palmer 30 (Holotype: MO-204133!; isotypes: A!, GH!) .

Quercus dunnii Kellogg , Pacific Rural Press 17(23): 371 (1879). This is an invalid name because Kellogg cited it as a provisional name.

Distribution: Mexico (Baja California, Baja California Sur, Chihuahua, Sonora), U.S.A. (Arizona, California).

Quercus peduncularis Neé, Anales Ci. Nat. 3: 270 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA26474 ! designated by Valencia-A. & Coombes (2020, p. 87); isolectotypes: MA!, UCSB (fragment p.p. as to smaller leaf from MA lectotype)!, UCSB (fragment)!).

Quercus affinis M.Martens & Galeotti non Scheidw. (1837), Bull. Acad. Roy. Sci. Bruxelles 10(1): 222 (1843). See Q. martensiana below.

Quercus barbanthera Trel., Mem. Natl. Acad. Sci. 20: 68, pl. 81 (1924). Type :— GUATEMALA: G. V. Skinner s. n. (1845) (Holotype at B destroyed. Lectotype (designated here): K000648587 !). Valencia-A. & Coombes (2020, p. 87) invalidly lectotypified the name based on Trelease’s image of the holotype at B which was destroyed. A paratype exists at K and according to Article 9.12 of the Code “If no isotype, syntype or isosyntype is extant, the lectotype must be chosen from among the paratypes if such exist” (Turland et al. 2018). The epitype, Miranda 6429 ( MEXU10381 as image!) which they selected is also invalid, because of the existence of the paratype which is now the lectotype.

Quercus barbanthera var. calva Trel., Mem. Natl. Acad. Sci. 20: 68 (1924). Type:— GUATEMALA, Chimaltenango: E. T. Heyde & E. Lux 3152 (Lectotype (designated here): K000512621 !; isolectotypes: NY!, US!). Trelease (1924. p. 68) cited two different collections in the protologue.

Quercus callosa Benth., Pl. Hartw. 91 (1842). Type:— GUATEMALA, Guatemala: T. Hartweg 616 (Lectotype: K000512622 ! designated by Trelease (1824, p. 218, pl. 82a); isolectotypes: CGE!, K!, LD (image)!, NY!). There are two sheets of Hartweg 616 in the Bentham Herbarium at K, one of which was effectively lectotypified by Trelease.

Quercus comasaguana Trel., Mem. Natl. Acad. Sci. 20: 43 (1924). Type:— EL SALVADOR, La Libertad: S. Calderon 1430 (Lectotype (designated here): ILL00007525 !; isolectotypes: GH!, US!). There is no indication in the protologue as to where the holotype was deposited, and both the ILL and US specimens are labeled as a type.

Quercus dolichopus E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 87 (1939). Type:— MEXICO, México: G. B. Hinton 6378 (Lectotype: K000575005 ! designated by Valencia-A. & Coombes (2020, p. 87); isolectotypes: BM!, F!, G!, GH!, K!, MEXU!, MO!, NY!, UCSB (fragment)!, US!) .

Quercus liebmannii Oerst. ex Trel. f. brevipes Trel., Mem. Natl. Acad. Sci. 20: 67, pl. 75b (1924). Type:— MEXICO, Oaxaca: F. M. Liebmann 3526 (Holotype: C not located; isotypes: C!, CAS!). The holotype of Q. liebmannii f. brevipes at C is missing although isotypes exist at C and CAS. A photo of part of the holotype is in Trelease (1924) pl. 75b which is Liebmann 3526 (Cupulif. Lbm. 136); hopefully the specimen with the correct Cupulif. Lbm. number and matching the photo will eventually be found. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. peduncularis as broadly interpreted.

Quercus martensiana Trel., Contr. U.S. Natl. Herb. 23(2): 182 (1922). Type:— MEXICO, Veracruz: H. Galeotti 90 (Holotype: BR0000009816213 !; isotypes: G!, K!, NY!, P!, UCSB (fragment)!). Nom. nov. for Q. affinis M. Martens & Galeotti (1843) non Scheidw. (1837).

Quercus martensiana f. perplexans Trel., Mem. Natl. Acad. Sci. 20: 60 (1924). Type :— MEXICO, Veracruz: F. M. Liebmann 3567 (Holotype: C 10012502!) .

Quercus peduncularis subsp. callosa (Benth.) A.Camus, Chênes Texte 2: 599 (1939).

Quercus peduncularis subsp. pedicularis var. callosa (Benth.) A.Camus, Chênes Atlas 2: 91 (1935).

Quercus peduncularis subsp. pilicaulis (Trel.) A.Camus, Chênes Atlas 2: 91 (1935).

Quercus peduncularis subsp. pilicaulis var. macrodonta (Trel.) A.Camus, Chênes Atlas 2: 91 (1935).

Quercus peduncularis subsp. pilicaulis var. obovalis (Trel.) A.Camus, Chênes Atlas 2: 91 (1935).

