Prionocerus bicolor Redtenbacher, 1868

Geiser, Michael, 2010, Studies on Prionoceridae (Coleoptera: Cleroidea). II. A revision of the genus Prionocerus Perty, 1831, Zootaxa 2328, pp. 1-48 : 22-34

Prionocerus bicolor Redtenbacher, 1868

Prionocerus pertii Castelnau, 1836: 27 , submitted to ICZN (see Nomenclatural notes below). Prionocerus bicolor Redtenbacher, 1868: 109 .

Idgia (Prionocerus) bicolor var. notaticollis Pic, 1910: 53 . Synonymised by Champion, 1919: 329.

Type material examined: P. b i c o l o r: Holotype Ƥ "Novara 1857–59 Reise"; " Prionocerus bicolor Redtb. Type! (Novara.)"; "Q"; NHMW, in good condition.

P. b i c o l o r var. notaticollis : 2 syntypes 3 "Sumatra (Rouyer)"; MNHP, in good condition. P. p e r t i i: The type specimen (or type series) of this species is not present at the NHML, and during a visit to MNHP, it could not be found either. As the Castelnau collection was distributed over many collectors and museums, the location of this type is entirely unknown at the moment. Unfortunately, it is not impossible that it has been destroyed or lost over the time.

Other material examined: INDIA: " India " 1 3 ( NHML); " India, Bowring 63 47*" 2 Ƥ ( NHML); "Assam, Coll. Lange, coll. Staudinger" 1 Ƥ ( SMTD); "Ober-Assam, Hartert S." 1 3 2 Ƥ ( MNHB); „Malabar, Nicol, coll. Melly“ 1 Ƥ ( MHNG). KERALA (?): "Munaar, S. India, C. E. Tottenham collection, B. M. 1975– 587" 2 Ƥ ( NHML). UTTAR PRADESH: "Allahabad, Bowring, 63 47*" 1 Ƥ ( NHML). WEST BENGAL: "Darj., Ribbe 79, Darjeeling, dedit Ribbe 1879" 1 Ƥ ( MCGD); "Darj., Himalaya Ribbe" 2 Ƥ ( MNHB); "Darj., Darjeeling" 1 Ƥ ( MNHB); "Nurbong, Darjeeling, 950–2240 ft.–14. H. Stevens, B. M. 1922–307" 1 Ƥ ( NHML); Distr. Darjeeling, Kalimpong, Yok Forest, 1200 m, 25. IV. 1991, N. Dangal leg. 2 3 ( NHMB); 12 km S Kalimpong, 27°05' N / 88°34' E, an Vitex negundo, 16.–17. VI. 2005, T. Osten leg. 7 3 3 Ƥ ( SMNS); Distr. Darjeeling, Kalimpong, Khani, 1200 m, 22.–28. IV. 1987, Narayan Dangal leg. 1 Ƥ ( NHMB); Distr. Darjeeling, Kalimpong, Khani-Busty, 900 m NN, 23. IV. 1991, N. Dangal leg. 8 3 4 Ƥ ( NKME), 1 3 1 Ƥ (CMG); Distr. Darjeeling, Kalimpong, Dumra, 1000 m NN, 29. III. 1991, N. Dangal leg. 1 3 2 Ƥ ( NKME); Distr. Darjeeling, Kalimpong, Vati-Dara, 900 m NN, 12. IV. 1988, N. Dangal leg. 6 3 1 Ƥ ( NKME); Distr. Darjeeling, Kalimpong, Seng-Say Forest, 1500 m, 7. IV. 1991, N. Dangal leg. 1 Ƥ ( NKME); Distr. Darjeeling, Kurseong, Golma Forest, 1300 m NN, 22. VII. 1987, N. Dangal leg. 1 Ƥ ( NKME); Gopaldhara, Rungbong Valley, 26. III. 1914, H. Stevens leg., Stevens Coll. B. M. 1916–218. 1 Ƥ ( NHML); "Gopaldhara, Rungbong Vall., H. Stevens, 1916–218" 14 Ƥ 6 3 ( NHML). SIKKIM: "Sikkim, India, coll. Staudinger" 1 3 ( SMTD); "Sikkim" 1 3 ( NHMB), 1 3 ( MNHP). ASSAM: "Margherita, Assam, IV. & V. 1889, Brit. Mus. 1923–320" 1 Ƥ ( NHML); "Jobat, Assam, H. E. Andrewes Bequest, B. M. 1922–221" 2 Ƥ ( NHML); "Assam, Sudiya, Fry Coll., 1905.100" 2 3 ( NHML). MEGHALAYA: "Assam, Patkai Mts., Doherty, Fry Coll., 1905.100" 1 3 ( NHML); "Khasia Hills, Godwin Austen, 96–135." 1 3 1 Ƥ ( NHML); "Khasia Hills, Assam, Coll. Kraatz" 3 Ƥ ( DEIM); "Khasis, 1898, Coll. Kraatz" 1 Ƥ ( DEIM); "Khasi Hills, Assam, 1000–3000 ft., H. E. Andrewes Bequest, B. M. 1922–221" 1 Ƥ ( NHML); " India, 2000, Khasia Hills" 2 Ƥ ( MNHP); Khasi Hills, 10 km S Cherrapunji, 19. VI. 2005, T. Osten leg. 1 3 ( SMNS); SW of Cherrapunjee, 25°13'–14'N 91°40' E, 900 m, 1.– 24. V. 2005, L. Dembický leg., BMNH (E), 2006- 48 1 3 1 Ƥ ( NHML); SW of Cherrapunjee, 25°14' N / 91°40' E, 900 m, 21. V. 2007, L. Dembický leg. 1 3 ( NHMB); SW of Cherrapunjee, 25°13–14' N / 91°40' E, 900 m, 23–25. VI. 2007, P. Pacholátko leg. 1 3 ( NHMB); SW of Shillong, 25°34' N / 91°51'20'' E, 1600 m, 14. V. 2004, L. Dembický leg. 1 3 ( NHMB); W-Garo Hills, Bagmara, ca. 100 m, 25° 11.5' N 90° 38.5' E, 19.– 21. V. 1996, Jendek & Sausa leg. 1 Ƥ ( NHMW).

