Pseudobiceros luteomarginatus ( Yeri & Kaburaki, 1918 )

Bahia, Juliana & Schrödl, Michael, 2016, Pseudobiceros wirtzi sp. nov. (Polycladida: Cotylea) from Senegal with revision of valid species of the genus, Zootaxa 4097 (1), pp. 101-117 : 103

Pseudobiceros luteomarginatus ( Yeri & Kaburaki, 1918)

Taxonomic remarks. This species is listed as synonymous to Pseudobiceros flavomarginatus by Faubel (1984). However, the original description ( Laidlaw 1902) divides Pseudoceros species in forms with a pair of penes or single penis, and P. flavomarginatus is under the species with single penis. This fact was already pointed out by Marcus (1950), who considered Pseudoceros flavomarginatus and luteomarginatus as separate species based on the number of gonopores and color information. The original Pseudoceros luteomarginatus description ( Yeri & Kaburaki 1918) stated clearly (p. 38; plate I, fig. 5) that this Pseudocerotidae has two male gonopores and smooth dorsal surface. So we think that the synonym and new combination presented by Faubel (1984) was result of some confusion about the literature consulted. Thus we argue that Pseudobiceros flavomarginatus should be left in Pseudoceros , as it has only one male gonopore, and that Pseudoceros luteomarginatus should be transferred to Pseudobiceros , as it has two male gonopores. Since the transference of P. luteomarginatus was indirectly done by Faubel (1984), we consider that he had established this new combination. It is, however, unclear why the species mentioned above were ignored from Newman & Cannon’s (1994, 1997) Pseudobiceros species list. Pseudoceros flavomarginatus was mentioned in the discussion of Pseudobiceros periculosus ( Newman & Cannon 1994) where it was wrongly stated that Laidlaw did not give any additional details about gonopores.