Quercus pilicaulis Trel., Mem. Natl. Acad. Sci. 20: 67, 77b (1924). Nom. nov. for Q. tomentosa var. abbreviata A. DC (1864) non Q. abreviata Vuk. (1889) .

Quercus pilicaulis f. macrodonta Trel., Mem. Natl. Acad. Sci. 20: 68, pl. 78 (1924). Type:— GUATEMALA: J. R. R. Warszewicz 15 (Holotype at B destroyed. Lectotype (designated here): G00390238!; isolectotype; UCSB (fragment from G lectotype)!). Trelease (1924, p. 218) in the caption to pl. 78a designated Warscewicz 15 at B as the holotype of Q. pilicaulis f. macrodonta and treated Q. tomentosa var. bullata A.DC. as a synonym for which Warsczwicz 15 at G was a syntype and which he effectively lectotypified in the caption to pl. 78b. We are lectotypifying Q. plicaulis f. macrodonta based on the specimen of Warsczwicz 15 at G.

Quercus pilicaulis f. obovalis Trel.,Mem.Natl.Acad.Sci. 20:68,pl.79b(1924). Type :— GUATEMALA, Chimaltenango: F. C. Lehmann 1725 (Holotype at B destroyed. Lectotype: UCSB000351 (fragment)! designated by Valencia-A. & Coombes (2020, p. 87); isolectotypes: BM!, G). Lectotypification of this name was based on a fragment at UCSB rather than the isotype at G from which the fragment was reportedly taken or the other isotype at BM.

Quercus schultzei Trel., Repert. Spec. Nov. Regni Veg. 33: 316 (1934). Type:— MEXICO, Guerrero: L. Schultze 115 (Holotype at B destroyed. Lectotype (designated here): ILL00007589 ! (fragment)).

Quercus splendens Neé, Anales Ci. Nat. 3: 275 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA25972 ! designated by Valencia-A. & Coombes (2020, p. 87); isolectotypes: MA!, UCSB (fragment)!).

Quercus tomentosa Willd. nom. illeg., Sp. Pl., ed. 4(1): 437 (1805). This is a superfluous name because Willdenow cites Q. peduncularis Neé (1801) , which is an older name, in synonomy.

Quercus tomentosa Willd. nom. illeg. var. abbreviata A.DC., Prodr. 16(2.1): 33 (1864). Type :— GUATEMALA: J. R. R. Warszewicz 43 (Holotype: G00421035 !; isotype: UCSB (fragment from G holotype)!). Although Valencia- A. & Coombes (2020, p. 87) lectotypified this name, it was not necessary as A. de Candolle clearly specified the holotype.

Quercus tomentosa Willd. nom. illeg. var. bullata A.DC., Prodr. 16(2.1): 33 (1864). Type :— GUATEMALA: J. R. R. Warszewicz 15 (Lectotype: G00390238 ! designated by Trelease (1924, p. 218, pl.78b); isolectotype: UCSB (fragment from the G lectotype)!). A. de Candolle (1864, p. 33) cited two syntypes.

Quercus tomentosa Willd. nom. illeg. var. communis A.DC., Prodr. 16(2.1): 33 (1864). Type :— GUATEMALA: J. Warszewicz 17 (Lectotype: G00390235 ! designated by Valencia-A. & Combes (2020, p. 87)). A. de Candolle (1864, p. 33) cites several syntypes.

Distribution: El Salvador, Guatemala, Honduras, Mexico (Aguascalientes, Chiapas, Ciudad de México, Colima,

Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México , Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Querétaro,

San Luis Potosí, Sinaloa, Tamaulipas, Veracruz, Zacatecas).

Quercus peninsularis Trel., Mem. Natl. Acad. Sci. 20: 124, pl. 225a (1924). Type:— MEXICO, Baja California: T. Brandegee s. n. (23 May 1893) (Holotype: A00062253 !; isotypes: UCSB (fragment)!, US) .

Distribution: Mexico (Baja California).

Quercus perseifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854) . Type:— MEXICO, Veracruz: F. M. Liebmann 3476 (Lectotype: CAS0006172 ! designated by Trelease (1924, p. 225, pl. 292a); isolectotypes: C!, UCSB (fragment from the CAS lectotype)!). The lectotype is sheet Cupulif. Liebm. 98. Trelease (1924, p. 149) described the fruit as annual although most collections we have seen are biennial. It is known that some species of Quercus have both annual and biennial fruit, not only within the same species, but even within the same population (Pons & Pausas 2012).

Quercus perseifolia f. microcarpa Trel., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854) . Type:— MEXICO, Veracruz: H. Galeotti 112 (Lectotype (designated here): BR!). Trelease cited two specimens without designating which one was the type.

Distribution: Mexico (Hidalgo, México, Michoacán, Nuevo León, Oaxaca, Puebla, San Luis Potosí, Tamaulipas,

Veracruz).

Quercus pinnativenulosa C.H.Mull., J. Arnold Arbor. 17(3): 171 (1936). Type :— MEXICO, Nuevo León : C. H. Muller & M. T. Muller 1342 (Holotype: A00055690 !; isotypes: CAS!, F!, MEXU!, TEX (image)!). This species may be the same as Q. perseifolia in the broad sense. Additional study is needed.