BANGLADESH: "Chandkhira, Sylhet, J. L. S., Brit. Mus. 1923–320" 4 Ƥ ( NHML); "Sylhet, Bowring, 63 47*" 4 Ƥ ( NHML); "Bengal, Chittagong, Hill Tracts, H. M. Parish, Brit. Mus. 1923–247" 1 Ƥ ( NHML); "Bengal, H. M. Parish, Brit. Mus. 1923–247" 1 Ƥ ( NHML).

NEPAL: W-Nepal, Parbat Distr., Kusma-Karkineta, 900–1600 m, 2. VII. 1986, J. Probst leg. 1 3 (NHMB); W-Nepal, Anklu-Khola, Arket-Sollentar, 1400–1610– 1000 m, 7. VI. 1990, Probst leg. 3 3 (NHMB); C-Nepal, Bagamati, Bhote-Koshi-Khola-Lamosangu, 700 m, 28. VI. 1987, Pasnang leg. 1 Ƥ (NHMB); Bagmati, Kabhre, Dhad Khola, 600 m, 31. V. 1989, M. Brancucci leg. 5 3 (NHMB); Bagmati, Sindhupalchok, Dhad Khola–Jarayetar, 600–800 m, 1. VI. 1989, M. Brancucci leg. 1 Ƥ (NHMB); Bagmati, Sindhupalchok, Dubhachaur–Sarmatang, 1600–2500 m, 3. VI. 1989, M. Brancucci leg. 1 3 (NHMB); C- Nepal, Bagmati Distrikt, Trisuli Bazar–Samri Bhanjyang, 540–1000 m, 8. VI. 1993, J. & J. Probst leg. 1 3 (NHMB); C-Nepal, Nuwakot Distr., Samri Bhanjyang–Tharpu, 1900–2000 m, 9. VI. 1993, Probst leg. 1 3 (NHMB); Z-Nepal, Kali Gandaki-Tal, Sikha, 2100 m, 4.–20. VI. 1977, D. Müting leg. 3 3 (NHMB); E Nepal, Arun valley, Tumlingtar-Sult., 750 m, 10. VI. 1988, J. Probst leg. 1 Ƥ (NHMB); E-Nepal, Arun V., Tumlingtar Khandbari, 450–1100 m, 27. V. 1983, M. Brancucci leg. 3 3 13 Ƥ (NHMB); E-Nepal, Arun V., Khandbari Arunthan 1100–1300 m, 29. V. 1983, M. Brancucci leg. 32 3 22 Ƥ (NHMB); O-Nepal, Khandbari–Arunthan, 1100–1300 m, 22. V. 1980, W. Wittmer leg. 10 3 10 Ƥ (NHMB); O-Nepal, Arunthan–Chichila, 1300–1950 m, 23. V. 1980, W. Wittmer leg. 1 Ƥ (NHMB); E-Nepal, Arun V., Arunthan–Chichila 1300–1950 m, 29. V. 1983, M. Brancucci leg. 1 3 (NHMB); E-Nepal, Arun V., Mure, 2000 m, 2.– 8. VI, 1983, M. Brancucci leg. 1 Ƥ (NHMB); O-Nepal, Num–Hedangna, 1500– 750–1100 m, 26. V. 1980, W. Wittmer leg. 1 Ƥ (NHMB); O- Nepal, Lamobagar Gao-Hedangna, 1000–1500– 1100 m, 4. VI. 1980, W. Wittmer leg. 1 Ƥ (NHMB); E-Nepal, Arun V., Arunthan–Tumlingtar, 1300- 450 m, 20. V, 1983, M. Brancucci leg. 2 Ƥ (NHMB); E-Nepal, Arun Valley, Pakhribas-Mongmaya, 1400– 350 m, 31. V. 1992, J. & J. Probst leg. 1 Ƥ (NHMB); E-Nepal, Arun Valley, Khandbari-Bhotebas, 950–1850 m, 5. VI. 1992, J. & J. Probst leg. 3 3 6 Ƥ (NHMB); E-Nepal, Arun Valley, Khandbari-Bhotebas, 400–1500 m, 1. VI. 1988, J. Probst leg. 3 3 3 Ƥ (NHMB); E-Nepal, Arun Valley, Mongmaya-Surtibari, 400 m, 1. VI. 1992, J. & J. Probst leg. 2 3 (NHMB); E-Nepal, Arun Valley, Mure-Num, 2050– 1600 m, 10. VI. 1992, J. & J. Probst leg. 1 Ƥ (NHMB); E-Nepal, Arun Valley, Num- Tashigaon, 1600– 900–1600 m, 10.–12. VI. 1992, J. & J. Probst leg. 3 3 1 Ƥ (NHMB); E-Nepal, Arun valley, Waleng Iswa-Khola–Waleng Siswa-Khola, 1200– 750 m, 15.–16. VI. 1992, J. & J. Probst leg. 1 3 1 Ƥ (NHMB); E-Nepal, Arun Valley, Dholikhorka-Mongmaya, 400–1250 m, 1. VI. 1988, J. Probst leg. 2 3 (NHMB); E-Nepal, Arun Valley, Dholikhorka-Mongmaya, 400–1250 m, 5. VI. 1988, J. Probst leg. 1 Ƥ (NHMB); Kosi # 4, Pakhribas, 27°03' N / 87°18' E –Mangmaya, 27°07' N / 87°15' E, 1700– 300 m, 29. V. 2001, NHMB Basel expedition to Nepal 2 3 1 Ƥ (NHMB); Kosi # 5, Mangmaya, 27°07' N / 87°15' E – Akibunkyabeshi, 27°12' N / 87°15' E, 300-400 m, 30. V. 2001, NHMB Basel expedition to Nepal 3 3 7 Ƥ (NHMB); Kosi # 7, Shawa Khola, 27°17' N / 87°13' E –Khandbari, 27°22' N / 87°12' E, 300–1000 m, 1. VI. 2001, NHMB Basel expedition to Nepal 2 Ƥ (NHMB); Kosi # 8, Khandbari, 27°22' N / 87°12' E –river S of Bhote Bash, 27°27' N / 87°11' E, 1000–1550 m, 2. VI. 2001, NHMB Basel expedition to Nepal 1 3 1 Ƥ (NHMB); Kosi # 10, Mure, 27°30'N / 87°16'E, 2000–2100 m, 6.–8. / 11.–12. VI. 2001, NHMB Basel expedition to Nepal 1 3 (NHMB); Kosi # 12a, Num Khola, 27°33'N / 87°18'E, 900–1000 m, 8.–10. VI. 2001, NHMB Basel expedition to Nepal 1 3 (NHMB); Kosi # 13, Depitar, 27°27'N / 87°17'E –Barabishe, 27°26' N / 87°18' E, 1250– 560 m, 12. VI. 2001, NHMB Basel expedition to Nepal 2 Ƥ (NHMB); Kosi # 14, Barabishe, 27°26' N / 87°18' E –Deorali, 27°24' N / 87°20' E, 560–1450 m, 13. VI. 2001, NHMB Basel expedition to Nepal 3 Ƥ (NHMB); Kosi # 15, Deorali, 27°24' N / 87°20' E –Peak, 27°24' N / 87°22' E, 1450–2800 m, 14. VI. 2001, NHMB Basel expedition to Nepal 1 3 (NHMB); Sankhua Sabha Distr., Arun Valley, betw. Darapangna and Khandbari, 1400– 1100 m, tree-rich cultural land, 21. VI. 1988, J. Martens & W. Schawaller leg., Nepal- Expeditionen Jochen Martens 5 Ƥ (SMNS); E-Nepal, Arun Valley, W. Iswa Khola–W.Siswa Kh., 750–1200 m, 15.–16. VI. 1992, Probst leg. 1 3 2 Ƥ (NHMW); E-Nepal, Arun Valley, Sobae Khola-Tumlingtar- Khandbari, 4–950 m, 4. VI. 1992, J. & J. Probst leg. 1 3 (NHMW); Dhankuta, Arun-Valley, Khandbari to Arunthan, 1100–1300 m, 22. V. 1980, C. Holzschuh leg. 3 3 1 Ƥ (NKME); Bhojpur Distr., Gothe to Majwa, 600–800 m, 27. V. 1997, M. Hauser leg. 2 3 1 Ƥ (SMNS).

