Promalactis lonchodes Wang, 2020

Wang, Shuxia & Liu, Chen, 2020, Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) IX. The maculosa species-group, with descriptions of eighteen new species, Zootaxa 4890 (1), pp. 38-66 : 43-44

Promalactis lonchodes Wang , sp. nov.

( Figs 5 , 23 , 38 )

Type material. CHINA, Guizhou: Holotype ♂, Dongdai (25.22°N, 108.03°E), Shuizu Town , Limingguan , Libo County, 720 m, 19.VII.2015, leg. MQ Yang & JE Lee, slide No. JYY17733 . Paratypes: 9♂, 6♀, same data as holotype, slide Nos. JYY17692 ♂, JYY17725 ♂, JYY17734 ♀, JYY17735 ♂, LC19427 ♀; 6♂, 9♀, Neila Village, Shuizu Town , Limingguan , Libo County, 800 m, 18–22.VII.2015, leg. MQ Yang & JE Lee, slide Nos. JYY17727 ♀, JYY17728 ♂, JYY17729 ♀; 2♂, 4♀, Pobao, Shuizu Town , Limingguan , Libo County, 740 m, 20.VII.2015, leg. MQ Yang & JE Lee, slide Nos. JYY17732 ♂, LC19474 ♀; 2♂, 3♀, Shuizu Town, Limingguan , Libo County, 820 m, 21.VII.2015, leg. MQ Yang & JE Lee, slide Nos. JYY17730 ♂, JYY17731 ♀; 2♀, Weng’ang Town , Libo County, 1345 m, 19.VII.2015, leg. JP Wan, slide No. LC19475 ; 1♀, Kuankuoshui , 1509 m, 30.VIII.2018, leg. ML Zheng et al., slide No. LC19216 .

Diagnosis. The new species can be distinguished from its congeners by the gnathos triangularly produced at the conjunction of the basal arm and the mesial plate. It is similar to P. diorbis Kim et Park, 2012 in the female genitalia, but can be distinguished by the lamella postvaginalis sub-semicircular and the antrum deeply incised from the middle of the posterior margin; in P. diorbis , the lamella postvaginalis is sub-trapezoidal and the antrum is deeply concave in the middle of the anterior margin ( Kim et al. 2012: 904, fig. 4D).

Description. Adult ( Fig. 5 ). Wingspan 9.0–12.0 mm. Head with vertex and occiput deep grey, frons grey. La-bial palpus with second segment brownish grey; third segment black except white at base and apex. Antenna with scape white dorsally, pale grey ventrally; flagellum black and white. Thorax deep grey, ochreous yellow apically; tegula deep grey basally, deep ochreous yellow distally. Forewing ochreous yellow; markings white edged with black scales: costal spot at basal 3/5, extending obliquely outward to above posterior angle of cell, with dense black scales forming an irregular black spot at outer margin and a small black spot at inner margin anteriorly; cell with a short stripe at basal 1/4; fold with a streak at base, a spot at middle and before distal 1/4 respectively; dorsum with three evenly spaced streaks reaching fold; apical spot ovate; terminal spot ovate, smaller, near apical spot; tornus with black scales diffused to costal spot, tinged with a few white scales; cilia ochreous yellow. Hindwing and cilia grey. Fore- and midlegs black, femora greyish brown ventrally, tibiae with a white spot at base and middle, with a tuft of white scales at apex dorsally, tarsi white at apices of basal two tarsomeres, white on entire apical tarsomere of midleg; hindleg brown ventrally, greyish brown dorsally, tarsus black except white at apices of basal four tarsomeres and on entire apical tarsomere.

Male genitalia ( Fig. 23 ). Uncus with basal 2/5 wide, subparallel laterally, narrowed from basal 2/5 to before apex; apex medially notched, forming small triangular lobes laterally. Gnathos almost as long as uncus; mesial plate triangular, obtuse at apex, produced triangularly at conjunction of basal arm and mesial plate; basal arm slender, longer than mesial plate. Tegumen slightly widened to middle, then divided; lateral arm narrowed to rounded apex. Valva wider at base, slightly narrowed to basal 2/3; distal 1/3 bilobed: dorsal lobe slender, clubbed, smooth, ventral lobe digitate, stronger and longer than dorsal lobe, setose on ventral margin; sacculus wide at base, narrowed to basal 3/5 length of valva. Saccus shorter than uncus, slightly narrowed to apex. Juxta narrowly connected at base, rounded on anterior margin, reaching before posterior margin of tegumen apically; lateral arm heavily sclerotized, widened to middle, then gradually narrowed to acute apex. Aedeagus straight, shorter than valva; cornutus 1/5 length of aedeagus, with more than six apical spines.

Female genitalia ( Fig. 38 ). Apophyses anteriores about 1/3 length of apophyses posteriores. Eighth sternal plate sub-semicircular, concave medially on posterior margin, with erect vertical flake at 1/3 length from each side. Antrum incised deeply from middle of posterior margin, infolded laterally. Ductus bursae about twice length of corpus bursae, posterior 3/4 inflated, anterior 1/4 narrow, with many spines in posterior 1/3. Corpus bursae membranous; two signa very large, round, with dense spicules.

Distribution. China (Guizhou).

Etymology. The specific epithet of the new species is derived from the Latin lonchodes , referring to the shape of the mesial plate of the gnathos.

Figures

FIGURES 1‒6. Adults of Promalactis spp. 1, P. coloristigmosa, paratype, ♂; 2, P. cruciate, holotype, ♂; 3, P. lateridentalis, holotype, ♂; 4, P. latispinata, holotype, ♂; 5, P. lonchodes, holotype, ♂; 6, P. magnispina, holotype, ♂. Scales = 2 mm.

FIGURES 19‒24. Male genitalia of Promalactis spp. 19, P. coloristigmosa, holotype, slide No. JYY17711, 19a, enlarged cornu- tus, slide No. JYY17804; 20, P. cruciate, holotype, slide No. HXM14127; 21, P. lateridentalis, holotype, slide No. JYY17706; 22, P. latispinata, holotype, slide No. LC19215, 22a, enlarged cornutus; 23, P. lonchodes, holotype, slide No. JYY17733, 23a, enlarged cornutus, slide No. JYY17725; 24, P. magnispina, holotype, slide No. JYY17718. Scales = 0.2 mm.

FIGURES 36‒41. Female genitalia of Promalactis spp. 36, P. coloristigmosa, paratype, slide No. LC19222; 37, P. cruciate, paratype, slide No. HXM14250; 38, P. lonchodes, paratype, slide No. JYY17727; 39, P. bomiensis, paratype, slide No. LC19306; 40, P. fasciserrata, holotype, slide No. JYY17767; 41, P. brevipalpa, paratype, slide No. HXM13120, 41a, enlarged ductus bursae. Scales = 0.2 mm.