Distribution: Mexico (Coahuila, Hidalgo, Nuevo León, Oaxaca, Puebla, Qurerétaro, San Luis Potosí, Tamaulipas,

Veracruz).

Quercus planipocula Trel., Mem. Natl. Acad. Sci. 20: 136, pl. 259a (1924). Type :— MEXICO, Nayarit: J. N. Rose 1970 (Holotype: US00089613!; isotypes: GH!, MO!, NY!, UCSB (fragment)!).

Distribution: Mexico (Colima, Guerrero, Jalisco, México, Michoacán, Nayarit, Oaxaca, San Luis Potosí, Sinaloa,

Tamaulipas). The occurrence of Q. planipocula in Oaxaca, San Luis Potosí, and Tamaulipas reported by Villaseñor

(2016) needs to be confirmed.

Quercus polymorpha Schltdl. & Cham., Linnaea 5(1): 78 (1830). Type :— MEXICO, Veracruz: C. J. W. Schiede 20 (Holotype at B destroyed. Lectotype (designated here): HAL0060344 !; isolectotypes: BR!, G!) .

Quercus germana Schltdl. & Cham. var. lemmonii Trel., Mem. Natl. Acad. Sci. 20: 52 (1924). Type :— MEXICO, Veracruz: J. G. Lemmon & J. G . Lemmon 103 (Holotype: US 00089539!; isotypes: E!, F!, GH!, MO!, NY!, UC!) .

Quercus guatimalensis A.DC, Prodr. 16(2.1): 78 (1864). Type :— GUATEMALA: J. R. R. Warszewicz 37 (Holotype at B destroyed. Lectotype (designated here): P00754147 !). Nom. nov. for Q. turbinata Liebm. (1854) non Blume (1825).

Quercus juergensenii Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854) . Type:— MEXICO, Puebla: F. M. Liebmann s. n. (Holotype: C (missing); isotype: UCSB (fragment from C holotype)!). Although the foliage is described by Liebmann, the only remaining specimens at C when Trelease visited there were acorns. These acorns are now missing at C and the only extant ones are a few now at UCSB. Trelease also considered this be a problematic species but perhaps synonymous with Q. polymorpha , which is a name that has priority.

Quercus petiolaris Benth., Pl. Hartw. 55 (1840). Type:— MEXICO, Jalisco: T. Hartweg 420 (Holotype: K000648582 !; isotypes: A!, BM!, CAS!, CGE!, E!, FI!, G!, LD (image)!, LE!, NY!, P!, UCSB (fragment)!).

Quercus polymorpha f. angustifolia C.H.Mull., J. Arnold Arbor. 17(3): 161 (1936). Type :— MEXICO, Nuevo León: C. H. Muller & M. T . Muller 1144 (Holotype: A00283982 !) .

Quercus turbinata Liebm. non Blume (1825), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 186 (1854). See Q. guatimalensis above.

Quercus varians M.Martens & Galeotti, Bull.Acad. Roy. Sci. Bruxelles 10(1): 214 (1843). Type:— MEXICO, Veracruz: H. Galeotti 92 (Lectotype: BR0000006916251 ! designated by Trelease (1924, p. 217, pl. 37); isolectotypes: BR!, G!, K!, L!, LE!, P!, UCSB (fragment)!). M. Martens & Galeotti (1843, p. 214) cited only one collection number, four sheets of which are at BR; one of these was designated by Trelease as “The type.”

Distribution: Guatemala, Honduras, Mexico (Chiapas, Coahuila, Guanajuato, Hidalgo, Jalisco, México, Michoacán,

Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Tamaulipas, Veracruz).

Quercus porphyrogenita Trel., Mem. Natl. Acad. Sci. 20: 51, pl. 39a (1924). Type:— MEXICO, Nuevo León: W. Trelease 117 (Holotype: MO-144179!; isotype: ILL!). The distinction between Q. porphyrogenita and Q. laceyi needs additional study.

Quercus microlepis Trel. & C.H.Mull. non Vuk. (1883), Bull. Torrey Bot. Club 63(3): 150 (1936). Type:— MEXICO, Nuevo León: C.H. Muller 578 (Holotype: ILL00007500 !; isotypes: MEXU (a image!), UCSB (fragment)!).

Distribution: Mexico (Nuevo León).

Quercus potosina Trel., Mem. Natl. Acad. Sci. 20: 84, pl. 123 (1924). Type :— MEXICO, San Luis Potosí: C. C. Parry & E . Palmer 838 (Holotype: MO-143894!; isotypes: BM!, GH!, K!, MO!, NY!, P!, PH!, UCSB (fragment)!, US!, WIS!) .

Quercus jaralensis Trel., Mem. Natl. Acad. Sci. 20: 86, pl. 133 (1924). Type :— MEXICO, Guanajuato: W. Schumann 1316 (Holotype at B destroyed. Lectotype (designated here): P00754117 !; isolectotype: P!) .