BHUTAN: "G. C. Dudgeon, 95–203" 1 Ƥ (NHML).

CHINA: YUNNAN: " China, Yunnan" 3 Ƥ (MTMB); Xishuangbanna, Jinghong city, Ufer d. Mekong, 22°00.42' N / 100°04.14' E, 10. XII.2007, A. Weigel leg. 1 Ƥ (NKME); Xishuangbanna, 29 km NW Jinghong, Da Nuo You (NNNR), 22°12.41' N / 100°38.29' E, 790 m, H KF, 24. V. 2008, A. Weigel leg. 1 Ƥ (NKME); Xishuangbanna, 26 km W Jinghong, vic. Meng Song (NNNR), 22°04.65' N / 100°33.98' E, 1400–1600 m, at flowers of Castanopsis trees, 30. V. 2008, A. Weigel leg. 2 3 (NKME); Xishuangbanna, 26 km W Jinghong, vi. Meng Song (NNNR), 22°04.65' N / 100°33.98' E, 1400–1800 m, at flowers of Castanopsis trees, 30. V. 2008, A. Weigel leg. 1 Ƥ (NKME); Xishuangbanna, 32 km W Jinghong, vic. Man Nuan Hui vill., 21°59.77' N / 100°29.49' E, 1150 m, KF, bloss. trees, 2. VI. 2008, A. Weigel leg. 1 Ƥ (NKME); W-Yunnan, Baoshan, 16. VI. 1993, C. Holzschuh leg. 2 3 (NHMB). GUANGXI ZHUANG: "Wuchong (Kwangsi) 16. V :: 54, H. C. Tao" 1 3 (MNHP); "Andé, Wuchow, Prov. Kwangsi" 1 3 1 Ƥ (NHMW).

TAIWAN: "Pilam ( Formosa), H. Sauter, VIII. 1912 " 1 3 2 Ƥ (DEIM); Kaohsiung Hsien, Paoshan, 4. VI. 1999, W. L. Chen leg. 1 Ƥ (KMNH).

