Pygovepres, WEIRAUCH, 2006

WEIRAUCH, CHRISTIAN E., 2006, New Genera and Species of Oak-Associated Phylini (Heteroptera: Miridae: Phylinae) from Western North America, American Museum Novitates 3521 (1), pp. 1-56 : 16-23

publication ID	https://doi.org/ 10.1206/0003-0082(2006)3521[1:NGNSAN]2.0.CO;2
persistent identifier	https://treatment.plazi.org/id/038487D4-F20F-0028-C483-FD103906F928
treatment provided by	Carolina
scientific name	Pygovepres
status	gen. nov.

Treatment

Pygovepres View in CoL View at ENA , new genus

Figures 1 View Fig , 4 View Fig , 6–12 View Fig View Fig View Fig View Fig View Fig View Fig View Fig

TYPE SPECIES: Psallus vaccinicola Knight, 1930 .

DIAGNOSIS: Recognized by the moderate size (male total length: 3.00–3.26), pale orange to reddish and whitish coloration with anterior crescent-shaped portion of cuneus transparent, and distal portion of cuneus darker orange or more reddish than rest of the corium; left side of the pygophore anterior to the genital opening with a protuberance bearing pointed processes each equipped with a single short seta (figs. 9F, G), the left paramere deeply excavated with the anterior process flattened laterally, the apex of the posterior process bent ventrally and truncate (fig. 4), and the slender C-shaped vesica with a small membranous lobe beset with spicules proximal to the secondary gonopore (figs. 4, 10).

DESCRIPTION: Male: Macropterous, size moderate (3.00–3.26), elongate ovoid. COLORATION (fig. 1): General coloration variable, rather uniform pale orange to reddish orange with cuneus often slightly darker than remaining hemelytron to pale red with dark red cuneus or entirely dark red, anterior crescent-shaped portion of cuneus and membrane whitish and transparent, sometimes anterior margin of anterior pronotal lobe, entire posterior pronotal lobe, and apical portion of scutellum whitish. Head: Coloration ranging from almost uniformly reddish orange to distinct orange-whitish color pattern; in former case, head orange with only subapical portion of clypeus and buccula and gula whitish; in latter case, vertex and frons whitish, suffused with orange at posterior margin, and with transverse paired fasciae and spot on inner rim of eye orange, clypeus whitish with base and two longitudinal stripes orange, sometimes with apex brown, mandibular plate whitish, whitish with central orange spot, or largely suffused with orange, maxillary plate and gena orange, buccula and gula whitish, neck orange to brownish; antennal segments 1 and 2 yellowish, segment 1 sometimes with pale brown basal and apical ring, segment 2 slightly darkened toward apex, segments 3 and 4 pale brown; labium yellowish, with segment 4 suffused with brown. Thorax: Pronotum varying from entirely reddish orange, to orange with only lateral margins of pronotum whitish to anterior lobe whitish with calli orange or reddish and posterior lobe largely pale with central and posterior parts suffused with orange or reddish; mesoscutum ranging from pale with orange lateral marks in pale specimens to uniformly orange or red in darker specimens; scutellum ranging from orange with whitish tip in pale specimens to entirely orange or reddish in darker specimens; propleura brownish or orange, often with margin of coxal cavity and dorsal margin whitish; mesepisternum uniformly reddish brown or brown, mesepimeron either uniformly reddish or whitish suffused with orange; metapleuron reddish, orange or pale brown; evaporatorium sometimes lighter or whitish with peritreme brownish. Legs: Pale with base of coxa suffused with pale brown; femur with small, sparsely distributed dark spots, at least proximal tibial spines with dark bases; tibia sometimes suffused with pale brown toward apex; tarsus pale with apical tarsomere suffused with brown. Hemelytra: Corium including clavus and cuneus orange to red, with cuneus often more deeply stained than remaining hemelytron, small spot on posterior margin distal to apex of clavus whitish or—in dark specimens—pale, a crescent-shaped anterior portion of cuneus whitish or transparent and sometimes anterior and proximal part of corium lighter; membrane translucent or slightly infuscate. Abdomen: Venter either entirely brown, reddish brown, or pale brown, sometimes suffused with reddish or yellow, occasionally with pale areas on ventral laterotergites. SURFACE AND VESTITURE: Dorsum and hemelytra shining, covered rath- er densely with two types of simple, long and slender, subadpressed setae, two types (black vs. white asterisks) differening slightly in their diameter (fig. 8C). STRUCTURE: Head (fig. 4): Only slightly broader than long, vertex wide, slightly convex, posterior margin straight, vertex and frons sloping, clypeus moderately produced, mandibular and maxillary plates short, latter sunken, buccula short, buccal cavity oval, gula short; eyes almost as high as head, only weakly emarginate posterior to fossa, posterolateral margin contiguous with anterolateral margin of pronotum; antennal insertion contiguous with anterior margin of eye, antennal segment 1 short and slender, slightly widened toward apex, segment 2 longest, about as long as greatest pronotal width, diameter similar to segment 1, very slightly increasing toward apex, segments 3 and 4 with diameter smaller than segments 1 and 2, together shorter than segment 2; labium reaching to between base of metacoxa and base of abdomen. Thorax: Pronotum trapeziform, anterior margin slightly sinuate, lateral margins weakly convex, posterior margin slightly concave, anterior and posterior pronotal lobes weakly demarcated, calli not developed, sometimes demarcated by distinct coloration; metapleural evaporatorium with mushroomlike cuticle area broad (fig. 6C), mushroomlike cuticle around mesothoracic spiracle well developed. Legs: Slender; claws slender, claw hairs present, pulvilli of moderate size, covering half of ventral claw surface, parempodia setiform (fig. 7). Hemelytra: Slightly convex, cuneus triangular. Abdomen: Reaching to apex of cuneus. GENITALIA (figs. 4, 9F–H, 10): Pygophore: Protuberance on left side anterior to genital opening beset with five spinelike processes (figs. 9F, G), four anterior processes closely grouped, fifth placed posterior to them, each bearing a very short and slender seta. Parameres: Right paramere typically phyline lanceolate (fig. 4); body of left paramere laterally flattened, dorsal lobe of body set apart by an indentation on dorsal rim between body and posterior process, anterior process flattened dorsoventrally (figs. 4, 9F). Phallotheca: Phallotheca slender and long, with a small process on posterior surface (figs. 4, 9H), slitlike opening ventral. Vesica: Almost circularly C-shaped, apex a slender simple spine slightly twisted to left, secondary gonopore subapical and of moderate size, a weakly sclerotized process on left side of vesical body proximal to secondary gonopore, beset with one to four lateral teeth, gonopore sclerite inconspicuous (figs. 4, 10).

FEMALE: Elongate ovoid, slightly stouter than male (fig. 1); coloration similar to male, sometimes somewhat paler; antennal segment 2 more slender than in male and covered less densely with setae. GENITALIA: See description of type species P. vaccinicola .

ETYMOLOGY: Named for the spiny process on the pygophore, combining ‘‘pygophore’’ and Latin noun vepres, meaning ‘‘thorny bush’’. The gender of the name is masculine.

DISCUSSION: A ‘‘cluster of four spines’’ ventrolaterally on the pygophore is a diagnostic character of Platyscytisca Costa & Henry, 1999 . Given the different structure and position of these ornamentations in Pygovepres and Platyscytisca , it appears unlikely that these structures are homologous.

Pygovepres vaccinicola (Knight) , new combination

Figures 1 View Fig , 4 View Fig , 6–12 View Fig View Fig View Fig View Fig View Fig View Fig View Fig

Psallus vaccinicola Knight, 1930: 128 [new species].

Psallus vaccinicola: Carvalho 1958: 133 [catalog].

Psallus vaccinicola: Henry & Wheeler 1988: 495 [catalog, distr.].

Psallus vaccinicola: Schuh 1995: 416 [catalog].

HOLOTYPE: Male: USA: Arizona: Coconino Co.: Grand Canyon, 36.05444 ° N 112.13861 ° W, 2134 m, 22 Jun 1925, A. A. Nichol, Vaccinium sp. (Ericaceae) , 13 ( AMNH _PBI 00069279) ( USNM).

