Prismognathus kanghianus ( Didier & Séguy, 1953 ), Didier & Seguy, 1953

Huang, Hao & Chen, Chang-Chin, 2012, A review of the genera Prismognathus Motschulsky and Cladophyllus Houlbert (Coleoptera: Scarabaeoidea: Lucanidae) from China, with the description of two new species, Zootaxa 3255, pp. 1-36 : 14-15

Prismognathus kanghianus ( Didier & Séguy, 1953)

( Figs. 69 , 101 , 126 , 154 , 190 , 214 , 235 )

Eligmodontus kanghianus Didier & Séguy, 1953: 44 [type locality: Chapa (northern Vietnam)]; Ikeda 1987: 29, Fig. 6 , male from Vietnam; Mizunuma & Nagai 1994: plate 155, Fig. 222 , male from Tonkin.

Prismognathus kanghianus: Nagai 2005: 36 , new combination, Fig. 18 , male from Sapa, northern Vietnam; Fujita 2010: plate 101, Figs. 507 (1–4), males and female from northern Vietnam.

Prismognathus kanghianus kittii: Nagai 2005: 37 [type locality: Chudo Razi, northeastern Myanmar], Fig. 19 , holotype male. Valid subspecies.

Material examined. CHINA: Yunnan: 1 3 (CLW), Honghe, Gejiu City, Hekou County, V.2007; 1 Ƥ (CLW), Honghe, Gejiu City, Hekou County, IX.1999.

Identification. This species was originally described from northern Vietnam in the genus Eligmodontus (type species E. arcuatus Houlbert ). There is no confusion in identifying this species as only one P. arcuatus -like species has been known from the area around the type locality in northern Vietnam and southern Yunnan. Fujita (2010) recorded and illustrated the female for the first time.

The male from southern Yunnan examined by us matches the original description of P. kanghianus kanghianus . The female from the same locality of the male examined by us was not collected together with the male; it was identified as P. kanghianus kanghianus because it has different genitalia from the other female Prismognathus from the same area. Therefore the correct identification of the female of P. kanghianus kanghianus needs to be confirmed.

Nagai (2005) described P. kanghianus kittii from northern Myanmar which differs from P. kanghianus kanghianus by the smaller eyes and the lateral angles of pronotum more sharply protruding.

Distribution. Southern Yunnan (Hekou); northern Vietnam, northeastern Myanmar.

Figures

FIGURE 235. Distribution of the other Chinese Prismognathus.

FIGURES 57 – 72. Head, pronotum and proleg of male in dorsal view. 57 – 60 — P. alessandrae; 57 — Weixi, Yunnan; 58 — Muli, Sichuan; 59 — Zhongdian, Yunnan, 60 — Pantiange, Weixi, Yunnan; 61 – 64 — P. shani; 61, 63 — paratypes, Siping, Yingjing, Sichuan; 62 — paratype, Erlangshan, Sichuan; 64 — holotype, Erlangshan; 65 — P. arcuatus, Siping, Sichuan; 66 — P. haojiani, holotype, Fanjingshan, Guizhou; 67 — P. sukkitorum, Tengchong, Yunnan; 68 — P. prossi, Siping, Sichuan; 69 — P. kanghianus kanghianus, Hekou, Yunnan; 70 – 71 — P. triapicalis ssp.; 70 — Dayaoshan, Guangxi; 71 — Fengyangshan, Zhejiang; 72 — P. prossi, Maoershan, Guangxi. Scale bar 10 mm.

FIGURES 95 – 110. Head, pronotum and proleg of female in dorsal view. 95 — P. arcuatus, Siping, Sichuan; 96 — P. haojiani, paratype, Fanjingshan, Guizhou; 97 — P. sukkitorum, Tengchong, Yunnan; 98 — P. dauricus, Liaoning; 99 – 100 — P. p ro s s i; 99 — Fanjingshan, Guizhou; 100 — Maoershan, Guangxi; 101 — P. kanghianus kanghianus, Hekou, Yunnan; 102 — P. formosanus, Taiwan; 103 – 105 — P. triapicalis subspecies; 103 – 104 — Dabashan, Chongqing; 105 — Fengyangshan, Zhejiang; 106 — P. triapicalis triapicalis, Erlangshan, Sichuan; 107 — P. s i n e n s i s, holotype, Kuatun; 108 – 110 — P. davidis davidis; 108 — Dabashan, Chongqing; 109 – 110 — Siping, Sichuan. Scale bar 10 mm.

FIGURES 111 – 131. Female canthus (not under the same scale). 111 — P. s h a n i, paratype, Labahe, Sichuan; 112 — P. nigricolor, paratype; 113 — P. arcuatus, Siping, Sichuan; 114 – 116 — P. p ro s s i; 114 — Siping, Sichuan; 115 — Fanjingshan, Guizhou; 116 — Maoershan, Guangxi; 117 – 119 — P. davidis davidis; 117 — Siping, Sichuan; 118 — Tianmushan, Zhejiang; 119 — Dabashan, Chongqing; 120 – 123 — P. triapicalis; 120 — Sichuan; 121 — Dabashan, Chongqing; 122 — Fengyangshan, Zhejiang; 123 — Dabashan, Chongqing; 124 — P. haojiani, paratype, Fanjingshan; 125 — P. dauricus, Liaoning; 126 — P. kanghianus kanghianus, Hekou, Yunnan; 127 — P. alessandrae, Zhongdian, Yunnan; 128 — P. sinen sis, holotype, Kuatun, Fujian; 129 — P. sukk ito rum, Tengchong, Yunnan; 130 — P. formosanus, Taiwan; 131 — P. siniaevi, Gongshan, Yunnan.