MYANMAR: "Burmah, H. Swale, 1913–117" 1 3 1 Ƥ (NHML); "Burmah, Bowring 63 47*" 1 3 1 Ƥ (NHML); "Carin Cheba, 900–1100 m, L. Fea, X.. 88" 1 Ƥ (MCGD); "Bhamo, Birmania, Fea, VIII. 1885 " 1 3 (MCGD); " Birmania, Shweego-Myo, Fea, Ottobre 1885" 1 3 (MCGD); "Shwegoo-Myo, Fea, Ottobre 1885" 1 3 (MNHB); Upper Burma, Nam Tamai Valley, Alt. 3000 ft., Lat. N. 27° 42', Long. E. 97° 54', 25. VIII. 1938, R. Kaulback leg., B. M. 1938–741. 2 Ƥ (NHML); Upper Burma, Nam Tamai Valley, Alt. 3000 ft., Lat. N. 27° 42', Long. E. 97° 54', 27. VIII. 1938, R. Kaulback leg., B. M. 1938–741. 2 Ƥ (NHML); "Rangoon, H.

E. Andrewes Bequest, B. M. 1922–221" 1 3 2 Ƥ (NHML); "Rangoon, 1902.294" 2 Ƥ (NHML); "Tharrawaddy, H. E. Andrewes Bequest, B. M. 1922–221" 2 Ƥ (NHML); "Tharrawaddy, Burma, 1902.194" 2 Ƥ (NHML); "Tharrawaddy, Burma, 1902.294." 1 Ƥ (NHML); "Maymyo, V. 10, H. L. Andrewes, 1919.26" 1 Ƥ (NHML); "Maymyo, VI. 10, H. L. Andrewes, 1919.26" 1 3 1 Ƥ (NHML); Mon State, Kyaikto, 1.–5. VI. 2003, M. Klícha leg. 1 3 (CMG); Kolaw, S. Shan States, 4000 ft., IV. 1916, F. M. Mackwood leg., B. M. 1916–205. 1 Ƥ (NHML); Shan State, Inle Lake, east Shore, 20°36,48’ N 96°56,53’ E, 12. V. 1999, Schuh & Schillhammer leg. 1 3 (CRS); SW Shan State, Kalaw, 70 km W Taunggyi, 10.–11. VI. 1997, J. Rejsek leg. 7 3 3 Ƥ (SMNS); "Tenasserim, coll. Helfer" 6 3 13 Ƥ (MNPR), 2 Ƥ (NHMB); "Helfer, Tenasserim" 1 3 1 Ƥ (NMPC); "Tenasserim, Tavoy, Fry Coll. 1905.100" 1 3 1 Ƥ (NHML); "Tenasserim, Meetan, Fea, Apr. 1887 " 1 Ƥ (MCGD).