REVISED DIAGNOSIS: Recognized by the orange to red coloration with anterior portion of cuneus transparent (fig. 1), the protuberance with several pointed processes, each bearing a single short and slender seta on the left hand side of the pygophore anterior to the genital opening (figs. 9F, G) and the slender and C-shaped vesica with a weakly sclerotized process proximal to the secondary gonopore (figs. 4, 10).

REVISED DESCRIPTION: Male: Size moderate, elongate ovoid; total length 3.00–3.26, length from apex of clypeus to cuneal fracture 2.02–2.26, width across pronotum 1.03–1.08. COLORATION (fig. 1): As in generic description. Head: Head including eyes, antennae and labium as in generic description. Thorax: Thorax including legs and hemelytra as in generic description. Abdomen: As in generic description. SURFACE AND VESTITURE: As in generic description. STRUCTURE: Head: Head including eyes, antennae, and labium as in generic description. Thorax: Thorax including legs and hemelytra as in generic description. Abdomen: As in generic description. GENITALIA: As in generic description.

FEMALE: As in generic description (fig. 1). Total length 3.12–3.72, length from apex of clypeus to cuneal fracture 2.20–2.53, width across pronotum 1.03–1.16. GENITALIA: Vestibulum very large, coiled, bursa copulatrix almost triangular, posterior margin with median indentation, sclerotized rings large and slender (fig. 11).

HOSTS: Recorded from more than ten species of Quercus (Fagaceae) , but also from Vaccinum sp. ( Ericaceae ), Ceanothus and Rhamnus (Rhamnaceae) , Amelanchier , Cercocarpus and Purshia (Rosaceae) , and some other plant species. The largest numbers of specimens were recorded from Quercus douglasii H. & A. (449 specimens), Q. john-tuckeri Nixon and Muller (255), Q. lobata Nee (250), Q. garryana Dougl. (195), and Amelanchier utahensis Koehne (215). The large specimen numbers collected on these hosts indicate that these plants may be breeding hosts of P. vaccinicola , even though no immature stages were collected. The host plant of the holotype of P. vaccinicola , as species of Vaccinium , was recorded for a total of only six specimens and thus more likely represents a sitting than breeding record.

DISTRIBUTION: Known from the Western United States (Arizona, California, Colorado, Idaho, Nevada , Oregon, Utah, Washington) and British Columbia, Canada (fig. 12) .

DISCUSSION: Knight (1930) described this species in Psallus Fieber but did not provide arguments for this taxonomic decision. The structure of the male genitalia clearly excludes this species from Psallus . Pygovepres vaccinicola is a widespread species, very variable in coloration, and breeds on a large number of, at least in part, not closely related hosts. However, examination of the male genitalia from numerous different hosts and localities did not reveal characters that could be used to recognize more than one species.

Pygovepres vaccinicola was recorded from the same localities as all the other species treated in this paper. For more detailed information on overlap in distribution of P. vaccinicola with these species, refer to the discussion section for the given species.

