Pusia paraleucozona ( Boettger, 1906 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 41-42

Pusia paraleucozona ( Boettger, 1906) nov. comb.

Figs 14A 1 –A 2 , B 1 –B 2

* Mitra (Uromitra) ebenus Lmk. var. paraleucozona n. var. — Boettger 1906: 8.

Vexillum (Uromitra) ebenus paraleucozona (Boettger) — Zilch 1934: 259, pl. 17, fig. 18.

Vexillum (Vexillum) paraleucozona ( Boettger, 1906) — Bałuk 1997: 35, pl. 9, fig. 6.

Type material. Lectotype: SMF 360345 (= SMF 12.2208 a), Senckenberg Museum, Frankfurt/Main, Germany, SL: 11.5 mm, MD: 5.0 mm, Coşteiu de Sus ( Romania), illustrated in Zilch (1934, pl. 17, fig. 18), figs 14B 1 –B 2 . Paralectotypes: SMF 360.2208 b–c, Senckenberg Museum , Frankfurt / Main, Germany , 17+ 7 specimens, Coşteiu de Sus ( Romania) .

Additional material. NHMW 1865/0036/0010, SL: 9.0 mm, MD: 3.9 mm, Drnovice u Vyškova ( Czech Republic), figs 14A 1 –A 2.

Revised description. Shell moderately large, broad fusiform, with distinctly incised suture, and broad conical to weakly cyrtoconoid spire. Protoconch unknown. Teleoconch of seven whorls. Spire whorls weakly convex, with periphery at abapical suture. Axial sculpture of orthocline, convex ribs, separated by interspaces of about same width (about 20–25 axial ribs on last whorl). Adapical tips of axial ribs may be slightly swollen resulting in slightly gradate spire. Last whorl strongly convex, rapidly contracting. Aperture wide, ovoid with indistinct anal sinus. Columellar callus indistinct. Columella with four prominent folds, weakening abapically. Outer lip thin, abapically contracting, about 10–12 lirae within. No spiral sculpture, except four to six spiral cords on weak fasciole, partly extending from columellar folds. Siphonal canal straight, narrow, moderately short, with shallow siphonal notch.

Shell measurements and ratios. SL: 9.0–12.0 mm, MD: 3.9–5.5 mm; AA: 45–55°, SL/MD: 2.3–2.4, AL/AW: 3.7, AH/S: 2.8–3.0.

Discussion. Boettger (1906) described this species as variety of “ Mitra ebenus ” [= Ebenomitra leucozona ( Andrzejowski 1830) ], which is clearly incorrect. Pusia paraleucozona has four prominent columellar folds and numerous delicate axial ribs, whereas E. leucozona has three folds and few blunt axial swellings. At first glance it is similar to Pusia moravica ( Hoernes & Auinger, 1880) in being broadly fusiform with narrow ribs persisting on all whorls, but that species is larger [reconstructed height about 13–16 mm vs 9.0–12.0 mm in Pusia paraleucozona ( Boettger, 1906) ], at the same size the axial ribs are wider spaced in P. moravica , and the spiral sculpture in the subsutural area is weak, but present, whereas in P. paraleucozona spiral sculpture is restricted to the siphonal fasciole.

Ebenomitra renauleauensis Landau, Ceulemans & Van Dingenen, 2019 , from the Tortonian of France, is reminiscent of P. paraleucozona but differs in its smaller size and the coarser axial ribs.

Palaeoenvironment. Unknown; probably offshore settings.

Distribution in Central Paratethys. Badenian (middle Miocene): North Alpine-Carpathian Foredeep: Drnovice u Vyškova ( Czech Republic); Korytnica Basin: Korytnica ( Poland) ( Bałuk 1997); Făget Basin: Coşteiu de Sus ( Romania) ( Boettger 1906).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 14 A1–A2. Pusia paraleucozona (Boettger, 1906), Drnovice u Vyškova (Czech Republic), NHMW 1865/0036/0010. B1–B2. Pusia paraleucozona (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360345. C1–C2. Pusia szobbiensis (Halaváts, 1884), lectotype, Inv. nr. M107, Szob (Hungary). D. Vexillum szobbiensis (Halaváts, 1884), paralectotype, Inv. nr. M107, Szob (Hungary). E1–E2. Vexillum pseudoschafferi Biskupič, 2020, holotype, Z 40033, Rohožník–Konopiská (Slovakia). E1–E2. Vexillum pseudoschafferi Biskupič, 2020, paratype, Z 40034, Rohožník–Konopiská (Slovakia). G. Vexillum pseudoschafferi Biskupič, 2020, paratype, Z 40035, Rohožník–Konopiská (Slovakia).