LAOS: " Annam, Laos " 1 Ƥ (MTMB); " Laos, Tonkin " 1 3 (NHMB); " Tonkin, Laos, coll. Staudinger" 3 Ƥ (SMTD); " Siam, Sen Kam, 28.–29. IV. 1920. R. V. de Salvaza" 1 3 (MNHP); " Siam, Vieng Vai, 23.–24. IV. 1920. R. V de Salvaza" 1 3 (MNHP); "Mekong River, Tanoun, 28. V. 1920. R. V. de Salvaza" 1 Ƥ (MNHP); Phongsaly Prov., Ban Sano Mai, 21°21' N / 102°03' E, ~ 1150 m, 19.–26. V. 2004, P. Pacholátko leg. 1 3 2 Ƥ (NHMB); Phongsaly Prov., Boun Neua (4 km E), 21°38' N / 101°57' E, ~ 1100 m, 26. V. 2004, Vit Kubán leg. 1 3 (NHMB); Phongsaly Prov., Ban Nam Li (5 km NW), 21°15' N / 102°14' E, ~ 900 m, 4. V. 2004, M. Brancucci leg. 1 3 (NHMB); Phongsaly Prov., Phongsaly env., 21°41–42' N / 102°06–08' E, ~ 1500 m, 28. V.– 30. VI. 2003, P. Pacholátko leg. 23 3 18 Ƥ (NHMB); Phongsaly Prov., Phongsaly env., 21°41–42' N / 102°06–08' E, ~ 1500 m, 28. V.–20. VI. 2003, M. Brancucci leg. 9 3 14 Ƥ (NHMB); Phongsaly Prov., Phongsaly env., 21°41–42' N / 102°06–08' E, ~ 1500 m, 28. V.–20. VI. 2003, Vit Kubán leg. 19 3 15 Ƥ (NHMB); Phongsaly Prov., Phongsaly env., 21°41' N / 102°06–08' E, ~ 1500 m, 6.–17. V. 2004, M. Brancucci leg. 1 3 (NHMB); Phongsaly Prov., Phongsaly env., 21°41' N / 102°06–08' E, ~ 1500 m, 6.–17. V. 2004, Vit Kubán leg. 2 3 (NHMB); Phongsaly Prov., Phongsaly env., Phu Fa, h: 1450–1600 m, Bergregenwald, 26. VII. 2006, M. Geiser leg. 1 3 1 Ƥ (CMG); Phongsaly Prov., Phongsaly env., Ausserhalb der Ortschaft, Kulturlandschaft, Auf Blüte ( Sambucus ), 27. VII. 2006, M. Geiser leg. 4 3 1 Ƥ (CMG); Phongsaly Prov., Phongsaly env., Ausserhalb der Ortschaft, Kulturlandschaft, Auf Blüte ( Sambucus ), 28. VII. 2006, M. Geiser leg. 1 Ƥ (CMG); Phongsaly Prov., Phongsaly env., Ausserhalb der Ortschaft, Kulturlandschaft, An Strauch, 27. VII. 2006, M. Geiser leg. 1 3 1 Ƥ (CMG); Louang Namtha Prov., Namtha ⊣ Muang Sing, 21°09' N / 101°19' E, 900–1200 m, 5.–31. V. 1997, Vit Kubán leg. 6 3 2 Ƥ (NHMB); Luang Namtha Distr., Lak Kham Mai, road between Luang Namtha–Muang Sing, 5.–9. V. 2007, M. Pejcha leg. 1 Ƥ (CMG); Luang Nam Tha Prov., ca. 20 km SE Muang Sing, 950 m, 12./ 13. VI. 1996, Schillhammer leg. 1 Ƥ (NHMW); Luang Nam Tha Prov., ca. 25 km SE Muang Sing, 900 m, 14./15./ 20.–22. VI. 1996, Schillhammer leg. 1 Ƥ (NHMW); Luang Namtha Distr., Luang Namtha env., 2.–9. V. 2007, M. Pejcha leg. 4 3 5 Ƥ (CMG); Louangphrabang prov., Thong Khan, 19°35' N / 101°58' E, ~ 750 m, 11.–21. V. 2002, Vit Kubán leg. 1 Ƥ (NHMB); Louangphrabang prov., Ban Song Cha, 1200 m, 20°33' N / 102°14' E, 24. IV.–16. V. 1999, Vit Kubán leg. 1 Ƥ (NHMB); Luang Phrabang Prov., Nam Sat stream, 150 m upstr. Hoi Sat Village, 340 m, 48Q 0 220680, UTM 2260258, Malaisetrap, loc 17, 28. IV.–2. V. 2005, N. Jönsson, T. Malm & B. Viklund leg. 1 3 (NRMS); Sayaboury Prov., Sayaboury, 17. V. 1965, Native Collector leg. 1 Ƥ (BIMH); Sayaboury Prov., Sayaboury, 25. III. 1965, Native Collector leg. 1 Ƥ (BIMH); N-Vientiane Prov., Vang-Vieng, 300 m, 18°55'23'' N / 102°26'55'' E, 10.– 15. V. & 1.–6. VII. 2001, Jiři Kolibáč leg. 2 3 2 Ƥ (NHMB); Vientiane prov., Lao Pako env., 55 km NE Vientiane, 200 m, 1.-4. V. 2004, J. Bezdëk leg. 1 3 (NMPC); "Vientiane, 31 / 5 1915, R. Vitalis de Salvaza" 1 Ƥ (MNHP); "Vientiane, 1.–15. IV. 1965, J. L. Gressitt Collector" 2 Ƥ (BIMH); "Umg. Vientiane, III.-VI. 1963 " 2 3 13 Ƥ (ZSMU) 1 Ƥ (CMG); Vientiane prov., Phou Khao Khouay N. P., 18°27' N / 102°49' E, 2005, P. Kresl leg. 5 3 1 Ƥ (NMPC); Vientiane Municipal., Phou Khao Khouay NBCA, Ban Van Hua env., 800– 1000 m, 5.–20. V. 2005, M. Pejcha leg. 8 3 1 Ƥ (CMG); Vientiane Prov., Ban Van Eue, 31. V. 1966 Native Collector leg. 1 Ƥ (BIMH); Houa Phan Prov., Sam Neua (Xam Nua), Stadtrand, Rodungsfläche, Käscherfang, 10. VIII. 2004, M. Geiser leg. 1 3 2 Ƥ (CMG); Houa Phan Prov., Vieng Xai, am Fenster eines Gebäudes, 11. VIII. 2004, M. Geiser leg. 3 3 1 Ƥ (CMG); Hua Phan prov., ~ 20°12' N / 104°01' E, Phu Phan Mt. 1500–1900 m, 17. V.–3. VI. 2007, M. Brancucci leg. 1 3 2 Ƥ (NHMB); Hua Phan prov., 20°12–13.5' N / 103°59.5'– 104°01' E, Ban Saluei ⊣ Phou Pane Mt., 1340–1870 m 15. IV.–15. V. 2008, Lao collectors leg. 1 Ƥ (NHMB); Hua Phan prov., 20°12' N / 104°01' E, Phu Phan Mt. ~ 1750 m, 17. V.–3. VI. 2007, Vit Kubán leg. 1 3 (NHMB); Hua Phan prov., Ban Saluei, Phu Phan mt., 20°13' N / 103°59' E, 1300–2000 m, 6.–18. V. 2004, J. Bezdëk leg. 3 3 1 Ƥ (NMPC); Houa Phan prov., Ban Saluei ⊣ Phou Pane Mt., 1340–1870 m, 20°12–13.5' N / 103°59.5– 104 °01' E, 10. V.–9. VI. 2009, C. Holzschuh & local coll. leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 15 3 5 Ƥ (NHMB), 1 3 1 Ƥ (CMG); Houa Phan prov., Ban Saluei ⊣ Phou Pane Mt., 1340–1500 m, 20°12–13.5' N / 103°59.5– 104 °01' E, 2.–9. VI. 2009, M. Brancucci leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 31 3 17 Ƥ (NHMB); Houa Phan prov., Ban Saluei ⊣ Phou Pane Mt., 1340–1500 m, 20°12–13.5' N / 103°59.5– 104 °01' E, 12.–16. VI. 2009, M. Brancucci leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 1 3 7 Ƥ (NHMB); Houa Phan prov., Ban Saluei ⊣ Phou Pane Mt., 1300–1900 m, 20°11–13' N / 103°59– 104 °01' E, flying around in village (daytime), 10.–17. VI. 2009, Michael Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 31 3 26 Ƥ (NHMB), 1 3 1 Ƥ (CMG); Houa Phan prov., Ban Saluei ⊣ Phou Pane Mt., 1300–1900 m, 20°11–13' N / 103°59– 104 °01' E, village, visiting flowers of Sambucus sp., 10.–17. VI. 2009, Michael Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 7 3 3 Ƥ (NHMB); Xieng Khouang Prov., 19°27' N / 103°13' E, Phonsavan env., 1150 m, 5.–6. VI. 2007, Vit Kubán leg. 1 Ƥ (NHMB); Xieng Khouang prov., Phonsavan town ⊣ Phou Phadeng Mt., 1100–1200 m, 19°26' N / 103°13' E, flying around in town (daytime), 9. V. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 8 3 6 Ƥ (NHMB), 1 3 1 Ƥ (CMG); Xieng Khouang prov., Phonsavan town ⊣ Phou Phadeng Mt., 1100–1200 m, 19°26' N / 103°13' E, flying around in town (daytime), 31. V. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 4 3 10 Ƥ (NHMB), 1 3 1 Ƥ (CMG); Xieng Khouang prov., Phonsavan town ⊣ Phou Phadeng Mt., 1100–1200 m, 19°26' N / 103°13' E, flying around in town (daytime), 1. VI. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 2 3 1 Ƥ (NHMB); Xieng Khouang prov., Phonsavan town ⊣ Phou Phadeng Mt., 1100–1200 m, 19°26' N / 103°13' E, flying around in town (daytime), 18.–24. VI. 2009, M. Geiser leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 7 3 2 Ƥ (NHMB); Xieng Khouang Prov., Phonsavan, Stadtgebiet, An Strauch, 7. VIII. 2006, M. Geiser leg. 1 3 (CMG); Xieng Khouang Prov., Phonsavan, Saylomyen-Golfplatz, An Strauch, 7. VIII. 2006, M. Geiser leg. 2 3 (CMG); Xieng Khoang Prov., by road no. 68, ca. 80 km NE from Phonsavan, 15. V. 2007, M. Brancucci leg. 1 3 (NHMB); Xieng Khuang prov., Nong Haet, 19°30' N / 104°03' E, 29.–31. V. 2001, J. Bezdëk leg. 2 3 1 Ƥ (NMPC); Xieng Khouang Prov., Phu Xane, 1200–2200 m, 3. IV. 2001, Local collector leg. 1 3 (KMNH); Xieng Khouang prov., Muang Khoun, 1070 m, 19°19.91' N / 103°21.99' E, flying around in town (daytime), 22. VI. 2009, M. Geiser & D. Hauck leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 4 3 (NHMB); Xieng Khouang Prov., Muang Khoun env., 10. V. 2009, Chantaseng Keomaravong leg. 1 3 2 Ƥ (CMG); Xieng Khouang Prov., road Ban Thaviang ⊣ Muang Khoun, ca. 20 km from Ban Thaviang, 700 m, 19°11.80' N / 103°21.20' E, 22. VI. 2009, D. Hauck leg., NHMB Basel, NMPC Prague, Laos 2009 Expedition, M. Brancucci, M. Geiser, Z. Kraus, D. Hauck, V. Kubáň. 1 3 (NHMB); Borikhane Prov., Pakkading, 17. V. 1965, Native Collector leg. 3 Ƥ (BIMH); Borikhane Prov., Pakkading, 14. VII. 1965, Native Collector leg. 4 Ƥ (BIMH); Borikhane Prov., Pakkading, 3. VIII. 1965, Native Collector leg. 1 Ƥ (BIMH); Borikhane Prov., Pakkading, 9. IX. 1965, Native Collector leg. 1 Ƥ (BIMH); Borikhane Prov., Pakkading, IX. 1965, Native Collector leg. 1 Ƥ (BIMH); Boli Kham Xai Prov., Ban Nape (8 km E), ~ 600 m, 18°21' N / 105°08' E, 1.–18. V. 2001, Pacholátko leg. 1 3 2 Ƥ (NHMB); Laksao 2. VI. 1998, S. Kawabe leg., Exot. Insect Coll., No. 16848. 1 Ƥ (KMNH); Boli Kham Xai prox., 70 km NEE of Vientiane, 18°16' N / 103°11' E, 150 m, 27.–30. IV. 1997, Vit Kubán leg. 2 Ƥ (NHMB); Boli Kham Xai prox., 70 km NEE of Vientiane, 18°16' N / 103°11' E, 150 m, 2.– 30. VI. 1997, Vit Kubán leg. 1 3 1 Ƥ (NHMB); Khammouane Prov., Phon Tiou, 28. IV. 1965, Native Collector leg. 3 Ƥ (BIMH); Khammouane Prov., Phon Tiou, 28. V. 1965, Native Collector leg. 3 Ƥ (BIMH); Khammouane Prov., Phon Tiou, 28. VI. 1965, Native Collector leg. 1 Ƥ (BIMH); Khammouane Prov., Phon Tiou, 6. VII. 1965, Native Collector leg. 4 Ƥ (BIMH); Khammouane Prov., Phon Tiou, 4. VIII. 1965, Native Collector leg. 1 Ƥ (BIMH) 2 Ƥ (CMG); Savannakhet Prov., Savannakhet, 20. VII. 1965, Native Collector leg. 1 Ƥ (BIMH); Sekong Prov., Bolavens Pl., N-slope, ca. 10 km N Mg. Tha Theng, 500–700 m, 29./ 30. V. 1996, Schillhammer leg. 8 3 2 Ƥ (NHMW); Champassak Prov., Bolavens Plateau, M. Paksong–B. Nam Thang, 800–1100 m, 26./ 27. V. 1996, Schillhammer leg. 1 Ƥ (NHMW); Attopeu Prov., Houei Kong, 16. V. 1965, Native Collector leg. 1 Ƥ (BIMH); Attopeu Prov., Houei Kong, 31. V. 1965, Native Collector leg. 1 Ƥ (BIMH).