Quercus potosina f. aperta Trel., Mem. Natl. Acad. Sci. 20: 84, pl. 125a (1924). Type:— MEXICO, San Luis Potosí: C. C. Parry & E. Palmer 838b (Holotype: MO-143893!).

Quercus potosina f. exilis Trel., Mem. Natl. Acad. Sci. 20: 84, pl. 124a (1924). Type :— MEXICO, San Luis Potosí: C. C. Parry & E . Palmer 838 (Holotype: MO-143892!; isotypes: BM!, GH!, K!, MO!, NY!, P!, PH!, UCSB (fragment)!, US!, WIS!) .

Distribution: Mexico (Aguascalientes, Chihuahua, Coahuila, Colima, Durango, Guanajuato, Hidalgo, Jalisco, México,

Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Veracruz, Zacatecas). The occurrence of Q.

potosina in México and Veracruz reported by Villaseñor (2016) needs to be confirmed.

Quercus praeco Trel., Mem. Natl. Acad. Sci. 20: 88, pl. 139 (1924). Type:— MEXICO, Jalisco: J. N. Rose 2590 (Holotype: US00089611!; isotypes: GH!, NY!, UCSB (fragment)!). This species is close to Q. xylina , and additional study is needed to determine if it is distinct.

Distribution: Mexico (Jalisco, Nayarit, Zacatecas).

Quercus pringlei Seemen, Bot. Jahrb. Syst. 29(1): 96 (1900). Type:— MEXICO, Coahuila: C. G. Pringle 2382 (Holotype at B destroyed. Lectotype (designated here): US00089609!; isolectotypes: A!, AC!, BM!, BR!, CAS!, COLO (image)!, E!, F!, G!, GH!, K!, KFTA (image)!, LL (image)!, MEL (image)!, MEXU!, MO!, NDG (image)!, NY!, P!, PH!, PUL (image)!, UC!, UCSB (fragment)!, US!, VT!, YU (image)!). Trelease (1924, p. 124) in the caption to pl. 148a designated Pringle 2382 at B as “The type” which was destroyed.

Quercus filiformis C.H.Mull., Amer. Midl. Naturalist 27(2): 473 (1942). Type:— MEXICO, Coahuila: C. H. Muller 3150 (Holotype: US00089513!; isotypes: GH!, UC!, UCSB (fragment)!). This name was also used in 1818 as a nom. nud. and as nom. inval. as a synonym of Q. bicolor var. platanoides .

Quercus opaca Trel., Mem. Natl. Acad. Sci. 20: 92, pl. 148b (1924). Type:— MEXICO, Hidalgo: J. N. Rose 9030 (Holotype: US00089571!; isotypes: GH!, NY!, UCSB (fragment)!).

Quercus pringlei f. dentata C.H.Mull., J. Arnold Arbor. 17(3): 168. (1936). Type :— MEXICO, Nuevo León: C. H. Muller 881 (Holotype: A00969953 !) .

Distribution: Mexico (Chihuahua, Coahuila, Durango, Hidalgo, Nuevo León, Querétaro, San Luis Potosí, Tamaulipas,

Veracruz, Zacatecas).

Quercus pungens Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171 (1854) . Type:— U.S.A., Texas: C. Wright 664 (Lectotype: NY00248762 (image)! designated by Trelease (1924, p. 221, pl. 157a); isolectotypes: BM!, GH!, K (image)!, NY!, P!, UCSB (fragment)!, US!, W!). The Hooker Herbarium collection of Wright 664 at K, which was a syntype of this species, was sent to NY, and this is the specimen Trelease effectively lectotypified. The only specimen of Wright 664 currently at K is the Bentham Herbarium specimen which becomes a lectotype. The other syntype is Lobb 106 from the Hooker Herbarium at K ( K000648585 !).

Quercus pungens var. vaseyana (Buckley) C.H.Mull., Cont. Texas Res. Found., Bot. Stud. 1:70 (1951).

Quercus undulata Torr. var. pungens (Liebm.) Engelm., Trans. Acad. Sci. St. Louis 3: 392 (1877).

Quercus undulata var. vaseyana (Buckley) Rydb., Bull. New York Bot. Gard. 2(6): 218 (1901).

Quercus vaseyana Buckley, Bull. Torrey Bot. Club 10(8): 91 (1883). Type:— U.S.A., Texas: S. B. Buckley s. n. (Lectotype: NY00253631 ! designated by Trelease (1924, p. 221, pl. 156a) and redundantly relectotypified by Dorr & Nixon (1985, p. 223); isolectotype: NY!) .

Distribution: Mexico (Chihuahua, Coahuila, Durango, Guanajuato, Hidalgo, Nuevo León, Querétaro, San Luis Potosí,

Sonora, Tamaulipas, Veracruz), U.S.A. (Arizona, New Mexico, Texas) .

Quercus purulhana Trel., Mem. Natl.Acad. Sci. 20: 63, pl. 64 (1924). Type:— GUATEMALA, Baja Verapaz: H. Pittier 163 (Holotype: US 00089604!; isotypes: NY!, UCSB (fragment)!, US!) .

