Protogoerius, COIFFAIT, 1956

SMETANA, ALEŠ & DAVIES, ANTHONY, 2000, Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae), American Museum Novitates 3287, pp. 1-1 : 1-

publication ID

https://doi.org/ 10.1206/0003-0082(2000)287<0001:ROTNTT>2.0.CO;2

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https://treatment.plazi.org/id/0384878F-D800-FB09-FEEE-68C50D9AFB94

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scientific name

Protogoerius
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PROTOGOERIUS COIFFAIT, 1956 View in CoL

P. brullei (nom. nov.) – Pa

P. brachypterus (Brulle´, 1839) (nec Geoffroy, 1785; nec Marsham, 1802)

OCYPUS LEACH, 1819 (stat. nov.) Subgenus Ocypus Leach, 1819 ( Goerius Westwood, 1827 ) ( Xanthocypus J. Müller, 1925 )

O. affinis Wollaston, 1864 – Pa

O. alfierii (Bernhauer, 1925) – Pa

O. auroguttatus (Cameron, 1932) – Pa, Or – (comb. nov.) (from Nudabemus )

O. baronii (Coiffait, 1982) – Pa – (comb. nov.) (from Nudabemus )

O. caerulescens (Coiffait, 1982) – Pa – (comb. nov.) (from Nudabemus )

O. cameroni (nom. nov.)

O. bimaculatus (Cameron, 1932) (nec Schrank, 1798; nec Gravenhorst, 1802; nec Marsham, 1802) (comb. nov.) (from Staphylinus )

O. curtipennis Motschulsky, 1849 – Pa

O. integer Abeille, 1900

O. gridellii (J. Müller, 1924)

O. libanoticus (J. Müller, 1950) (as subsp. of O. gridellii ) (syn. nov.)

O. kyproticus Coiffait, 1964 (as subsp. of O. gridellii )

O. pseudolens Coiffait, 1964 (as subsp. of O. gridellii )

O. cyaneopubens (Reitter, 1913) – Pa

O. festae (J. Müller, 1926) – Pa

O. miwai (Bernhauer, 1943) – Pa

O. olens (O. Müller, 1764) – Pa, Ne

O. olens olens (O. Müller, 1764)

O. olens azoricus (Méquignon, 1942)

O. ophthalmicus (Scopoli, 1763) – Pa

O. ophthalmicus ophthalmicus (Scopoli, 1763)

O. cyanochloris Hochhuth, 1849 O. baicalensis (Eppelsheim, 1887) (as variety of O. ophthalmicus )

O. ophthalmicus atrocyaneus (Fairmaire, 1860)

O. ibericus Coiffait, 1964 (as subsp. of O. ophthalmicus )

O. pernigrus Coiffait, 1964 (as subsp. of O. ophthalmicus ) (syn. nov.)

O. ophthalmicus hypsibatus (Bernhauer, 1899)

O. ophthalmicus balcanicus (J. Müller, 1923) O. ophthalmicus balearicus (J. Müller, 1926) O. ophthalmicus rodopensis Coiffait, 1971 O. ophthalmicus benoiti Drugmand, 1998 O. ophthalmicus brigitteae Drugmand, 1998

O. quadrimaculatus (Cameron, 1932) – Pa, Or – (comb. nov.) (from Nudabemus )

O. rhaeticus Eppelsheim, 1873 – Pa

O. wasmanni (Bernhauer, 1920) – Pa – (comb. nov.) (from Nudabemus )

O. weisei Harold, 1877 – Pa – (comb. nov.) (from Agelosus )

Subgenus Matidus Motschulsky, 1860 (stat. nov.)

O. albanicus (J. Müller, 1943) – Pa

O. alpestris (Erichson, 1840) – Pa

O. bernhaueri (J. Müller, 1925) – Pa

O. biharicus (J. Müller, 1926) – Pa

O. brenskei (Reitter, 1884) – Pa

O. bisetosus (J. Müller, 1943) (as form of O. brenskei ) (syn. nov.)

O. holosetosus (J. Müller, 1943) (as subsp. of O. brenskei ) (syn. nov.)

O. brevipennis (Heer, 1839) – Pa

O. brevipennis brevipennis (Heer, 1839)

O. brevipennis pseudoalpestris (J. Müller, 1926)

O. brunnipes (Fabricius, 1781) – Pa O. brunnipes brunnipes (Fabricius, 1781) O. brunnipes intermedius (J. Müller, 1911)

O. caroli (Jarrige, 1943) – Pa

O. cerceticus Coiffait, 1964 – Pa

O. chevrolatii (Baudi, 1848) – Pa

O. alsaticus Coiffait, 1964 (as subsp. of O. chevrolatii ) (syn. nov.) O. pseudobrevipennis Coiffait, 1964 (as subsp. of O. chevrolatii ) (syn. nov.) O. vesubiensis Coiffait, 1964 (as subsp. of O. chevrolatii ) (syn. nov.)