OTHER SPECIMENS EXAMINED: CANADA: British Columbia: Kaleden, 49.4 ° N 119.6 ° W, 2 Jul 1975, L. A. Kelton, alfalfa, 2♀ (CNC). Keremeos, 49.2 ° N 119.81666 ° W, 2 Jul 1975, L. A. Kelton, Spiraea sp. (Rosaceae) , 2♀ (CNC). Oliver, 49.18333 ° N 119.55 ° W, 2 Jul 1974, L. A. Kelton, Ceanothus sp. (Rhamnaceae) , 43 (CNC). Oliver, Madden Lake, 49.18333 ° N 119.55 ° W, 3 Jul 1959, L. A. Kelton, 53, 4♀ (CNC). Oliver, Sawmill Lake, 49.18333 ° N 119.55 ° W, 30 Jun 1959, L. A. Kelton, 13, 2♀ (CNC). USA: Arizona: Coconino Co.: 7 miles N of Jacob Lake on Route 89, 36.82698 ° N 112.25061 ° W, 25 Jun 1980, R. T. Schuh, Amelanchier utahensis (Rosaceae) , det. B. Ertter 1980, 93, 10♀ (AMNH). Gila Co. : 8 mi SW jct Rts 87 and 188 (off Rt 87), Tonto National Forest, 33.55989 ° N 111.21341 ° W, 1219 m, 27 May 1983, R. T. Schuh and G. M. Stonedahl, Cercocarpus montanus (Rosaceae) , 33, 13♀ Amelanchier sp. (Rosaceae) , 83, 15♀, Light Trap, 73, 5♀ (AMNH). Yavapai Co.: Mud Tanks Mesa, George Crook Road, 33.985 ° N 112.41583 ° W, 1981 m, 14 Jun 1983, R. T. Schuh and M. D. Schwartz, Amelanchier utahensis (Rosaceae) , 13, 2♀ (AMNH). California: Alpine Co.: 4 mi SE of Markleeville, 38.65411 ° N 119.7268 ° W, 2484 m, 10 Jun 1966, W. Gagne, 143, 7♀ (UCB). 6 miles S Markleeville, 38.60801 ° N 119.77917 ° W, 1829 m, 8 Jul 1966, C. W. O’Brien, 13, 4♀ (CAS). Woodfords, 38.77778 ° N 119.82083 ° W, 11 Jun 1966, C. W. O’Brien, 13 (UCB). Butte Co.: 5 miles W Paradise, 39.75968 ° N 121.7151 ° W, 13 May 1961, G. W. Frankie, 13 (UCB). Big Bend Mountain, 39.69722 ° N 121.44639 ° W, 23 May 1928, H. H. Keifer, Ceanothus integerrimus (Rhamnaceae) , 43, 4♀ (CAS). Chico, 39.72861 ° N 121.83639 ° W, 18 Apr 1928, H. H. Keifer, Quercus lobata (Fagaceae) , 13 (CAS). Oroville, 39.51389 ° N 121.55528 ° W, 17 Apr 1928, H. H. Keifer, Quercus douglasii (Fagaceae) , 13, 1♀, 3;u Quercus lobata (Fagaceae) , 63 Ceanothus cuneatus (Rhamnaceae) , 13, 1♀ (CAS); 29 Apr 1928, H. H. Keifer, Quercus wislizenii (Fagaceae) , 13 Ceanothus cuneatus (Rhamnaceae) , 23 (CAS); 9 Apr 1928, H. H. Keifer, Quercus douglasii (Fagaceae) , 13 (CAS); 29 Apr 1927, H. H. Keifer, Quercus douglasii (Fagaceae) , 13 Quercus lobata (Fagaceae) , 63, 6♀ (CAS); 2 May 1928, H. H. Keifer, 13 Quercus sp. (Fagaceae) , 23 (CAS); 1 May 1928, H. H. Keifer, Quercus douglasii (Fagaceae) , 33 (CAS). Calaveras Co.: 4.8 km S of West Point, 38.35607 ° N 120.52639 ° W, 22 May 1981 - 24 May 1981, S. C. Williams, 43 (CAS). Colusa Co. : 7.9 mi NW Sites, 39.37939 ° N 122.38378 ° W, 30 Apr 1980, F. G. Andrew, S. Kuba and T. D. Eichlin, 13 (CAFA). Rumsey Canyon, 38.925 ° N 122.32534 ° W, 10 May 1958, E Jessen, 13 (UCB). Contra Costa Co.: 2 miles S Curry Creek Park along Marsh Creek, 37.9824 ° N 121.68876 ° W, 13 May 1977, D. D. Wilder, 103 (CAS). 3 mi E Crockett, 38.05248 ° N 122.1567 ° W, 18 Jun 1965, C. E. Hawkins, 23 (ORSU). Lafayette, 37.88583 ° N 122.11694 ° W, 12 May 1952, R. A. Stirton, 13 (UCB). Mount Diabolo, 37.88159 ° N 121.91384 ° W, 18 May 1947, H. P. Chandler, 13 (CAS). Mount Diabolo State Park Stagecoach, 37.88159 ° N 121.91384 ° W, 10 Jun 1975, R. Kawin, 13, 1♀ (UCB). El Dorado Co.: Blodgett Forest, 13 mi E Georgetown, 38.90669 ° N 120.59532 ° W, 1295 m, 2 Jun 1973, J. Powell, 13 (UCB). Fallen Leaf Lake, 38.88306 ° N 120.07167 ° W, 11 Jul 1931, O. H. Swezey, 1♀ (CAS). Pilot Creek, 38.8433 ° N 121.05769 ° W, 26 Jun 1967, W.J. Turner, Ceanothus integerrimus (Rhamnaceae) , 13 (UCB). Quinette, El Dorado county, 38.76667 ° N 120.51667 ° W, 24 Jun 1967, J. Powell, Ceanothus integerrimus (Rhamnaceae) , 13 (UCB). Ceanothus integerrimus (Rhamnaceae) , 13 (UCR). Tallac Lake, Tahoe, 38.91194 ° N 120.11056 ° W, 25 Jul 1915, E. P. Van Duzee, 13 (UCB). Fresno Co.: 7 miles E Piedra, 36.81021 ° N 119.25431 ° W, 5 May 1963, R. Orbeck, 13 (UCB). Glenn Co.: 5 mi N Elk Creek, 39.6029 ° N 122.53662 ° W, 7 Jun 1984, J. D. Pinto, Quercus douglasii (Fagaceae) , 13 (UCR). Humboldt Co.: 3 miles N Garberville, 40.14376 ° N 123.79389 ° W, 23 May 1976, R. Wharton and R. Coville, 13 (UCB). Blocksburg, 40.27611 ° N 123.63528 ° W, 25 May 1941, B. P. Bliven, 23 (CAS). Bridgeville, 40.46944 ° N 123.79861 ° W, 20 Jun 1959, Kelton and Madge, Ceanothus sp. (Rhamnaceae) , 13 Spiraea sp. (Rosaceae) , 13 (CNC). Dinsmores, 40.49167 ° N 123.60583 ° W, 4 Jun 1939, B. P. Bliven, 43 (CAS). Garberville, 40.10028 ° N 123.79389 ° W, 5 May 1940, B. P. Bliven, 23 (CAS). Laribee Valley, 32.65306 ° N 116.70639 ° W, 3 Jun 1934, Brunson P. Bliven, 23 (CAS); 19 Jun 1938, B. P. Bliven, 23 (CAS). Redway, 40.12028 ° N 123.82222 ° W, 15 May 1938, B. P. Bliven, 53 (CAS). Kern Co.: E of Caliente on Caliente Creek Rd, 35.31251 ° N 118.5042 ° W, 820 m, 22 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Mimulus aurantiacus pubescens (Torr) D. Thompson (Scrophulariaceae) , det. A. Sanders UCR 140565, 13 (AMNH). E of Caliente on Caliente Creek Rd, Twin Oaks, 35.30702 ° N 118.4233 ° W, 836 m, 22 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, 33 Juniperus californica Carr. (Cupressaceae) , det. A. Sanders UCR 140634, 13 Quercus douglasii H. and A. ( Fagaceae ), det. K. Nixon 2004, 1173, 179♀ Quercus X alvordiana Eastwood ( Fagaceae ),det. K. Nixon 2004, 223, 43♀ (AMNH). E of Caliente on Walker Basin Road, 35.38774 ° N 118.425 ° W, 1220 m, 22 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus sp. (Fagaceae) , 43, 15♀ Quercus wislizenii A. DC. (Fagaceae) , det. K. Nixon 2004, 193, 22♀ Ribes quercetorum Greene (Grossulariaceae) , 13, 14♀ Ceanothus sp. (Rhamnaceae) , 73, 1♀ (AMNH). S of Bodfish on Caliente-Bodfish Road, 35.56089 ° N 118.5053 ° W, 1094 m, 22 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Fremontodendron californicum (Sterculiaceae) , det. A. Sanders 2004 UCR 140607, 23, 2♀ Fremontodendron californicum (Torr.) Corville (Sterculiaceae) , 23, 2♀ Quercus sp. (Fagaceae) , 