Quercus aguana Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 9(4): 281 (1940). Type:— HONDURAS, Lempira: T. G. Yuncker et al. 8180 (Holotype: ILL00007476 !; isotypes: BM!, F!, G!, GH!, K!, MICH!, MO!, NY!, S (image)!, US!) .

Quercus barbeyana Trel., Mem. Natl. Acad. Sci. 20: 67, pl. 76 (1924). Type :— GUATEMALA, Chiquimula: F. C. Lehmann 1711 (Holotype: G00358171 !; isotype: BM!) .

Distribution: Belize, Guatemala, Honduras, Mexico (Chiapas), Nicaragua.

Quercus radiata Trel., Proc. Amer. Philos. Soc. 60: 33, pl. 3 (1921). Type :— MEXICO, Nayarit: J. N. Rose 2230 (Holotype: US 00089601!; isotypes: GH!, NY!) .

Distribution: Mexico (Durango, Jalisco, Nayarit, Zacatecas).

Quercus rekonis Trel., Mem. Natl. Acad. Sci. 20: 60, pl. 57 (1924). Type:— MEXICO, Oaxaca: C. Conzatti et al. 3121 (Holotype: US00089631!; isotypes: MEXU!, MO!). The collectors of the type were C. Conzatti, B.P. Reko & Makrinius, but only the the name B.P. Reko is on the US holotype.

Distribution: Mexico (Chihuahua, Durango, Guerrero, Jalisco, México, Michoacán, Nayarit, Oaxaca, Sinaloa).

Quercus resinosa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 182 (1854) . Type :— MEXICO: B. Seemann 1972 (Holotype: K000512615 !; isotypes: GH!, K!, UCSB (fragment)!). There are two specimens of Seemann 1972 in the Hooker Herbarium at K, but only one of them has a determination in Liebmann’s hand which we consider to be the holotype.

Distribution: Mexico (Aguascalientes, Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán,

Nayarit, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Zacatecas). The occurrence of Q.

resinosa in Guerrero and Hidalgo reported by Villaseñor (2016) needs to be confirmed.

Quercus rugosa Neé, Anales Ci. Nat. 3: 275 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA26471 ! designated by Bartholomew & Almeda (2022, p. 29); isolectotypes: MA!, UCSB (fragment from MA lectotype)!, UCSB (fragment)!).

Quercus conglomerata Trel. non Pers. (1807), Mem. Natl. Acad. Sci. 20: 77, pl. 106a (1924). Type:— MEXICO, Michoacán: T. Hartweg 429 (Holotype: B destroyed; isotypes: BM!, E!, G!, K!, L!, NY!, P!, UCSB (fragment)!). Because Q. conglomerata Trel. is a later homonym, we are not lectotypifying the name.

Quercus decipiens M.Martens & Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 214 (1843). Type:— MEXICO, Veracruz: H. Galeotti 131 (p.p.) (Lectotype: BR0000006914295! designated by Trelease (1924, p. 219, pl. 107); isolectotypes: BR!, G!, P!, UCSB (fragment)!). M.Martens & Galeotti cite only one collection, Galeotti 131, four sheets of which are at BR. Two of them are types of Q. decipiens , one (BR000006915612) is a mixed sheet in which only the upper left specimen on the sheet is a type of Q. decipiens , and the other one (BR0000006913953) is not a type of Q. decipiens . The lectotype is the sheet designated by Trelease as “The type.”

Quercus diversicolor Trel., Mem. Natl. Acad. Sci. 20: 73, pl. 92a (1924). Type :— MEXICO, Chihuahua: C. G. Pringle 2021 (Holotype at B destroyed. Lectotype (designated here): US 01013683!; isolectotypes: BM!, BR!, CAS!, G!, GH!, K!, M!, MEL (image) !, MO!, NDG (image) !, NY!, P!, PH!, UC!, UCSB (fragment) !, US!, VT !).

Quercus diversicolor var. mearnsii Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 94 (1924). Type:— U.S.A., Arizona: E. A. Mearns 2230 (Holotype: NY01465003 !; isotype: US!) .

Quercus diversicolor var. socorronis Trel., Mem. Natl. Acad. Sci. 20: 74 (1924). Type:— U.S.A., New Mexico: O. B. Metcalfe 669 (Lectotype (designated here): US 00089497!; isolectotypes: NDG (image)!, UCSB (fragment)!). Trelease (1924, p. 74) did not indicate where the type collections were deposited.

Quercus durangensis Trel., Mem. Natl. Acad. Sci. 20: 73, pl. 91a (1924). Type:— MEXICO, Durango: Mathews s. n. (Nov 1904) (Holotype: MO-143914!; isotype: UCSB (fragment from MO holotype)!).

Quercus macrophylla Neé var. rugosa (Neé) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 196 (1884).

Quercus peduncularis Neé subsp. hurteri (Trel.) A.Camus, Chênes Atlas 2: 92 (1935).

Quercus pilicaulis Trel. f. armata Trel., Mem. Natl. Acad. Sci. 20: 68, pl. 79b (1924). Type:— GUATEMALA, Quezaltenango: E. W. Nelson 3721 (Holotype: US 00089588!) .