VIETNAM: " Tonkin Centr., N.-E. de Tuyen-Quan, Qium-Quam-Thuong, A. Weiss 1901, avril-juin" 1 3 2 Ƥ (MNHP); " Tonkin Centr., Région de Tuyen-Quan et de Phu-An-Binh, A. Weiss 1901, avril-juin" 1 Ƥ (MNHP); " Tonkin Central, Région de Tuyen-Quan et de Dong-Chau, A. Weiss 1901, avril-juin" 1 3 (MNHP); " Tonkin N., Rég. d'Ha-Giang (H te riviére claire), Siébens-Olivier 1916" 1 Ƥ (MNHP); "M es du Ht Song Chai, Rabier 258–95" 3 Ƥ (MNHP); "Song-Chay, 1908, Indo Chine, coll. Dussault" 1 Ƥ (NHMB); " Tonkin, Hoabinh, Aug. 1918, R. V. de Salvaza" 2 Ƥ (MNHP); Hoa Binh, coll. W. Wittmer" 3 3 9 Ƥ (NHMB); " Tonkin, Env. de Hoa-Binh, J. Laisi 1901" 1 Ƥ (MNHP); 52 km SW of Lang Son, 21°35' N 106°30' E, 370 m, 27. IV.–6. V. 1996, Pacholátko & Dembický leg. 4 3 (NHMB); Shonla Pr., Shonla, 10. V. 1991, Murzin leg. 2 Ƥ (NHMB); " Tonkin, Laokay, 10 / 8 1913, R. Vitalis de Salvaza" 1 Ƥ (MNHP); " Annam " 1 Ƥ (NHMB); " Annam, Mus. Pragense" 1 Ƥ (NHMB).