43 Quercus douglasii H. and A. ( Fagaceae ), det. K. Nixon 2004, 163, 25♀ Quercus douglasii H. and A. ( Fagaceae ), det. K. Nixon 2004, 163, 25♀ Artemisia tridentata parishii (Gray) Hall and Clements (Asteraceae) , 13 (AMNH). S of Tehachapi at jct Tehachapi Willow Springs Rd and Oak Creek Rd, 35.05019 ° N 118.3525 ° W, 1235 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, 83 Quercus lobata Nee (Fagaceae) , det. K. Nixon 2004, 923, 131♀, 1;u Quercus X alvordiana Eastwood ( Fagaceae ), det. K. Nixon 2004, 83, 22♀ (AMNH). S of Tehachapi on Tehachapi Willow Springs Rd, 35.05502 ° N 118.3696 ° W, 1329 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus john-tuckeri Nixon and Muller (Fagaceae) , 963, 75♀ Quercus john-tuckeri Nixon and Muller (Fagaceae) , det. A. Sanders 2004 UCR-140663, 963, 75♀ Quercus john-tuckeri Nixon and Muller (Fagaceae) , det. A. Sanders 2004, 963, 75♀ (AMNH). S of Tehachapi on Water Canyon Rd, 35.09563 ° N 118.4909 ° W, 1445 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, 23 Quercus chrysolepis Liebm. (Fagaceae) , det. K. Nixon 2004, 143, 2♀ Quercus douglasii H. and A. ( Fagaceae ), 73, 12♀, 1;u Quercus douglasii H. and A. ( Fagaceae ), det. K. Nixon 2004, 73, 12♀, 1;u Quercus john-tuckeri Nixon and Muller (Fagaceae) , det. K. Nixon 2004, 433, 48♀ Rhamnus ilicifolia (Rhamnaceae) , det. A. Sanders 2004, 93, 12♀ (AMNH). Tehachapi Loop near Hwy 58, 35.13222 ° N 118.44806 ° W, 25 Mar 1981, M. D. Schwartz, Ribes sp. (Grossulariaceae) , 83, 6♀ (AMNH). Tehachapi Pass, 35.1 ° N 118.29444 ° W, 6 Jun 1929, R. L. Usinger, 13, 5♀ (CAS); 6 Jun 1929, E. P. Van Duzee, 33 (CAS). Wofford Heights, 35.70694 ° N 118.45528 ° W, 12 Jun 1914, H. F. Howden, Light Trap, 13 (CNC). Lake Co.: 10 mi E Clear Lake Oaks, 38.99578 ° N 122.52323 ° W, 8 May 1985, W. F. Chamberlain, 13 (TAMU). 5 mi W Colusa County line on Hwy 20, 39.22718 ° N 122.13384 ° W, 16 May 1981, R. S. McClay, 13 (TAMU). Lassen Co.: 3 mi W of Nubieber, 41.09583 ° N 121.23948 ° W, 1405 m, 6 Jul 1979, R. T. and Joe Schuh, Purshia tridentata (Rosaceae) , 23 (AMNH). Susanville, 40.41639 ° N 120.65194 ° W, 17 Jun 1959, Kelton and Madge, oak, 253, 24♀ (CNC). Susanville, 40.41639 ° N 120.65194 ° W, 17 Jun 1959, Kelton and Madge, oak, 43, 8♀ (CNC). Los Angeles Co.: 22 mi S of Palmdale, 34.26139 ° N 118.11556 ° W, 914 m, 30 May 1981, J. T. Polhemus, 4♀ (JTP). Palmdale, 34.57944 ° N 118.11556 ° W, 14 Apr 1932, E. P. Van Duzee, 1♀ (CAS). Pasadena, 34.14778 ° N 118.14361 ° W, 25 May 1909, Grinnell, 13, 1♀ (CAS). Tanbark Flat, 33.69111 ° N 116.67056 ° W, 24 Jun 1950, W. C. Bentinck, Quercus sp. (Fagaceae) , 13 (CAS); 23 Jun 1952, J. K. Hester, 13 (UCD); 24 Jun 1952, A. A. Grigarick, 13 (UCD); 24 Jun 1952, R. L. Anderson, 13 (UCD). Madera Co. : 1.8 mi SE of Route 41 at Route 274 and Pettit Rd, 37.00969 ° N 119.79426 ° W, 1036 m, 22 May 2001, T. J. Henry, Quercus sp. (Fagaceae) , 23 (AMNH). Coarsegold, 37.26222 ° N 119.7 ° W, 610 m, 28 May 1973, A. Gilbert, Ceanothus sp. (Rhamnaceae) , 13 (CAFA). Marin Co.: Mill Valley, 37.89083 ° N 122.5225 ° W, 25 Apr 1926, M. C. Van Duzee, 43, 1♀ (CAS). Phoenix Lake, 37.95583 ° N 122.57417 ° W, 14 May 1927, H. H. Keifer, 13, 2♀ Quercus lobata (Fagaceae) , 13 (CAS). Samuel P Taylor State Park, 38.02583 ° N 122.72667 ° W, 5 Aug 1962, J. T. Doyen, 13 (UCB). Terra Linda E of Rt 101 at Freitas Prwy, 38.00417 ° N 122.54861 ° W, 25 Apr 1980, Russell and Schwartz, oak, 13 (AMNH). Mariposa Co. : NW of Mariposa off Rt 140 on Bear Valley Rd, 37.56694 ° N 120.1324 ° W, 663 m, 25 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus douglasii H. and A. ( Fagaceae ), det. K. Nixon 2004, 8♀ Quercus wislizenii A. DC. (Fagaceae) , det. K. Nixon 2004, 33, 5♀ (AMNH). W of Mariposa near Mt. Bullion, 37.49936 ° N 120.0435 ° W, 675 m, 25 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus wislizenii A. DC. (Fagaceae) , det. K. Nixon 2004, 13, 7♀ (AMNH). W of Mariposa on Old Toll Rd, 37.50248 ° N 120.0689 ° W, 585 m, 25 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus douglasii H. and A. ( Fagaceae ), det. K. Nixon 2004, 53, 5♀ (AMNH). Yosemite National Park, 37.85 ° N 119.56667 ° W, 1201 m, 26 May 1931, [collector unknown], 13 (UCB). Yosemite National Park, 37.85 ° N 119.56667 ° W, 1201 m, 16 May 1931, E. O. Essig, 23, 4♀ (CAS); 21 May 1931, E. O. Essig, 43, 1♀ (CAS). Mendocino Co.: 6 mi SE of Hopland Field Station Headquarters, pond area, 38.94041 ° N 123.00446 ° W, 19 May 1973, J. Cate, 13, 1♀ (UCB). 7 mi W of Eel River Ranger Station, Mendocino National Forest, 39.62653 ° N 123.34493 ° W, 442 m, 10 Jun 1972, J. Doyen, Quercus sp. (Fagaceae) , 13 Achillea millefolium (Asteraceae) , 13 (UCB). Hopland, 38.97306 ° N 123.11528 ° W, 9 May 1926, E. P. Van Duzee, 13, 1♀ (CAS). Hopland Experiment Station, Kelsey Cab. Orchard area, 38.97306 ° N 123.11528 ° W, 808 m, 17 May 1974, A. Hajek, 13 (UCB). UC HoplandField Station, near Headquarters, 39.00194 ° N 123.08361 ° W, 18 May 1968, W. J. Turner, 23 (CAS). 