Quercus pilicaulis f. concava C.H.Mull., Amer. Midl. Naturalist 18(5): 851 (1937). Type :— GUATEMALA, Chimaltenango: A. F. Skutch 685 (Holotype: US 00089587!; isotypes: A!, F!, ILL!, UC!) .

Quercus pilicaulis f. elongata C.H.Mull., Amer. Midl. Naturalist 18(5): 851 (1937). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 687 (Holotype: US00089586!; isotypes: A!, BR!, DS!, F!, ILL!, K!, UC!).

Quercus pilicaulis f. exserta C.H.Mull., Amer. Midl. Naturalist 18(5): 852 (1937). Type :— GUATEMALA, Chimaltenango: A. F. Skutch 661 (Holotype: US 00089585!; isotypes: A!, UC!) .

Quercus pilicaulis f. hurteri Trel., Mem. Natl.Acad. Sci. 20: 68, pl. 80 (1924). Type :— GUATEMALA, Quezaltenango: W. Trelease 37 (Holotype: ILL00007598 !) .

Quercus purpusii Trel., Mem. Natl. Acad. Sci. 20: 76, pl. 105 (1924). Type:— MEXICO, México: C. A. Purpus 1794 (Holotype: MO-143916!; isotypes: F!, GH!, NY!, UC!, UCSB (fragment)!, US!) .

Quercus reticulata Bonpl., Pl. Aequinoct. 2: 40, pl. 86 (1809). Type:— MEXICO, Guanajuato: A. Bonpland s. n. (Lectotype (designated here): P00129698!; isolectotypes: P!, UCSB (fragment)!). There are four Bonpland s. n. collections from Santa Rosa, Guanajuato, Mexico which are syntypes of Q. reticulata . The lectotype has the name “ Quercus coccolobaefolia sp. n. ” in Bonpland’s hand. Bonpland, however, used the name Q. reticulata in his publication. The syntypes of Q. reticulata are not the same collection as Bonpland 4269 which is the collection number for the type of Q. coccolobifolia Trel.

Quercus reticulata f. longa Trel., Mem. Natl.Acad. Sci. 20: 75, pl. 99a (1924). Type:— MEXICO, Hidalgo: C. Ehrenberg 897 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 99a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus reticulata subsp. reticulata var. conglomerata (Trel.) A.Camus, Chênes Atlas 2: 93 (1935).

Quercus reticulata subsp. reticulata var. squarrosa (Trel.) A.Camus, Chênes Atlas 2: 94 (1935).

Quercus reticulata subsp. rhodophlebia (Trel.) A.Camus, Chênes Atlas 2: 94 (1935).

Quercus reticulata subsp. rhodophlebia f. concava (Trel.) A.Camus, Chênes Texte 2: 581 (1939).

Quercus reticulata f. squarrosa Trel., Mem. Natl. Acad. Sci. 20: 75, pl. 99b (1924). Type:— MEXICO, Hidalgo: G. J. Graham 328 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 99b)!). Trelease (1924. p. 219) wrote in the caption to pl. 99b that the type was an Ehrenberg collection at B. He cited five of Ehrenberg’s collections earlier on p. 75. Because they were all destroyed, it is impossible to know which collection number (903, 904, 905, 906, or 908) was the holotype. The image in pl. 99b is probably of a collection made in Hidalgo, Mexico, because all the Ehrenberg collections around these numbers are from Hidalgo. Trelease also cited two Graham collections, of which we have found only Graham 328 which is a different species, so we are not lectotypifying the name based on either of the paratypes but rather on the image of the holotype in plate 99b.

Quercus rhodophlebia Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 95a (1924). Type:— MEXICO, Zacatecas: J. N. Rose 2806 (Holotype: US 00089626!) .

Quercus rhodophlebia f. concava Trel., Mem. Natl. Acad. Sci. 20: 74, pl. 95b (1924). Type:— MEXICO, Jalisco: J. N. Rose 2998 (Holotype: US00089623!; isotypes: GH!, MO!, NY!, UCSB (fragment)!).

Quercus sideroxyla Bonpl. f. aquifolia Trel., Mem. Natl. Acad. Sci. 20: 170 (1924). Type:— MEXICO, Guanajuato: A. Dugès 4 (Holotype: GH00969956 !). The specimen of Dugès 4 at US is Q. tridens so it cannot be considered an isotype.

Quercus suchiensis Warb., J. Roy. Hort. Soc. 58(1): 187, fig. 55 (1933). Type:— MEXICO, México / Michoacán: R. Heatley s. n. (Holotype: K000575026 ! and K000575027 !). The holotype was originally mounted on one sheet but now it is mounted on two sheets. The type collection is “on the boundaries of the states of Mexico and Michoacan.”

