Architipula, Handlirsch, 1906

Kopeć, Katarzyna, Krzemiński, Wiesław, Skowron, Karina & Coram, Robert, 2017, The genera Architipula Handlirsch, 1906 and Grimmenia Krzemiński and Zessin, 1990 (Diptera: Limoniidae) from the Lower Jurassic of England, Palaeontologia Electronica 15 (32), pp. 1-7 : 2

Genus ARCHITIPULA Handlirsch, 1906

Synonymy. Liassotipula Tillyard, 1933 (new synonymy); Eoasilidea Bode, 1953 (synonymised by Krzemiński and Kovalev, 1988, p. 55-56); Protipula Handlirsch, 1906 (synonymised by Evenhuis, 1994, p. 62); Paratipula Bode, 1953 (synonymised by Evenhuis, 1994, p. 62).

Type species. Architipula seebachiana Handlirsch, 1906 (= Architipula seebachi [ Geinitz, 1884]), Toarcian (Lower Jurassic), Dobbertin in Mecklenburg, Germany.

Included species. Of the approximately 50 named species, the following, from the Lower Jurassic of Germany, can be confidently assigned to the genus Architipula : A. arculifera Bode, 1905 ; A. bodeisimilis Bode, 1953 ; A. formosa Bode, 1953 ; A. fragmentosa ( Bode, 1953) ; A. maior Handlirsch, 1939 ; A. stigmatica Handlirsch,1906 ; and A. latipennis Handlirsch, 1906 . Re-examination of the holotypes of other named species is required to establish membership of Architipula since their generic affiliation cannot be determined on the basis of existing descriptions and drawings. Architipula youngi Krzemiński from the Norian (Upper Triassic) of North America is the oldest representative of the genus ( Krzemiński, 1992a).

Revised diagnosis. Vein Sc ends after fork of vein Rs into R 2+3+4 and R 5; Rs long, subequal to or a little shorter than veins R 2+3 and R 3 combined; d-cell characteristically deeply indented in basal part ( Figures 1-2 ), upper part of d-cell 1.5-2x longer than its lower part (from the forking of Mb to cross-vein m-cu); cross-vein m-m between M 1+2 and M 3 distinctly inclined, its bottom part shifted towards the wing’s base (unlike in most Limoniidae ); vein M 3 usually distinctly sinuous; cross-vein m-cu situated close to origin of the fork of M 3+4 into M 3 and M 4; A 2 usually short and straight.

Figures

FIGURE 1. Wing venation of Architipula seebachi (Geinitz, 1884). SGWG 122/82, holotype of A. seebachiana Handlirsch, 1906.

FIGURE 2. Wing venation of Architipula seebachi (Geinitz, 1884). SGWG 119/6, holotype.