173, 16♀ (UCB). UC Hopland Field Station, near Headquarters, 39.00194 ° N 123.08361 ° W, 268 m, 18 May 1969, W. J. Turner, 93, 9♀ (CAS). Modoc Co.: 18 mi S of Rt 139 towards Lookout, 40.94783 ° N 121.15417 ° W, 1420 m, 6 Jul 1979, R. T. and Joe Schuh, Quercus garryana (Fagaceae) , 53 (AMNH). Fandango Pass Summit, 41.80222 ° N 120.20583 ° W, 1890 m, 3 Jul 1979, R. T. Schuh and B. M. Massie, Cercocarpus ledifolius (Rosaceae) , 1♀ (AMNH). Mono Co.: Mono Lake at Hwy 395, 38.01667 ° N 119.00833 ° W, 2187 m, 11 Jul 1980, G.M. Stonedahl, Ribes sp. (Grossulariaceae) , 153, 23♀ (AMNH). Mono Lake, N edge at Rt 395, 38.01667 ° N 119.00833 ° W, 2187 m, 11 Jul 1980, R. T. Schuh, G.M. Stonedahl, Ribes sp. (Grossulariaceae) , 163, 12♀, 1;u (AMNH). Monterey Co.: Bryson, 35.80667 ° N 121.08944 ° W, 18 May 1920, E. P. Van Duzee, 23 (CAS); 20 May 1920, E. P. Van Duzee, 23 (CAS); 17 May 1920, E. P. Van Duzee, 13 (CAS). Jolon, 35.97083 ° N 121.175 ° W, 11 May 1959, T. R. Haig, 13 (UCD). Pleyto, 35.86028 ° N 120.9925 ° W, 21 May 1920, E. P. Van Duzee, 33, 3♀ (CAS). Napa Co.: Howell Mountain, N side, 2 mi NNE Angwin, 38.60254 ° N 122.43473 ° W, 396 m, 18 Apr 1986, H. B. Leach, 23 (CAS). Lake Berryessa, Steele Park Resort, unnamed creek, 38.51361 ° N 122.10278 ° W, 100 m, 6 May 1978, P. H. Arnaud, Salix hindsiana (Salicaceae) , 13 (CAS). Pope creek at Walter Springs road, 38.64605 ° N 122.35853 ° W, 29 Apr 1978, H. B. Leech, Ceanothus sp. (Rhamnaceae) , 13 (CNC); 6 May 1978, P. H. Arnaud, 23 (CAS). Snell valley, Stage Coach Canyon road at Spanish Valley trail, 38.67833 ° N 122.38278 ° W, 4 May 1980, H. B. Leech, 13 (CAS). Soda Creek, 38.35335 ° N 122.29027 ° W, 3 May 1931, H. H. Keifer, mahogany, 13 (CAFA). Nevada Co. : Donner Lake, 39.32772 ° N 120.26557 ° W, 4 Jul 1934, E. P. Van Duzee, 13 (CAS). Truckee, 39.32806 ° N 120.18222 ° W, 6 Jul 1927, E. P. Van Duzee, Quercus sp. (Fagaceae) , 1♀ (CAS). Orange Co.: Cleveland National Forest, Lower San Juan Campground on Hwy 74, 33.59889 ° N 117.46 ° W, 405 m, 12 May 1978, J. D. Pinto and R. T. Schuh, Lonicera subspicata (Caprifoliaceae) , 43, 1♀ (AMNH_PBI 00059353) Marah macrocarpus (Cucurbitaceae) , 173, 11♀ (AMNH). Marah sp. (Cucurbitaceae) , 33, 1♀ (UCR). Green River Camp, Lower Santa Ana Canyon , 33.74556 ° N 117.86694 ° W, 9 May 1933, E. P. Van Duzee, 33 (CAS). Placer Co.: G Alpine Cr, Lake Tahoe, 39.16667 ° N 120.14333 ° W, 5 Jul 1915, E. P. Van Duzee, 33 (UCB). Riverside Co.: 12 mi E of Hemet, San Bernardino National Forest, 33.70871 ° N 116.7611 ° W, 961 m, 20 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus wislizeni A. DC. var. frutescens Engelm. (Fagaceae) , det. K. Nixon 2004, 2♀ (AMNH). 5 mi N of Idyllwild on Rt 243, San Bernardino National Forest, 33.78326 ° N 116.7489 ° W, 1775 m, 20 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Ceanothus leucodermis (Rhamnaceae) , det. A. Sanders 2004 UCR-140683, 13 (AMNH). 6 mi W of Murrieta, Santa Rosa Plateau Area, 33.55 ° N 117.32074 ° W, 625 m, 1 May 1985, R. T. Schuh and B. M. Massie, Quercus agrifolia oxyadenia (Fagaceae) , det. K. Nixon 1985, 83 (AMNH_PBI 00095264), 7♀ (AMNH_PBI 00095268) Quercus dumosa (Fagaceae) , det. K. Nixon 1985, 43, 6♀ Quercus engelmanni (Fagaceae) , 373, 50♀ Quercus engelmanni (Fagaceae) , det. K. Nixon 1985, 373, 50♀ Quercus engelmanni (Fagaceae) , det. K. Nixon 1985, 373, 50♀ Quercus engelmanni Greene (Fagaceae) , det. K. Nixon 1985, 373, 50♀ Cercocarpus betuloides (Rosaceae) , 13 (AMNH). Bautista Canyon, 33.69778 ° N 116.85194 ° W, 5 May 1987, R. D. Goeden and D. W. Ricker, Encelia virginensis (Asteraceae) , 13 (UCR). Poppet Flat, 33.85 ° N 116.85167 ° W, 22 May 1976, B. A. Bowers, 23 (UCR); 28 May 1976, D. S. Hughes, oak, 23 (UCR). San Jacinto Mountains, S on J. River, 33.75028 ° N 116.66667 ° W, 30 May 1940, R. Husbands, Cercocarpus sp. (Rosaceae) , 13 (UCB). San Jacinto Mountains, jct of Two Pines Rd and Rt 243, 33.88083 ° N 116.84722 ° E, 20 May 2000, M. D. Schwartz, Quercus palmeri Engelm. (Fagaceae) , det. L. Raz 2002, 13, 1♀ (AM). Quercus palmeri Engelm. (Fagaceae) , det. L. Raz 2002, 43, 8♀ (AMNH). Quercus palmeri Engelm. (Fagaceae) , det. L. Raz 2002, 1♀ (CNC). Quercus palmeri Engelm. (Fagaceae) , det. L. Raz 2002, 13 (ZISP). San Jacinto Mountains, junction Poppet Flat Road and Route 243, 33.75028 ° N 116.66667 ° W, 20 May 2000, M. D. Schwartz, Quercus palmeri (Fagaceae) , det. L Raz 2002, 333, 11♀ Quercus palmeri (Fagaceae) , det. L. Raz 2002, 333, 11♀ Quercus palmeri Engelm. (Fagaceae) , det. L. Raz 2002, 333, 11♀ Rhamnus tomentella (Rhamnaceae) , det. L Raz 2002, 13, 3♀ Rhamnus tomentella (Rhamnaceae) , det. L. Raz 2002, 13, 3♀ (AMNH). Quercus palmeri Engelm. (Fagaceae) , det. L. Raz 2002, 13 (CNC). Quercus palmeri Engelm. (Fagaceae) , det. L. Raz 2002, 13, 1♀ (USNM). Quercus palmeri Engelm. (Fagaceae) , det. L. Raz 2002, 1♀ (ZISP). Santa Rosa Plateau Reserve , 33.52417 ° N 117.27528 ° W, 29 Apr 1985, J. D. Pinto, Quercus englemanni (Fagaceae) , 23, 3♀ (UCR). Tenaja Road W of Murrieta, 33.55 ° N 117.21667 ° W, 410 m, 12 May 1978, J. D. Pinto and R. T. Schuh, Mimulus aurantiacus (Scrophulariaceae) , 23, 2♀ (AMNH). Sacramento Co.: Carmichael, 38.61722 ° N 121.32722 ° W, 19 Apr 1960, R. W. Wilkey, Spiraea sp. (Rosaceae) , 113, 25♀ (CAFA). Spiraea sp. (Rosaceae) , 23 (UCD). Citrus Heights, 38.60667 ° N 121.26556 ° W, 3 Jun 1967, A. D. and G. J. Keuter, 23 (CAS); 10 Jun 1967, A. D. and G. J. Keuter, 53, 2♀ (CAS). Sacramento, 38.58167 ° N 121.49333 ° W, 27 Apr 1981, R. L. Bugg, mahogany, 13 (CAFA). San Bernardino Co.: 2 km NW of I-15 on Rt 138, N of San Bernardino, 34.31089 ° N 117.4971 ° W, 1555 m, 18 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Fremontodendron californicum (Torr.) Corville (Sterculiaceae) , 243, 5♀ Quercus sp. (Fagaceae) , 243, 14♀ Quercus douglasii H. and A. ( Fagaceae ), 93, 14♀ Quercus wislizeni A. DC. var. frutescens Engelm. (Fagaceae) , det. K. Nixon 2004, 453, 48♀ Ceanothus sp. (Rhamnaceae) , 313, 8♀, 1;u Ceanothus leucodermis Greene (Rhamnaceae) , 23 (AMNH). 2 mi E of Wrightwood on Rt 2, 34.36083 ° N 117.59747 ° W, 1563 m, 30 Jun 1980, R. T. Schuh, Quercus dumosa (Fagaceae) , det. B. Ertter 1980, 133, 66♀ Quercus dumosa (Fagaceae) , det. B. Ertter 1980, 133, 66♀ Quercus dumosa Nutt. (Fagaceae) , det. B. Ertter 1980, 133, 66♀ (AMNH). Mill Creek, 34.08721 ° N 117.11396 ° W, 1829 m, 1 Jun 1976, D. S. Hughes, 23 (UCR). Yucaipa, 34.03361 ° N 117.04222 ° W, [date unknown], Drake, Paddock, Rowe and Worthy, Hoarhoond, 23 (CAFA). just E of Wrightwood on Rt 2, 34.35774 ° N 117.6105 ° W, 1700 m, 18 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus berberidifolia Liebm. (Fagaceae) , det. K. Nixon 2004, 103, 2♀, 2;u (AMNH). San Diego Co.: 5 mi E of Alpine on Rt 8, 32.83861 ° N 116.63333 ° E, 890 m, 21 May 2000, M. D. Schwartz, Platanus racemosa (Platanaceae) , det. L. Raz, 2002, 43, 2♀ (AMNH). 