Quercus uhdeana Trel., Mem. Natl. Acad. Sci. 20: 75, pl. 100 (1924). Type :— MEXICO: Uhde 276 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 100)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Distribution: Guatemala, Honduras, Mexico (Aguascalientes, Baja California, Baja California Sur, Chiapas, Chihuahua,

Ciudad de México, Coahuila, Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos ,

Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz,

Zacatecas), U.S.A. (Arizona, New Mexico, Texas) .

Quercus rysophylla Weath., Proc. Amer. Acad. Arts 45: 423 (1910). Type:— MEXICO, Nuevo León: C. G. Pringle 10225 (Lectotype: GH00069922 ! designated by Trelease (1924, p. 225, pl. 279a); isolectotypes: ASU (image)!, CAS!, DAO (image)!, DUKE (image)!, ENCB (image)!, F!, IBUG (image)!, LE!, LL (image)!, MICH!, MSC (image)!, RSA (image)!, UC!, UCR (image)!, UCSB (fragment)!, US!, VT!, WIS!, YU (image)!).

Quercus tenuiloba f. hirsuta C.H.Mull., J.Arnold Arbor. 17(3): 178 (1936). Type:— MEXICO, Nuevo León: C. H. Muller 2122 (Holotype: A00055647 !; isotypes: MEXU!, MO!, P!, TEX (image)!, UCSB (fragment)!, US!). Although the species epithet is based on a probable hybrid collection (see Appendix 2), the type of this forma is Q. rysophylla .

Distribution: Mexico (Coahuila, Hidalgo, Nuevo León, Oaxaca, Querétaro, San Luis Potosí, Tamaulipas, Veracruz).

Quercus sagrana Nutt., N. Amer. Sylv. 1(1): 17 (1842). Type :— CUBA, Pinal del Rio: R. de la Sagra 418 (Holotype: PH00023400!; isotypes: G!, P (as s.n.)!, UCSB (as s.n.) (fragment)!).

Quercus cubana A.Rich., Hist. Fis. Cuba 11: 230 (1850). This is an illegitimate name because it is based on an isotype of Q. sagrana Nutt., R. de la Sagra [J. M. Valenzuela] s. n., which is at P as P00754080!. J.M. Valenzuela was a collector for R.de la Sagra and all of the type specimens of both Q. sagrana and Q. cubana were almost certainly collected by him. Richard (1850) cited J.M.Valenzuela as the collector of Q. cubana , but Valenzuela’s name is not on the any of the specimens distributed by de la Sagra.

Quercus oleoides Schltdl. & Cham. var. sagrana (Nutt.) C.H.Mull., Amer. Midl. Naturalist 65(1): 37. (1961).

Quercus oleoides subsp. sagrana (Nutt.) Borhidi, Acta Bot. Acad. Sci. Hung. 17: 6 (1971[1972]).

Quercus virginiana Mill. var. sagrana (Nutt.) Trel., Mem. Natl. Acad. Sci. 20: 113, pl. 189b (1924).

Distribution: Cuba.

Quercus salicifolia Neé, Anales Ci. Nat. 3: 265 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA25957 ! designated by Muller & McVaugh (1972, p. 520); isolectotypes: MA!, UCSB (fragment)!). We are treating Q. salicifolia in a broad sense to include what could be recognized as many separate species if a narrower species concept were adopted. Further study of this species complex is needed.

Quercus acapulcensis Trel., Mem. Natl. Acad. Sci. 20: 153, pl. 302a (1924). Type:— MEXICO, Guerrero: E. Palmer 334 (Holotype: GH00069901 !; isotypes: CAS!, DS!, F!, K!, NY!, UCSB (fragment)!, US!) .

Quercus boquetensis Standl., Publ. Field Mus. Nat. Hist., Bot. Ser. 22(1): 13 (1940). Type:— PANAMA, Chiriquí: M. E. Davidson 780 (Holotype: F!; isotypes: GH!, MO!, US!). Additional study is needed to determine if Q. boquetensis might actually be a good species.

Quercus borucasana Trel., Mem. Natl.Acad. Sci. 20: 161, pl. 315a (1924). Type:— COSTA RICA, Cartago: A. S. Oersted 3491 (Holotype: C10014102 !). Nom. nov. for Q. granulata Liebm. (1854) non Raf. (1838).

Quercus citrifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 187 (1854) . Type :— COSTA RICA, Cartago: A. S. Oersted 3461 (Lectotype (designated here): C 10012469!; isolectotypes: C!, CAS!, G!, UCSB (fragment)!). Trelease (1924) in pl. 288 shows two different specimens, but in his caption for pl. 288 states “The type (Costa Rica, Oersted, 6=3461).” Specimen 6=3461 is the bottom image of pl. 288 and is being lectotypified here, and 8=3461, the top image, which is now at CAS , becomes an isolectotype. A third specimen, 7=3461, is an isolectotype at C.