THAILAND: " Siam, Bowring 63 47*" 2 Ƥ (NHML); "S. Siam, H. Way, 99–135." 1 3 (NHML); " Siam, Gyldenst." 2 Ƥ (NRMS); " Siam, [locality written in Thai], 18. Jun. 1936, Lot 316, To: Czecho, List: 068–05, Lot: 316/19" 1 Ƥ (NHMB); " Siam, [locality written in Thai], 1000 ft., 28. Jun. 1936, Lot 321, To: Czecho, List: 068–05, Lot: 321/19" 1 Ƥ (NHMB); " Siam, [locality written in Thai], 2. Jun. 1938, Lot 529, To: Czecho, List: 068–05, Lot: 529/19" 1 Ƥ (NHMB); " Siam, [locality written in Thai], 8. Jan. 1938, Lot 534, To: Czecho, List: 068–05, Lot: 534/19" 2 Ƥ (NHMB); "Doi Vieng Par, Siam, Gyldenst." 1 Ƥ (NRMS); Mae Hong Son Prov., Ban Huai Po, 1600 m, 9.–16. V. 1991, Pacholátko leg. 1 Ƥ (NHMB); Mae Hong Son Prov., Ban Huai Po, 19° 19' N 97° 59' E, 1600–2000 m, 17.–23. V. 1991, L. Dembický leg. 3 Ƥ (NHMW); Mae Hong Son Prov., Ban Huai Po, 1600–2000 m, 8.–18. V. 1992, J. Horak leg. 1 3 (NHMB); Mae Hong Son Prov., Ban Huai Po, 1600 m, 8.–17. V. 1992, S. Bily leg. 1 Ƥ (NHMB); Mae Hong Son prov., Soppong env., 600 m, 28. V.–2. VI. 1999, M. Riha leg. 1 3 1 Ƥ (NHMB); NW-Thailand, Pai, 500 m, 24.–25. IV. 2004, W. Schawaller leg. 1 3 (SMNS); Chiang Mai prov., San Pakia vill., 19°19' N / 98°50' E, 1.–15. V. 1988, V. Kubán leg. 1 3 (NHMB); Chiang Mai Prov., Ban San Pakia, 1700 m, 25. IV.–7. V. 1996, Sv. Bilý leg. 1 3 (NHMB); Chiang Mai prov., San Pakia vill., 19°19' N / 98°50' E, 1400 m, 1.–15. V. 1998, V. Kubán leg. 1 3 (NHMB); Chiang Dao, Ban San Pakia, 1200 m, 5.–10. V. 2004, Sv. Bilý leg. 3 Ƥ (NMPC); Chiang Mai, 56 km NW, 99°25' / 19°05', 7.–14. VI. 1995, M. Snizek leg. 1 3 1 Ƥ (NHMB); Chiang Mai Pref., Doi Suthep, 1.–5. VI. 1986, H. Hirasawa leg., Exot. Insect Coll., Nos. 9870–9876. 5 3 2 Ƥ (KMNH); Chiang Mai pref., Chiang Mai city, 19. VI. 1988, O. Furuta leg., Exot. Insect Coll., No. 16849. 1 3 (KMNH); "Thanon Thong Chai", Umphang riv., 1000 m, 16°07' N / 99°00', 28. IV.–6. V. 1991, D. Král & V. Kubán leg. 1 3 (NHMB); Loei prov., Phu Kradung N. P., 1300 m, 16°53' N / 101°47' E, 11–15. V. 1999, M. Riha leg. 1 3 (NHMB); Saraburi, III. 1989, Sab. Steinke leg. 1 Ƥ (SMNS); "Renong, Siam (Doherty), Brit. Mus., 1923–320" 1 Ƥ (NHML).