O. zimmermanni Korge, 1965 (as subsp. of O. chevrolatii ) (syn. nov.)

O. coreanus (J. Müller, 1925) – Pa

O. deuvei Coiffait, 1978 – Pa

O. forficularius (Motschulsky, 1860) – Pa O. korgei Smetana, 1965

O. frater Smetana, 1965 – Pa

O. heinzi Smetana, 1965 – Pa

O. hissaricus Dvořák, 1984 – Pa

O. hochhuthi Eppelsheim, 1878 – Pa

O. italicus (Aragona, 1830) – Pa

O. italicus (Erichson, 1840) (syn. nov.) O. garganicus Fiori, 1894 (syn. nov.) O. silensis Fiori, 1894 (as variety of O. italicus ) (syn. nov.)

O. mirtensis J. Müller, 1943 (as subsp. of O. italicus ) (syn. nov.)

O. khnzoriani Coiffait, 1967 – Pa

O. kuntzeni (J. Müller, 1926) – Pa

O. longimanus Smetana, 1965 – Pa

O. macrocephalus (Gravenhorst, 1802) – Pa

O. manceps Smetana, 1965 – Pa

O. matilei Jarrige, 1971 – Pa

O. iranicus (Smetana, 1971)

O. megalocephalus (Nordmann, 1837) – Pa

O. milleri Quedenfeldt, 1882 – Pa

O. milleri milleri Quedenfeldt, 1882 O. milleri robustus Coiffait, 1964

O. nitens (Schrank, 1781) (stat. nov.)

O. similis (Fabricius, 1792) (misidentification, nec Paykull, 1789)

O. nero (Faldermann, 1835) (syn. nov.) O. nitens nitens (Schrank, 1781)

O. nitens semialatus (J. Müller, 1904) O. nitens grigiensis (Reitter, 1918)

O. nitens ochropus (J. Müller, 1950)

O. nubigena Smetana, 1965 – Pa

O. ormayi (Reitter, 1887) – Pa

O. ottomanus Fauvel, 1900 – Pa

O. pedemontanus (J. Müller, 1924) – Pa

O. pedemontanus pedemontanus (J. Müller,

1924)

O. pedemontanus pyrenaeus (J. Müller,

1924)

O. pedemontanus cantabricus (J. Müller,

1926) O. ponticus Smetana, 1968 – Pa O. pullus Hochhuth, 1849 – Pa O. quadraticeps (Ménétriés, 1832) – Pa O. reimoseri (Bernhauer, 1906) – Pa O. simulator Eppelsheim, 1878 – Pa O. solarii (J. Müller, 1923) – Pa O. syriacus Baudi, 1848 – Pa

O. syriacus syriacus Baudi, 1848

O. syriacus primigenius (J. Müller, 1923) O. tenebricosus (Gravenhorst, 1846) – Pa O. depolii (J. Müller, 1924) O. torvus Smetana, 1965 – Pa O. trapezensis Coiffait, 1964 – Pa O. turcicus (Bernhauer, 1923) – Pa

Subgenus Pseudocypus Mulsant and Rey, 1876 (stat. nov.) ( Protocypus J. Müller, 1923 ) ( Atlantogoerius Coiffait, 1956 ) ( Fortunocypus Coiffait, 1964 ) ( Nudabemus Coiffait, 1982 )