8.2 mi E of Dulzura on Rt 94 at mp 36.50, 32.64417 ° N 116.63977 ° W, 15 May 1982, M. D. Schwartz, Quercus dumosa (Fagaceae) , 23, 2♀ (AMNH). Laguna Mountain Road (Old Hwy 80) just SE jct with Rt 8, 32.8054 ° N 116.50872 ° W, 1180 m, 21 May 2000, M. D. Schwartz, Ceanothus greggii perplexans Engelm. (Rhamnaceae) , 43, 6♀ (AMNH). Laguna Mountains, Kitchen Creek Rd 1.4 mi N of Rt 8, 32.73222 ° N 116.48778 ° W, 1100 m, 21 May 2000, M. D. Schwartz, Ceanothus greggii perplexans (Rhamnaceae) , det. H. D. Hammond 2002, 283, 2♀ Ceanothus greggii perplexans (Rhamnaceae) , det. H. D. Hammond, 2002, 283, 2♀ (AMNH). San Diego, 32.71528 ° N 117.15639 ° W, 3 Jun 1913, W. S. Wright, 43, 4♀ (CAS); 3 May 1913, W. S. Wright, 13 (CAS). San Diego County, 32.71528 ° N 117.15639 ° W, 11 Apr 1913 - 3 Jun 1913, E. P. Van Duzee, 103, 4♀ (CAS); 29 Mar 1914 - 1 Jun 1914, E. P. Van Duzee, 13, 1♀ (CAS). San Diego County, 33.03333 ° N 116.8 ° W, 5 May 1913, E. P. Van Duzee, 13 (CAS); 4 May 1913, E. P. Van Duzee, 13 (CAS). Torrey Pines, 32.91476 ° N 117.25253 ° W, 30 May 1955, W. E. Simonds, 13, 1♀ (CAFA). just S of Santa Ysabel , 33.10917 ° N 116.67222 ° W, 1060 m, 30 Apr 1985, R. T. Schuh and B. M. Massie, Quercus agrifolia oxyadenia (Fagaceae) , det. K. Nixon 1985, 2♀ (AMNH). San Luis Obispo Co.: Arroyo Grande Creek SW of San Luis Obispo, 35.11644 ° N 120.58609 ° W, 160 m, 8 May 1985, R. T. Schuh and B. M. Massie, Quercus lobata Nutt. (Fagaceae) , 63 (AMNH). Avenales Guard Station Jct, Huasna Valley E of Arroyo Grande, 35.11861 ° N 120.58972 ° W, 840 m, 9 May 1985, R. T. Schuh and B. M. Massie, 143, 12♀ Quercus douglasii (Fagaceae) , 253, 15♀, 1;u (AMNH). E of Arroyo Grande, Husana Valley, Stony Creek Campground, 35.20933 ° N 120.25846 ° W, 9 May 1985, R. T. Schuh and B. M. Massie, Quercus agrifolia oxyadenia (Fagaceae) , 23 (AMNH). Morro Road near Atascadero, 35.48944 ° N 120.66972 ° W, 14 May 1962, B. P. Bliven, 13 (CAS). San Mateo Co.: San Mateo, 37.56444 ° N 122.35056 ° W, 19 May 1918, E. P. Van Duzee, 33, 5♀ (CAS). Santa Barbara Co. : Lake Cachuma (Reservoir), 34.58667 ° N 119.98 ° W, 25 Jun 1964, W. F. Chamberlain, 13 (TAMU). Upper Oso Campground off Rt 154, 34.55583 ° N 119.75389 ° W, 305 m, 7 May 1985, R. T. Schuh and B. M. Massie, 103, 1♀ Quercus dumosa (Fagaceae) , det. K. Nixon 1985, 383, 23♀ (AMNH). Santa Clara Co. : Flamingo Avenue, San Jose, 37.33688 ° N 121.88802 ° W, 23 May 1953, O. H. Swezey, oak, 23 (ORSU). Palo Alto, 37.44194 ° N 122.14194 ° W, 18 May 1922, E. O. Essig, 13 (UCB). Shasta Co.: Hat Creek, 40.83047 ° N 121.50891 ° W, 31 May 1947, W. F. Chamberlain, Ceanothus sp. (Rhamnaceae) , 13 (TAMU). Platina, 40.35972 ° N 122.89361 ° W, 8 Jun 1984, J. Pinto, Quercus sp. (Fagaceae) , 33, 7♀ (UCR). Redding, 40.58667 ° N 122.39056 ° W, 1 May 1977, T. R. Haig, 273, 22♀ (CAFA); 7 May 1974, T. R. Haig, 13 (CAFA). Siskiyou Co.: 1.1 mi E Klamath R, 41.52666 ° N 124.01592 ° W, 28 May 1971, Oman, 13 (ORSU). 3.7 mi W of McCloud, 41.25583 ° N 122.20948 ° W, 1390 m, 9 Jul 1980, R. T. Schuh and G. M. Stonedahl, Ceanothus cordulatus (Rhamnaceae) , 53, 3♀ (AMNH); 9 Jul 1980, G.M. Stonedahl, Ceanothus sp. (Rhamnaceae) , 83, 1♀ (AMNH). Happy Camp, 41.79333 ° N 123.37833 ° W, 22 Jun 1964, A. D. and G. J. Keuter, 13 (CAS). Lava Beds National Monument, Valentine Caves, 41.75333 ° N 121.50556 ° W, 30 Jun 1963, V. L. Vesterby, 13 (UCD). Mount Shasta City exit on Interstate 5, 41.3054 ° N 122.31701 ° W, 22 May 1981, J. D. Lattin, 23, 1♀ (ORSU). Sonoma Co. : Mark West Springs, 38.54917 ° N 122.71917 ° W, 10 May 1930, E. P. Van Duzee, 13 (CAS). Petaluma Marsh, 38.26139 ° N 122.53583 ° W, 24 Apr 1980, J. T. Polhemus, oak, 43, 3♀ (JTP). Stanislaus Co.: Del Puerto Canyon, Frank Raines Park, 37.48833 ° N 121.20583 ° W, 341 m, 12 Apr 1974, G. Sahara, 23 (UCB). Del Puerto Canyon, N Fork Del Puerto Creek, 37.48673 ° N 121.20517 ° W, 305 m, 22 May 1971, R. Hinslop, 13 (UCB). Frank Raines Park, Del Puerto Canyon, 18 mi W Patterson, 37.47167 ° N 121.45648 ° W, 16 Apr 1977, R. C. Berg, 13 (UCD). Sutter Co.: Marysville Buttes, 39.15926 ° N 121.64102 ° W, 2 May 1928, H. H. Keifer, manzanita, 23, 12♀ (CAS). Tehama Co. : 2 miles N Red Bluff junction of route 36 & 15, 40.20759 ° N 122.23472 ° W, 26 Apr 1980, Russell and Schwartz, Quercus garryana (Fagaceae) , 93, 6♀ (AMNH). Red Bluff, 40.17861 ° N 122.23472 ° W, 30 Apr 1953, H. T. Osborn, clover, 33 (CAFA). Red Bluff, Samson Slough, 40.18859 ° N 122.22188 ° W, 29 Apr 1984, D. S. Chandler, Quercus sp. (Fagaceae) , 163, 8♀, 33 (UNHP). Trinity Co.: 12 mi W of Whiskeytown, 40.66138 ° N 122.78896 ° W, 14 May 1997, W. F. Chamberlain, 13 (TAMU). 5 mi E of Lewiston, 40.70746 ° N 122.7108 ° W, 3 Jun 1992, G. M. Chamberlain, 13 (TAMU). Mountain Meadow Ranch, head of Coffee Creek, 41.10157 ° N 122.77786 ° W, 1859 m, 8 Jul 1969, J. Powell, 33 (UCB). Tulare Co.: 12 miles N of Kernville, 35.92877 ° N 118.42444 ° W, 10 May 1997, W. F. Chamberlain, 13, 1♀ (TAMU). 