Pterostichus brevicornis brevicornis (Kirby, 1837)

Bousquet, Yves, 2012, Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico, ZooKeys 245, pp. 1-1722 : 544-545

Pterostichus brevicornis brevicornis (Kirby, 1837)

Argutor brevicornis Kirby, 1837: 31. Type locality: "Lat. 65° [= apparently region of Great Bear Lake, Northwest Territories]" (original citation), restricted to "Good Hope, N[orth]W[est] T[erritories]" by Ball (1966a: 105). Lectotype (♂), designated by Ball (1966a: 105), in BMNH.

Feronia subtilis R.F. Sahlberg, 1844: 35. Type locality: circa Okhotsk, Khabarovsk Kray, Siberia, Russia (inferred from introduction and title of the paper). Lectotype [as holotype] (♀), designated by Ball (1966a: 105), in ZMH. Synonymy established by Ball (1966a: 105).

Cryobius fastidiosus Mannerheim, 1853: 131. Type locality: "ad sinum Woskresenk [= Resurrection Bay] peninsulae Kenai [Alaska]" (original citation). Lectotype (♂), designated by Silfverberg (1987: 16), in ZMH. Synonymy established by Lindroth (1954b: 131).

Pseudocryobius quinquepunctatus Motschulsky, 1860: 93. Type locality: "Kamtschatka [Russia]" (original citation). One syntype in ZMMU (Keleinikova 1976: 213). Synonymy established by Ball (1966a: 105).

Feronia fragilis Mäklin, 1878: 20. Type locality: "Sopotschnaja Korga (71°40' n. lat.) [Siberia, Russia]" (original citation). Syntype(s) location unknown. Synonymy established by Ball (1966a: 105). Note. The lectotype selected by Ball (1966a: 105), apparently in ZMH though not listed in Silfverberg’s (1987) type list, is labeled “Waigatsch” (= Vaygach Island in Siberia) and is probably not a syntype.

Feronia infima Mäklin, 1878: 20 [primary homonym of Feronia infima Chaudoir, 1868]. Type locality: "Jenisej, nemligen åtminstone från Mesenkin åt söder ända nedom 61 breddgraden [Russia]" (original citation). Syntype(s) location unknown (possibly in NRSS). Synonymy established, under the name Pterostichus fastidiosus (Mannerheim), by Poppius (1906b: 192), confirmed by Lindroth (1954b: 132).

Feronia arctica J.R. Sahlberg, 1880: 31. Replacement name for Feronia infima Mäklin, 1878.

Feronia epipleuralis J.R. Sahlberg, 1885b: 49. Type locality: Port Clarence, Alaska (inferred from title of the paper). Syntype(s) [3 originally cited] in ZMH. Synonymy established, under the name Pterostichus fastidiosus (Mannerheim), by Poppius (1906b: 192), confirmed by Ball (1966a: 105).

Platysma aquilonium Tschitschérine, 1904: 125. Type locality: "insula Kolgujev: Bugrino-Stanovishtshe [in Barents Sea, Russia]" (original citation). Syntype(s) [2 ♂ originally cited] probably in ZILR. Synonymy established, under the name Pterostichus fastidiosus (Mannerheim), by Poppius (1906b: 192).

Cryobius carbo Poppius, 1906b: 182. Type locality not stated. Lectotype (♀), designated by Ball (1966a: 105), in ZMH. Synonymy established by Ball (1966a: 105).

Cryobius fastidiosus minusculus Poppius, 1906b: 199. Type locality: "Bulkur, Unterste Lena [Yakutia, Siberia, Russia]" (original citation). Lectotype (♀), designated by Ball (1966a: 105), in ZMH. Synonymy established by Ball (1966a: 105).

Cryobius delicatus Casey, 1918: 375. Type locality: "S[ain]t Paul Island, Alaska" (original citation). Lectotype [as holotype] (♂), designated by Ball (1966a: 89), in USNM [# 47079]. New synonymy. Note. Ball (1966a) retained this name for a subspecies of Pterostichus brevicornis endemic to the Bering Sea Islands but, as pointed out by himself (Ball 1966a: 121) and Lindroth (1966: 523), there are no constant structural character states between the two forms and intermediate specimens are known. Based on these facts, I prefer not to retain this form as a valid subspecies.

Distribution.

This Holarctic subspecies ranges from the Kola Peninsula in northern European Russia eastwards to Newfoundland (Ball 1966a: 119); the species is represented by isolated populations in the Gaspé Peninsula, Maine, New Hampshire, Vermont, New York, Michigan (Ball 1966a: 119-121), northeastern Minnesota (Cook County, CNC), and northern Wyoming (Big Horn County, FFPC) [see Ball and Currie 1997: Fig. 4]. The record from northern Colorado (Armin 1963: 217) is likely in error. Fossil remnants, older than 33,000 years B.P., have been unearthed in southwestern Ontario (Warner et al. 1988: 37); others from a Plio-Pleistocene sequence have been found in northwestern Greenland ( Böcher 1995: 28).

Records.

CAN: AB, BC, LB, MB, NF, NT, NU, ON, QC, YT USA: AK, ME, MI, MN, NH, NY, VT, WY - Holarctic

Note.

The subspecies Pterostichus brevicornis yasudai Morita is endemic to the island of Hokkaidō, Japan.