O. addendus (Lindberg, 1953) – Pa – (comb. nov.) (from Pseudocypus )

O. aeneocephalus (DeGeer, 1774) – Pa, Ne – (comb. nov.) (from Pseudocypus )

O. aereus Cameron, 1928 – Pa

O. aethiops (Waltl, 1835) – Pa – (comb. nov.) (from Pseudocypus )

O. luigionii (J. Müller, 1926) (as subsp. of O. aethiops ) (comb. nov., syn. nov.) (from Pseudocypus )

O. almensis Coiffait, 1967 – Pa

O. angustulus Eppelsheim, 1888 – Pa

O. anophthalmicus (Hernandéz and Aguiar, 1988) – Pa – (comb. nov.) (from Pseudocypus )

O. apterus (Scheerpeltz, 1976) – Pa – (comb. nov.) (from Staphylinus )

O. canariensis Gemminger and Harold, 1868 – Pa – (comb. nov.) (from Pseudocypus )

O. densissimus (Bernhauer, 1933) – Pa – (comb. nov.) (from Staphylinus )

O. dorsalis Sharp, 1889 – Pa

O. eppelsheimi Reitter, 1887 – Pa – (comb. nov.) (from Pseudocypus )

O. excisus (J. Müller, 1950) – Pa – (comb. nov.) (from Pseudocypus )

O. libanicus Coiffait, 1956 (as subsp. of O. picipennis ) (comb. nov.) (from Pseudocypus )

O. fortunatarum Wollaston, 1871 – Pa – (comb. nov.) (from Pseudocypus )

O. fujiensis Naomi, 1992 – Pa

O. fulvipennis Erichson, 1840 – Pa – (comb. nov.) (from Pseudocypus )

O. fulvotomentosus Eppelsheim, 1889 – Pa

O. hauseri (Bernhauer, 1933) (comb. nov., syn. nov.) (from Staphylinus )

O. fuscatoides (Coiffait, 1964) – Pa – (comb. nov.) (from Pseudocypus )

O. fuscatus (Gravenhorst, 1802) – Pa – (comb. nov.) (from Pseudocypus )

O. abbreviatus (Motschulsky, 1858) (comb. nov.) (from Pseudocypus )

O. fuscoaeneus Solsky, 1871 – Pa – (comb. nov.) (from Pseudocypus )

O. graeseri Eppelsheim, 1887 – Pa – (comb. nov.) (from Pseudocypus )

O. graeseri graeseri Eppelsheim, 1887 O. graeseri brunneopubens (J. Müller, 1926) (comb. nov.) (from Pseudocypus )

O. hagai Naomi, 1992 – Pa

O. hakusanus Naomi, 1992 – Pa

O. helleni (J. Müller, 1926) – Pa – (comb. nov.) (from Pseudocypus )

O. almorensis (Cameron, 1932) – Pa – (comb. nov., syn. nov.) (from Pseudocypus )

O. himalayicus (Cameron, 1935) – Pa – (comb. nov., syn. nov.) (from Staphylinus )

O. hidanus Watanabe, 1987 – Pa

O. himalaycus (Coiffait, 1982) – Pa – (comb. nov.) (from Pseudocypus )

O. impennis (Fauvel, 1882) – Pa – (comb. nov.) (from Staphylinus )

O. inexspectatus Eppelsheim, 1887 – Pa – (comb. nov.) (from Pseudocypus )

O. iyonus Naomi, 1992 – Pa

O. izayoi Naomi, 1992 – Pa

O. jeannei Coiffait, 1980 – Pa

O. kendappa Naomi, 1992 – Pa

O. kiimontanus Naomi, 1992 – Pa

O. kisonus Naomi, 1992 – Pa

O. lewisius Sharp, 1874 – Pa

O. kobensis Cameron, 1930 (syn. nov.)

O. mateui (Coiffait, 1954) – Pa – (comb. nov., stat. nov.) (from Pseudocypus )

O. auricomus (Lindberg, 1953) (nec Brulle´, 1842; nec Cameron, 1929) (comb. nov., syn. nov.) (from Pseudocypus )

O. montanus (Cameron, 1942) – Pa – (comb. nov.) (from Staphylinus )

O. mus (Brulle´, 1832) – Pa – (comb. nov.) (from Pseudocypus )

O. transadriaticus (J. Müller, 1926) (as subsp. of O. mus ) (comb. nov., syn. nov.) (from Pseudocypus )

O. tauricus (J. Müller, 1932) (as subsp. of O. mus ) (comb. nov.)

O. nemotoi Naomi, 1992 – Pa

O. nepalicus (Coiffait, 1981) – Pa – (comb. nov.) (from Pseudocypus )

O. nigroaeneus Sharp, 1889 – Pa

O. rambouseki (J. Müller, 1925)

O. obscuroaeneus Fairmaire, 1852 – Pa – (comb. nov.) (from Pseudocypus )

O. obscuroaeneus obscuroaeneus Fairmaire, 1852

O. algiricus (J. Müller, 1932) (as subsp. of O. obscuroaeneus ) (comb. nov.) (from Pseudocypus )

O. atlasicus (Coiffait, 1974) (as subsp. of O. obscuroaeneus ) (comb. nov., syn. nov.) (from Pseudocypus )

O. horsti (Coiffait, 1974) (as subsp. of O. obscuroaeneus ) (comb. nov., syn. nov.) (from Pseudocypus )

O. obscuroaeneus schatzmayri (J. Müller, 1923) (comb. nov.) (from Pseudocypus )

O. ohtsukaorum Naomi, 1992 – Pa

O. orientis (nom. nov.) – Pa

O. orientalis (Bernhauer and Schubert, 1914) (nec Motschulsky, 1858) (comb. nov., syn. nov.) (from Pseudocypus )

O. picipennis (Fabricius, 1792) – Pa – (comb. nov.) (from Pseudocypus )

O. picipennis picipennis (Fabricius, 1792) O. abbruzzensis (J. Müller, 1926) (as subsp. of O. picipennis ) (comb. nov.) (from Pseudocypus )