4 mi NE of Lemoncove, 36.42366 ° N 118.97279 ° W, 13 May 1963, S. W. Earnshaw, 13 (UCB). Horse Creek, 3 miles SW Three Rivers, 36.38543 ° N 118.94415 ° W, 2 May 1979, J. Powell, Quercus wislizenii (Fagaceae) , 43 (UCB). NW of Springville on Balch Park Drive, 36.26184 ° N 118.7677 ° W, 916 m, 23 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus wislizenii A. DC. (Fagaceae) , det. K. Nixon 2004, 13 (AMNH). NW of Springville on Wokohl Rd, 36.26025 ° N 118.7432 ° W, 1228 m, 23 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Quercus kellogii Newb. (Fagaceae) , det. K. Nixon 2004, 73, 10♀ Ceanothus integerrimus (Rhamnaceae) , det. A. Sanders 2004 UCR-140570, 73, 13♀ (AMNH). Sequoia National Forest, Headquarters Camp, 36.00028 ° N 118.50028 ° W, 10 May 1997, W. F. Chamberlain, 13 (TAMU). Sequoia National Park, 36.56417 ° N 118.77278 ° W, 29 May 1949, W. D. Pierce, Ceanothus sp. (Rhamnaceae) , 13, 3♀ (LACM). Woodlake, 36.41361 ° N 119.09778 ° W, 24 Apr 1932, E. P. Van Duzee, 23, 1♀ (CAS). Tuolumne Co.: Dardanelles Outlook on Route 108, 38.34555 ° N 119.93305 ° E, 1800 m, 27 Jul 1999, M. D. Schwartz, Quercus kellogii (Fagaceae) , 13, 3♀ (CNC). Yolo Co.: 5 mi NE Rumsey, 38.93956 ° N 122.17054 ° W, 21 Apr 1973, J. C. Miller, 13 (UCD). Creek Canyon, 37.83167 ° N 122.18667 ° W, 30 Apr 1954, E. I. Schlinger, 13 (UCD); 30 Apr 1954, W. H. Lange, 13 (UCD). Davis, 38.545 ° N 121.73944 ° W, 3 Jun 1948, E. I. Schlinger, 13 (UCD); 1 May 1966, J. S. Buckett, 13 (UCD). Putah Canyon, 37.83167 ° N 122.18667 ° W, 9 Apr 1961, D. Q. Cavagnaro, 13 (UCD). Rumsey, 38.88833 ° N 122.23639 ° W, 21 Apr 1973, L. T. Wilson, 23 (UCD); 21 Apr 1973, Y. B. Ibrahim, 13 (UCD). Yuba Co.: 18 mi NE of Marysville, Sierra Foothill Rge F.S., 39.32983 ° N 121.35239 ° W, 7 May 1977, B. D. Wahl, 13 (UCD). 3 mi N of Smartville, Sierra Foothill Field Station, 39.25087 ° N 121.2975 ° W, 427 m, 2 May 1980, J. DeBenedictis, Quercus wislizenii (Fagaceae) , 13 (UCB). Camp Far West Reservoir, 39.05051 ° N 121.31724 ° W, 6 May 1980, J. A. Powell, 13 (UCB). Dry Creek, 4 miles NW Smartville, 39.25036 ° N 121.35185 ° W, 3 May 1980, J. T. Doyen, 23 (UCB). Harry L. Englebright Lake Dam, 39.24012 ° N 121.27162 ° W, 4 May 1980, J. DeBenedictis, Heteromeles arbutifolia (Rosaceae) , 13 (UCB). Sierra Foothill Field Station, 5 miles N Smartville, 39.27998 ° N 121.2975 ° W, 427 m, 4 May 1980, M. E. Buegler, 13 (UCB); 4 May 1980, J. T. Doyen, Quercus kellogii (Fagaceae) , 13, 1♀ Rhamnus sp. (Rhamnaceae) , 2♀ (UCB). [Unknown county]: Morgan Hill, 33.36972 ° N 116.92083 ° W, 1 Jun 1922, [collector unknown], 13 (CAS). Colorado: Eagle Co.: S of Wolcott, 39.70278 ° N 106.67806 ° W, 22 Jun 1980, J. T. Polhemus, 133, 15♀ (JTP). Vail, 39.64028 ° N 106.37361 ° W, 26 Jun 1977, J. T. Polhemus, 13, 2♀ (JTP). Mesa Co. : Colorado National Monument, Liberty Cap Trailhead, 39.05972 ° N 108.69917 ° W, 1829 m, 10 Jun 1982, M. D. Schwartz, Amelanchier utahensis (Rosaceae) , 143, 9♀ (AMNH). Moffat Co.: Escalante Overlook on Harpers Corner Road, 8.1 miles N route 40, 40.32611 ° N 108.945 ° W, 2350 m, 8 Jul 1982, M. D. Schwartz, Amelanchier utahensis (Rosaceae) , 13 (AMNH). Montezuma Co. : Hovenweep National Monument, Holly Canyon, 37.44278 ° N 108.98056 ° W, 28 May 1983, D. A. and J. T. Polhemus, 93 (AMNH_PBI 00063624, AMNH_ PBI 00063625), 10♀ (JTP). Routt Co.: Steamboat Springs, 40.485 ° N 106.83111 ° W, 2134 m, 24 Jul 1983, J. T. and D. A. Polhemus, 23, 4♀ (JTP); 23 Jul 1983, D. A. and J. T. Polhemus, Vaccinium sp. (Ericaceae) , 43, 1♀ (JTP). Idaho: Franklin Co.: Thomas Spring, 42.14583 ° N 111.64111 ° W, 28 Jun 1974, G. F. Knowlton, 13 (USU). Nevada: Elko Co. : 18 mi SE of Halleck on Rt 11, Secret Canyon, T34N R60E Sec15, 40.829 ° N 115.1915 ° W, 1905 m, 26 Jul 1982, M. D. Schwartz, Crataegus douglasii (Rosaceae) , 13 (AMNH). Nye Co.: Nevada Atomic Test Site, Shoshone Mountains , 9–10.5 miles SW Tippapah Hwy on Mine Mountain Road, 36.88336 ° N 116.27896 ° W, 7 Jun 1983, R. T. Schuh, M. D. Schwartz, G. M. Stonedahl, Amelanchier utahensis (Rosaceae) , 203, 16♀ (AMNH). Oregon: Benton Co.: 10 mi NW Philomath, Woods Crk Rd, 44.64259 ° N 123.51006 ° W, 259 m, 1 Jul 1971, J. D. Lattin, 13 (ORSU). Corvallis, 44.56472 ° N 123.26083 ° W, 14 Jun 1981, J. D. Lattin, Quercus garryana (Fagaceae) , 93, 18♀ (ORSU); 1 Jun 1981, J. D. Lattin, Quercus garryana (Fagaceae) , 23, 6♀ (ORSU); 1 Jun 1981, J. D. Lattin, Quercus garryana (Fagaceae) , 13 (ORSU). Deschutes Co.: Pioneer Ford FC Metolius River, 44.38669 ° N 121.28962 ° W, 7 Jul 1959, K. M. Fender, 1♀ (ORSU). Douglas Co.: Buzzard Bay County Park, 43.6375 ° N 123.57222 ° W, 16 Jun 1971, Oman, 13 (ORSU). S of Roseburg at Round Prairie Exit on I-5, 43.08833 ° N 123.36833 ° W, 12 Jun 1979, R. T. Schuh, Quercus garryana Engelm. (Fagaceae) , 13 (AMNH). Jackson Co.: 0.5 mi E of Pinehurst, 42.11778 ° N 122.35525 ° W, 1082 m, 17 Jun 1979, J. D. Lattin, Amelanchier alnifolia (Rosaceae) , 1♀ (ORSU). 0.5 mi S of Siskiyou Summit on Old Rt 99, Old Siskiyou Rd, 42.06777 ° N 122.60583 ° W, 1311 m, 27 Jun 1979, J. D. Lattin, Quercus garryana (Fagaceae) , 123, 13♀ (ORSU); 27 Jun 1979, M. D. Schwartz, Quercus garryana (Fagaceae) , 83, 12♀ (AMNH). 0.5 mi S of Siskiyou Summit on Old Rt 99, Old Siskiyou Rd, 42.06777 ° N 122.60583 ° W, 1346 m, 27 Jun 1979, M. D. Schwartz, Quercus garryana (Fagaceae) , 13 (AMNH); 26 Jun 1979, R. T. and Joe Schuh, Quercus garryana (Fagaceae) , 23, 37♀ (AMNH). 1 mi E Pinehurst on Hwy 66, 42.11777 ° N 122.34546 ° W, 27 Jun 1979, G. Stonedahl, Quercus garryana (Fagaceae) , 43 (AMNH). 8 mi S Siskiyou Pass, old highway, 42.01018 ° N 122.61164 ° W, 1463 m, 29 Jun 1977, J. D. Lattin, 1♀ ( Rosaceae ), 13 (ORSU). 9 mi SE of Butte Falls, 42.4541 ° N 122.43613 ° W, 19 Jul 1973, Ritcher and Carlson, 13 (ORSU). Siskiyou, 42.075 ° N 122.60583 ° W, 14 Jun 1959, Kelton and Madge, 153, 7♀ (CNC). Siskiyou Summit, Old Road, 42.075 ° N 122.60583 ° W, 1260 m, 22 Jul 1999, Schwartz, Gillespie, Quiring, Quercus garryana (Fagaceae) , 13, 2♀ (CNC). Siskiyou Summit, Old Siskiyou Hwy and Frontage Rd, 42.075 ° N 122.60583 ° W, 1314 m, 1 Jul 1994, M. D. Schwartz, Quercus garryana (Fagaceae) , 163 (AMNH_PBI 00072273), 12♀ (AMNH). Quercus garryana (Fagaceae) , 63, 3♀ (CNC). Jefferson Co.: Camp Sherman, 44.46056 ° N 121.64 ° W, 5 Jul 1978, J. D. Lattin, Amelanchier sp. (Rosaceae) , 13, 2♀ (ORSU). Josephine Co.: just S of Rough and Ready Wayside, S of Cave Junction, 42.16306 ° N 123.64694 ° W, 12 Jun 1979, R. T. Schuh, Quercus breweri Engelm. (Fagaceae) , 23 (AMNH). 13 (CAS). Klamath Co.: 1 mi E of Hayden Mountain Summit on Highway 66,