Quercus eugeniifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 185 (1854) . Type:— COSTA RICA: J. R. R. Warszewicz s. n. (C) (Lectotype (designated here): image in Trelease (1924, pl. 316a)!). Liebmann (1854, p. 185) wrote “Guatimala [sic] et Costa rica [sic] leg. Warsewicz [sic] (in hb. reg. Berol.!) Species affinis Q. salicifoliae Née. ” Trelease (1924, p. 161, pl. 316a) effectively lectotypified Warszewicz C at B. The photograph in pl. 316a is Q. salicifolia . The lectotype at B was destroyed, and because there are no extant specimens of Warszewicz at C, we are lectotypifying the name based on pl. 316a. Trelease cites Warszewicz 29 as also being Q. eugeniifolia , although without stating where he saw the specimens. If this specimen was at B it was destroyed, and the specimens of Warszewicz 29 at G and P are Q. acutifolia .

Quercus eugeniifolia f. petiolata Trel., Mem. Natl. Acad. Sci. 20: 161, pl. 316b (1924). Type :— COSTA RICA: C. Hoffmann 863 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 316b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus granulata Liebm. non Raf. (1838), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 186 (1854). See Q. borucasana above.

Quercus mulleri Martínez, Anales Inst. Biol. Univ. Nac. Autón. México 24(2): 51, fig. 1, 2 (1953[1954]). Type :— MEXICO, Oaxaca: T. MacDougall 2435 (Holotype: MEXU00011238 !; isotypes: CAS!, ENCB (image)!, UCSB (fragment)!).

Quercus nixoniana S.Valencia & Lozada-Pérez, Novon 13(2): 261, f. 1 (2003). Type :— Mexico, Guerrero: R. de Santiago 747 (Holotype: FCME; isotypes: BH, IBUG, MEXU, MO, XAL). The relegation of this epithet to synonymy is based on the protologue and illustration as well as examination of numerous paratypes.

Quercus obtusanthera Trel., Repert. Spec. Nov. Regni Veg. 33: 318 (1934). Type:— MEXICO, Guerrero: L. Schultze 343 (Holotype: at B destroyed). The determination of Q. obtusanthera as probably Q. salicifolia is based on Trelease’s (1933) description as no images or original specimens exist.

Quercus petiolata (Trel.) A.E.Murray non Schur (1857), Kalmia 13: 27 (1983).

Quercus rubramenta Trel., Repert. Spec. Nov. Regni Veg. 33: 318 (1934). Types: MEXICO, Guerrero. L. Schultze 371 and 372 (Syntypes: at B destroyed). The determination of Q. rubramenta as Q. salicifolia is based on Trelease’s (1934) description as no images or original specimens exist.

Quercus salicifolia var. seemannii (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 207 (1884).

Quercus seemannii Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 188 (1854) . Type:— PANAMA, Veraguas: B. Seemann 1228 (Lectotype: K000575040 ! designated by Trelease (1924, p. 226, pl. 317a); isolectotypes: BM!, G!, GH? (as s.n.)!, K!, UCSB (fragment)!). Liebmann (1854, p. 188) wrote “leg. Seemann (Coll. n. 1228 in hb. Hook.!)” but there are two collections of Seemann 1228 in the Hooker Herbarium, both of which are named by Liebmann in his hand as Q. seemanni Liebm.

Quercus tahuasalana Trel., Mem. Natl.Acad. Sci. 20: 154, pl. 302b (1924). Type:— MEXICO, Michoacán: E. Langlassé 211 (Syntypes at B destroyed. Lectotype (designated here): G00358146 !; isolectotypes: GH!, K!, P!, UCSB (fragment from G lectotype)!, UCSB (fragment)!, US!). Trelease (1924, p. 154) wrote “ Langlassé, 217, June 18, 1898, the type” but on p. 226 in the caption to pl. 302b he wrote “The type (Mexico, Langlassé, 211).” The image detail in pl. 302b is from B now lost. Both Langlassé 211 and 217 are Q. salicifolia and are considered to be syntypes.

Distribution: Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Colima, Durango, Guerrero, Hidalgo,

Jalisco, México, Michoacán, Morelos, Nayarit, Oaxaca, Puebla, San Luis Potosí, Sinaloa, Veracruz, Zacatecas ),

Nicaragua, Panama. The occurrence of Q. salicifolia in México, San Luis Potosí, Sinaloa, and Zacatecas reported by

Villaseñor (2016) needs to be confirmed.

Quercus sapotifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 185 (1854) . Type :— GUATEMALA: G. V. Skinner 6 (Holotype: K000512931 !) .

Quercus amissiloba Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 357 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6379 (Holotype: ILL00007474 !; isotypes: F!, G!, GH!, K!, MO!, NY!, US!) .

Quercus apanecana Trel., Mem. Natl. Acad. Sci. 20: 152 (1924). Type:— EL SALVADOR, Ahuachapan: P. C. Standley 20187 (Holotype: US 00089403!) .

Quercus donnell-smithii Trel., Mem. Natl. Acad. Sci. 20: 162, pl. 318 (1924). Type:— GUATEMALA, Guatemala: J. D. Smith 1967 (Lectotype (designated here): US00089494!; isolectotypes: GH!, NY!). Although Trelease cited Smith 1967 as the type, there is no indication of a collection locality. In pl. 318 two different specimens are shown which Trelease says “are two variants of the type.” Neither of these is a good match for the NY or US specimens.