O. aprutianus (J. Müller, 1926) (as subsp. of O. picipennis ) (comb. nov.) (from Pseudocypus )

O. herzegovinensis (J. Müller, 1932) (as subsp. of O. picipennis ) (comb. nov., syn. nov.) (from Pseudocypus ) O. picipennis graecus (Scheerpeltz, 1958) (comb. nov., syn. nov.) (from Pseudocypus )

O. anatolicus (Coiffait, 1964) (as subsp. of O. picipennis ) (comb. nov.) (from Pseudocypus )

O. andorranus (Coiffait, 1964) (as subsp. of O. picipennis ) (comb. nov.) (from Pseudocypus )

O. pindensis (Coiffait, 1964) (as subsp. of O. picipennis ) (comb. nov.) (from Pseudocypus )

O. bulgaricus (Coiffait, 1970) (as subsp. of O. picipennis ) (comb. nov., syn. nov.) (from Pseudocypus )

O. rodopensis (Coiffait, 1971) (as subsp. of O. picipennis ) (comb. nov., syn. nov.) (from Pseudocypus )

O. ponticus (Coiffait, 1978) (as subsp. of O. picipennis ) (comb. nov., syn. nov.) (from Pseudocypus )

O. picipennis altaiensis (J. Müller, 1926) (comb. nov.) (from Pseudocypus )

O. picipennis barbarus (J. Müller, 1926) (comb. nov.) (from Pseudocypus )

O. marocanus (Coiffait, 1964) (as subsp. of O. picipennis ) (comb. nov., syn. nov.) (from Pseudocypus )

O. picipennis caucasicus (J. Müller, 1926) (comb. nov.) (from Pseudocypus )

O. picipennis fallaciosus (J. Müller, 1926) (comb. nov.) (from Pseudocypus ) O. maritimus Coiffait, 1956 (as subsp. of O. picipennis ) (comb. nov.)

O. picipennis nevadensis (J. Müller, 1926) (comb. nov.) (from Pseudocypus ) O. asturicus Coiffait, 1956 (as subsp. of O. picipennis ) (comb. nov.) (from Pseudocypus )

O. cerdanicus Coiffait, 1956 (as subsp. of O. picipennis ) (comb. nov.) (from Pseudocypus )

O. gallicus Coiffait, 1956 (as subsp. of O. picipennis ) (comb. nov.) (from Pseudocypus )

O. teruelensis (Coiffait, 1976) (as subsp. of O. picipennis ) (comb. nov., syn. nov.) (from Pseudocypus )

O. plagiicollis (Fairmaire, 1891) – Pa – (comb. nov.) (from Staphylinus )

O. sabi Naomi, 1992 – Pa

O. scutiger Sharp, 1889 – Pa

O. semenowi Reitter, 1887 – Pa

O. septentrionalis Watanabe, 1984 – Pa

O. sericeicollis (Ménétriés, 1832) – Pa – (comb. nov., stat. nov.) (from Pseudocypus )

O. cupreus (Rossi, 1790) (nec Fourcroy, 1785) (comb. nov., syn. nov.) (from Pseudocypus )

O. angustatus Stephens, 1832 (comb. nov., syn. nov.) (from Pseudocypus )

O. confinis Stephens, 1932 (comb. nov., syn. nov.) (from Pseudocypus )

O. rossii (Jarrige, 1954) (comb. nov., syn. nov.) (from Pseudocypus )

O. fulvicupreus (Coiffait, 1956) (comb. nov., syn. nov.) (from Pseudocypus )

O. sericeomicans (Bernhauer, 1931) – Pa – (comb. nov.) (from Staphylinus )

O. serotinus (Adám, 1992) – Pa – (comb. nov.) (from Pseudocypus )

O. smetanai Naomi, 1992 – Pa

O. subaenescens Wollaston, 1864 – Pa – (comb. nov.) (from Pseudocypus )

O. svozili Dvořák, 1984 – Pa

O. sylvaticus Wollaston, 1865 – Pa – (comb. nov.) (from Pseudocypus )

O. umbricola Wollaston, 1864 – Pa – (comb. nov.) (from Pseudocypus )

O. yamato Naomi, 1992 – Pa

O. yoroi Naomi, 1992 – Pa

O. yuinus Naomi, 1992 – Pa

Subgenus Aulacocypus J. Müller, 1925

O. gloriosus Sharp, 1874 – Pa

O. kansuensis (Bernhauer, 1933) (comb. nov.) (from Staphylinus ) (see comment under Ocypus above)

O. parvulus Sharp, 1874 – Pa

O. subtilis Tikhomirova, 1973 – Pa

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

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