FIGURES 150 – 167. Aedeagus in ventral view. 150 – 151 — Prismognathus arcuatus, Sichuan; 152 – 153 — P. haojiani; 152 — holotype, Fanjingshan; 153 — paratype, Leigongshan, Guizhou; 154 — P. kanghianus kanghianus, Hekou, Yunnan; 155 — P. sukkitorum, Tengchong, Yunnan; 156 – 157 — P. triapicalis triapicalis, Sichuan; 158 – 161 — P. triapicalis subspecies; 158 – 159 — Dabashan, Chongqing; 160 — Fengyangshan, Zhejiang; 161 — Dayaoshan, Guangxi; 162 – 163 — P. prossi; 162 — Siping, Sichuan; 163 — Fanjingshan, Guizhou; 164 – 165 — P. siniaevi; 164 — Jinping, Yunnan; 165 — Gongshan, Yunnan; 166 — Tetrarthrius sukkiti, Gongshan, Yunnan; 167 — Cyclorasis fukinukii, Motuo, Tibet. Scale bars 1 mm for all.

FIGURES 168 – 199. Flagellum of the median lobe. 168 – 171 — P. davidis davidis; 168 — Qinling, Shaanxi; 169 — Erlangshan, Sichuan; 170 — Tianmushan, Zhejiang; 171 — Dabieshan, Anhui; 172 — P. davidis cheni, Taiwan; 173 — P. nigricolor, Gongshan, Yunnan, same specimen as in Fig. 226; 174 – 178 — P. klapperichi; 174 — Simianshan, Chongqing; 175 — Wuyishan, Fujian; 176 — Maoershan, Guangxi; 177 — Nanling, Guangdong; 178 — Baishanzu, Zhejiang; 179 — P. dauricus, Liaoning; 180 — P. s h a n i, paratype, Siping; 181 — P. alessandrae, Zhongdian, Yunnan; 182 – 184 — P. p ro s s i; 182 — Siping, Sichuan; 183 — Fanjingshan, Guizhou; 184 — Maoershan, Guangxi; 185 — P. siniaevi, Gongshan; 186 – 187 — P. arcuatus, Sichuan; 188 – 189 — P. haojiani; 188 — holotype, Fanjingshan; 189 — paratype, Leigongshan, Guizhou; 190 — P. kanghianus kanghianus, Hekou, Yunnan; 191 — P. sukkitorum, Tengchong, Yunnan; 192 – 194 — P. triapicalis triapicalis, Sichuan; 195 – 196 — P. triapicalis subspecies; 195 — Dabashan, Chongqing; 196 — Fengyangshan, Zhejiang; 197 — P. formosanus, Taiwan; 198 — Tetrarthrius sukkiti, Gongshan, Yunnan; 199 — Cyclorasis fukinukii, Motuo, Tibet. Figs. 168 – 198 under the same scale. Scale bars 1 mm for all.

FIGURES 212 – 223. Female genitalia and the central protrusion of the 9 th tergites (at left bottom or right bottom). 212 — Prismognathus arcuatus, Sichuan; 213 — P. haojiani, paratype, Fanjingshan; 214 — P. kanghianus kanghianus, Hekou, Yunnan; 215 — P. sukkitorum, Tengchong, Yunnan; 216 – 217 — P. triapicalis triapicalis, Sichuan; 218 – 219 — P. triapicalis subspecies; 218 — Dabashan, Chongqing; 219 — Fengyangshan, Zhejiang; 220 — P. davidis davidis, Siping, Sichuan; 221 — P. siniaevi, Gongshan; 222 — P. prossi, Maoershan, Guangxi; 223 — P. formosanus, Taiwan. Scale bars 1 mm for all.

FIGURES 1 – 20. Cladophyllus, Tetrarthrius and Cyclorasis species. 1 – 4 — Cladophyllus oberthuri, male (Weixi, Yunnan); 5 – 8 — Cladophyllus bousqueti, male (Pianma, Yunnan); 9 – 12 — Cladophyllus nobuhikoi, male (Jinping, Yunnan); 13 – 14 — Cladophyllus oberthuri, female (Weixi, Yunnan); 15 – 16 — Cladophyllus bousqueti, female (Pianma, Yunnan); 17 – 18 — Tetrarthrius sukkiti, female (Motuo, Tibet); 19 – 20 — Cyclorasis nosei, female (Motuo, Tibet). 1, 5, 9, 13, 15, 17, 19 — head and pronotum in dorsal view; 2, 6, 10 — 9 th abdominal segment; 3, 7, 11 — aedeagus; 4, 8, 12 — flagellum; 14, 16, 18, 20 — female genitalia. Scale bars 5 mm for habitus, 1 mm for genitalia.