Note: Text continues on page 37 42.11571 ° N 122.07866 ° W, 26 Jun 1979, G. Stonedahl, Amelanchier alnifolia (Rosaceae) , 13, 6♀ (AMNH). 1 mile E of Cascade Summit, Highway 66, 43.57943 ° N 122.02111 ° W, 26 Jun 1979, G.M. Stonedahl, Amelanchier alnifolia (Rosaceae) , 5♀ (AMNH_PBI 00095266) (AMNH). 10 mi NW of Klamath Falls, Above Geary Ranch, 42.44515 ° N 122.11093 ° W, 3 Jul 1954, Joe Schuh, 33, 1♀ (ORSU). 13 mi W of Keno on Rt 66, 42.12667 ° N 122.18228 ° W, 1402 m, 27 Jun 1979, M. D. Schwartz, Amelanchier alnifolia (Rosaceae) , 23, 2♀ (AMNH); 27 Jun 1979, J. D. Lattin, Amelanchier alnifolia (Rosaceae) , 13, 2♀ (ORSU). 5 miles SE Keno, 42.07542 ° N 121.85982 ° W, 27 Jun 1979, G. M. Stonedahl, Amelanchier alnifolia (Rosaceae) , 33 (AMNH_PBI 00095262), 1♀ (AMNH). Hayden Mountain Summit, W of Keno, 42.10694 ° N 122.09861 ° W, 1475 m, 27 Jun 1979, R. T. and Joe Schuh, Ceanothus velutinus (Rhamnaceae) , 23, 4♀ Ceanothus venosus (Rhamnaceae) , 23 Amelanchier alnifolia (Rosaceae) , 63, 7♀ (AMNH). Keno on Route 66, 42.12667 ° N 121.92889 ° W, 4 Jul 1982, G.M. Stonedahl and T.J. Henry, Amelanchier alnifolia (Rosaceae) , 43, 6♀ (AMNH). Siskiyou Summit on I5, 42.075 ° N 122.60583 ° W, 4 Jul 1982, G.M. Stonedahl and T.J. Henry, Quercus garryana (Fagaceae) , 43, 5♀ Ceanothus integerrimus (Rhamnaceae) , 93, 10♀ (AMNH). Ceanothus integerrimus (Rhamnaceae) , 13 (USU). Upper Klamath Lake, Dennie Creek, 42.39833 ° N 121.88139 ° W, 3 Jul 1954, Joe Schuh, 13, 1♀ (ORSU). between Worden and Keno, 42.08806 ° N 121.90009 ° W, 1280 m, 27 Jun 1979, R. T. and Joe Schuh, Amelanchier alnifolia (Rosaceae) , 13 (AMNH). Utah: Cache Co.: 5 mi NE of Logan, Green Canyon (trail), T12N R2E sec 21, 41.7616 ° N 111.729 ° W, 1905 m, 3 Jul 1982, M. D. Schwartz, Cercocarpus ledifolius (Rosaceae) , 13 (AMNH). Blacksmith Fork Canyon, on UT State route 101, T10N R3E, 41.6289 ° N 111.80274 ° W, 1646 m, 25 Jun 1981, M. D. Schwartz, Melilotus officinalis (Fabaceae) , 13, 1♀ (AMNH). Logan Canyon, 41.74028 ° N 111.79306 ° W, 9 Jul 1935, R. E. Nye, 13 (USU). Tony Grove Canyon, 41.89611 ° N 111.55278 ° W, 10 Jul 1975 - 16 Jul 1975, Knowlton and Hanson, 13 (USU). Carbon Co.: 8 mi NW of Helper, Price Canyon Recreation Area, T12E R9E, 39.76594 ° N 110.96028 ° W, 2438 m, 9 Jul 1982, M. D. Schwartz, Amelanchier utahensis (Rosaceae) , 23, 3♀ (AMNH). Pine Canyon, 16 miles NE of Wellington, T12E R12E, 39.70629 ° N 110.52206 ° W, 2286 m, [date unknown], M. D. Schwartz, Amelanchier utahensis (Rosaceae) , 23, 5♀ (AMNH). Garfield Co.: 14.3 mi S of Rt 95 on Rt 276, 3.4 mi N of Starr Springs Campground turnoff, 37.87632 ° N 110.56773 ° W, 1524 m, 19 Jun 1983, R. T. Schuh and M. D. Schwartz, Light Trap, 13, 3♀ (AMNH). Capitol Reef National Park, Grand Wash, Cobab Canyon Trail, 38.15 ° N 111.16667 ° W, 1827 m, 21 Jun 1983, R. T. Schuh, M. D. Schwartz, Cercocarpus ledifolius (Rosaceae) , 33 Amelanchier utahensis (Rosaceae) , 33, 14♀ (AMNH). Kane Co.: Mount Carmel Hwy, 37.24722 ° N 112.66361 ° W, 18 Jun 1935, G. F. Knowlton, snowball, 13 (USU). Rich Co.: Garden City, 41.88917 ° N 111.385 ° W, 7 Jul 1969, G. F. Knowlton, 23 (USU). Salt Lake Co.: Emigration Canyon, 40.75056 ° N 111.81028 ° W, 17 Jun 1932, O. H. Swezey, 13 (CAS). San Juan Co.: 2.7 mi W of Rt 95 on Rt 263, T37S R17E, Natural Bridges National Monument, 37.52902 ° N 110.04838 ° W, 1829 m, 18 Jun 1983, R. T. Schuh and M. D. Schwartz, Amelanchier utahensis (Rosaceae) , 83 (AMNH_ PBI 00095265), 34♀ (AMNH_PBI 00095269) (AMNH). Brushy Basin Rim Road, NW of Blanding, County road # 227 0.5 E mp 116, T37S R12E, 37.7419 ° N 109.55267 ° W, 1737 m, 12 Jun 1982, M. D. Schwartz, Purshia tridentata (Rosaceae) , 73, 14♀ Amelanchier utahensis (Rosaceae) , 1♀ (AMNH). Canyonlands National Park, Needles Overlook, T29S R20E Sec23 sw1/4 sw1/4, 38.2687 ° N 109.6877 ° W, 11 Jun 1982, M. D. Schwartz, Amelanchier utahensis (Rosaceae) , 13, 2♀ (AMNH). Grand Flat near Collins Canyon, 37.42167 ° N 110.16056 ° W, 1707 m, 1 Jun 1982, D. A. and J. T. Polhemus, 103, 7♀ Shepherdia rotundifola (Elaeagnaceae) , 1♀ (JTP); 26 May 1986, D. A. and J. T. Polhemus, 53, 7♀ (JTP). Summit Co.: Kamas, 40.64306 ° N 111.28 ° W, 4 Jul 1922, E. P. Van Duzee, 2♀ oak, 2♀ (CAS). Uintah Co.: 5–10 miles SW Bonanza, T10S R24E Sec14, 39.9456 ° N 109.1842 ° W, 1615 m, 4 Jun 1981, M. D. Schwartz, Amelanchier utahensis (Rosaceae) , 123, 9♀ (AMNH). Utah Co.: Mount Nebo Loop, 39.81239 ° N 111.76152 ° W, 3 Jul 1971, G. F. Knowlton, 13 (UCD). Provo Canyon, route 92, 1 mile N of Sundance Ski Resort, 40.31306 ° N 111.65583 ° W, 2438 m, 4 Jul 1982, M. D. Schwartz, Quercus gambelii (Fagaceae) , 133, 15♀ (AMNH_PBI 00095267) Quercus gambelii Nutt. (Fagaceae) , 133, 15♀ (AMNH). Wasatch Co.: Dock Flat, 1 mi NE of Rt 40, T28S R12W Sec 9, 40.32583 ° N 111.2525 ° W, 2438 m, 14 Aug 1982, M. D. Schwartz, Crataegus douglasii (Rosaceae) , 23, 5♀ (AMNH). Heber, 40.50689 ° N 111.41323 ° W, 5 Jul 1922, E. P. Van Duzee, 13, 6♀ Amelanchier sp. (Rosaceae) , 1♀ (CAS). Washington Co.: Saint George, 37.10417 ° N 113.58333 ° W, 4 Jun 1963, G. E. Bohart, Stanleya pinnata (Brassicaceae) , 103, 6♀ (USU). Zion National Park, 7 mi NE Rt 9 on Kolob Reserve Rd, 37.24861 ° N 112.95583 ° W, 1372 m, 25 May 1981, M. D. Schwartz, Quercus gambelii (Fagaceae) , 133 (AMNH_PBI 00095263), 5♀ Quercus gambelii Nutt. (Fagaceae) , 133, 5♀ Amelanchier utahensis (Rosaceae) , 183, 19♀ (AMNH). Washington: Klickitat Co.: Brooks Memorial State Park, 45.98778 ° N 120.67361 ° W, 18 Jun 1951, W. H. Lange, 13 (UCD).