Promalactis cribrata Wang, 2019

Wang, Shuxia & Jia, Yanyan, 2019, Taxonomic study of the genus Promalactis Meyrick, 1908 (Lepidoptera: Oecophoridae) IV. The sakaiella species group, with descriptions of fifteen new species, Zootaxa 4563 (3), pp. 491-515 : 504-506

Promalactis cribrata Wang , sp. nov.

( Figs 4 , 29 , 39 )

Type material. MALAYSIA: Holotype ♂, Mt. Trusmadi (5.44°N, 116.45°E), Sabah, 1223 m, 6.II.2018, coll. Mujie Qi, slide Nos. JYY17647 ♂. Paratypes (10♂, 8♀): 9♂, 8♀, 6–11.II.2018, other same data as holotype, slide Nos. JYY17637 ♂, JYY17646 ♀, JYY17648 ♀, JYY17665 ♀; 1♂, Native mountain, Kundasang Golf Course, Mt. Kinabalu , Sabah, 1500 m, 17–20.V.1989, coll. K. R. Tuck, slide No. BMNH-33577 ( BMNH) .

Diagnosis. This species is more similar to P. oliviformis sp. nov. in the male genitalia. It can be distinguished by the valva deeply concave inward ventrodistally, and the costa with a thorn at distal 1/3; in P. oliviformis sp. nov., the valva is obliquely straight ventrodistally, and the costa lacks a thorn at basal 2/3. Further differentiation with other similar species can be found in the diagnosis of the preceding species.

Description. Adult ( Fig. 4 ) wingspan 13.0 mm. Head with vertex snowy white, frons and occiput dark brown. Labial palpus with second segment yellow on inner surface, basal 2/3 black and distal 1/3 yellow on outer surface; third segment approximately as long as second segment, basal 1/5 white, distal 4/5 black, ventral surface with a white dot at distal 1/3. Antenna with scape white; flagellum with basal several flagellomeres white, remaining flagellomeres white annulated with black on dorsal surface, dark brown on ventral surface. Thorax and tegula shinning blackish brown. Forewing deep ocherous yellow, black along basal 2/3 of costal margin; dorsum with two spots: basal spot sub-elliptical, blackish brown mixed with black scales, edged with a white stripe along outer margin, distal spot reaching middle of cell, sub-triangular, deep ocherous brown mixed with black scales, edged with a stright white streak along inner margin and crossing fold anteriorly; postmedian fascia deep grey, from basal 3/5 of costal margin extending straightly to tornus, narrowed medially, its outer margin with a diffused white spot anteriorly; distal patch deep grey mixed with black, extending along termen, sometimes meeting postmedian fascia at tornus; cilia yellow around apex and along termen, deep grey along distal part of costal margin and dorsum. Hindwing and cilia deep grey. Foreleg black, tarsus white at apex of each tarsomere on outer side; midleg black, femur with a longitudinal white stripe on ventral side; hindleg with femur grey, tibia and tarsus yellow on ventral side, grey on dorsal side.

Male genitalia ( Fig. 29 ): Uncus broad U-shaped, joined by a wide band basally; lateral arms curved, extending upward to rounded apex. Gnathos with lateral arms slender, slightly curved, narrowed to pointed apex; median process cylindrical, thick and straight, as long as lateral arms. Tegumen broad, sclerotized along anterior margin, concave in triangle anteromedially. Valva wide at base, slightly narrowed to distal 1/4, with dense spines in distal half roughly grouped in two rows: one row from distal 2/5 to apex below costa, another row along distal half above ventral margin; distal 1/4 with ventral margin deeply concave inward, forming two processes: dorsal process elongate triangular, narrowly rounded at apex, curved downward, ventral process being a small or large spine. Costa with a large sub-triangular basal process extending obliquely inward and crossing inner margin of vinculum, with a thorn at distal 1/3 that has a round cribiform base bearing dense spines. Sacculus broad at base, abruptly narrowed to aproximately middle of valva ventrally. Saccus wide at base, narrowed to a rounded apex. Juxta small Y shaped; lateral process ovate, apically with a tuft of strong spines of varied length; basal lobe slender, clubbed. Aedeagus slender, slightly curved, pointed at apex, approximately as long as valva; cornutus a tiny spine set before apex.

Female genitalia ( Fig. 39 ): Apophyses anteriores about half length of apophyses posteriores. Ostium bursae small, sclerotized and hood-like posteriorly. Ductus bursae thick, almost uniform, more than twice length of corpus bursae. Corpus bursae round; signum being a large spine arising from middle of a rhombic basal plate, with a smaller denticle near its base.

Distribution. Malaysia.

Etymology. This epithet refers to the cribiform base of the thorn at distal 1/3 of the costa.

Figures

FIGURES 1–4. Adults of Promalactis spp. 1. P. apicidigitata sp. nov., holotype, ♂; 2. P. biprocessa sp. nov., holotype, ♂; 3. P. bicornea sp. nov., holotype, ♂; 4. P. cribrata sp. nov., holotype, ♂. Scales =1.0 mm

FIGURES 27–31. Male genitalia of Promalactis spp. 27. P. grandilobata sp. nov., holotype, slide No. DZH12117; 28. P. arciformis sp. nov., holotype, slide No. JYY17636; 29. P. cribrata sp. nov., paratype, slide No. BMNH-33577; 30. P. bornensis sp. nov., holotype, slide No. MNHU-NK011; 31. P. oliviformis sp. nov., holotype, slide No. BMNH-33555. Scales =0.2 mm, 29a, b =0.1mm (not available for No. 30).

FIGURES 37‒40. Female genitalia of Promalactis spp. 37. P. bicornea sp. nov., paratype, slide No. JYY17547; 38. P. arciformis sp. nov., paratype, slide No. JYY17635; 39. P. cribrata sp. nov., paratype, slide No. JYY17665; 40. P. bornensis sp. nov., paratype, slide No. MNHU-NK009. Scales =0.2 mm (not available for No. 40).