MALAYSIA: "Fed. Malay States, 1909, C. J. Brooks, B. M. 1931–570. 1 3 3 Ƥ (NHML); " Malacca " 1 Ƥ (MNHP); " Malacca, Coll. Franklin Müller" 4 Ƥ (DEIM); " Malacca, Tengah Gebirge, P. Zobris. V." 2 Ƥ (MNHB). KEDAH: "Talan Sintok, N. Kedah, IV.–VII. 1938, A. J. Slatter, ex. mass of tropical vegetation" 1 3 (NHML); Langkawi Island, 23. III.–14. IV. 1992, G. & M. Novak leg. 1 Ƥ (NHMB). PENANG: "P. Pinang, Batu Ferringghi", 28. VII. 1975, Kling leg. 1 3 (NHML); "Penang, Lamb. 93.60." 1 Ƥ (NHML); "Penang Is., Oct. 1913, G. E. Bryant, G. Bryant Coll. 1919–147" 1 3 1 Ƥ (NHML); "Penang, G. E. Bryant, XI. 1913, G. Bryant Coll. 1919–147" 3 Ƥ (NHML); "Penang, G. E. Bryant, 3. XI. 13 " 1 Ƥ (NHML); "Penang, G. E. Bryant, 4. XI. 13 " 1 3 (NHML); "Penang, 14. XI. 13, G. Bryant Coll. 1919–147" 1 Ƥ (NHML); "Penang, Bowring 63 47*" 1 3 5 Ƥ (NHML); "Penang, 96–85." 1 Ƥ (NHML); "Penang, H. N. Ridley, 97–1900–111." 1 Ƥ (NHML).

PERAK: " Malakka, Perak" 1 Ƥ (MNHP); "Perak, Doherty, Fry Coll., 1905.100" 1 Ƥ (NHML); Cameron Highland, 30 km SE of Ipoh, Ringlet, 900 m, 18.–22. I. 1999, P. Čechovský leg. 2 Ƥ (CMG); Ipoh (30 km SE), 900 m, Cameron Highland, Ringlet, 18.–22. I. 1999, P. Cechovský leg. 1 3 (NHML); Banjaram Titi Wangsa, Fraser's Hill, 9. III. 1997, Ivo Jeniš leg. 1 Ƥ (NMPC); "Tapah, Perak, G. B. Cerruti 1900" 2 Ƥ (MNHP). PAHANG: Cameron Highlands, Tanah Rata, 20.–24. II. 1998, A. Kudrna jr. leg. 1 3 (CMG); Cameron Highlands, Tanah Rata, 13.–17. II.1997, Ivo Jeniš leg. 2 Ƥ (NMPC); Cameron Highlands, Tanah Rata, 1600 m, 11.–27. II. 2000, P. Pacholátko leg. 1 3 (NHMB); Cameron Highlands, Tanah Rata, 1500–1800 m, 2.–26. III. 2004, P. Pacholátko leg. 1 3 (NHMB); Cameron Highlands, Tanah Rata, 1500–1800 m, 22.–26. I. 1999, P. Pacholátko leg. 1 3 (NHMB); Cameron Highlands, Tanah Rata, 1600 m, 11.–27. II. 2000, P. Pacholátko leg. 1 3 (NHMB); Cameron Highlands, Tanah Rata, 1500–1700 m, 1.–13. II. 2003, P. Pacholátko leg. 2 Ƥ (NHMB); Cameron Highlands, Tanah Rata, 1500–1800 m, 2.–26. III. 2004, P. Pacholátko leg. 10 3 5 Ƥ (NHMB); Cameron Highlands, Tanah Rata, V. 1996, Wong Tet Fatt leg. 1 Ƥ (MTMB); Cameron Highlands, 1600 m, IV.-VI. 1993, G. J. Minet leg. 1 3 1 Ƥ (NHMB); M. Fraser's Hill, 1250 m, 27. I. 1948, F. E. Leiler leg. 1 3 (NRMS). KELANTAN: Banjaran Titi Wangsa Mts., Ladang Padrak, 1500–1800 m, 9.–11. IV. 1997, P. Čechovský leg. 2 3 (NHMB). TERENGGANU: Tasik Kenyir, 26. VII.–2. VIII. 1993, H. Forster leg. 2 3 1 Ƥ (NHMW). KUALA LUMPUR: "K. Lumpur, 12. 1. 10, Fed. Malay States, London Sch. Trop. Med., Pres. by Imp. Bur. Ent., Brit. Mus. 1927–503" 1 Ƥ (NHML). SARAWAK: " 1907–1909, C. J. Brooks, B. M. 1936–681. 2 Ƥ (NHML); "G. J. Brooks, B. M. 1928–193" 1 3 (NHML); "Borneo, Sarawak, 1865–66, coll. G. Doria" 7 Ƥ (MCGD), 4 Ƥ (MNHB), 3 Ƥ (MNHP); "Quop, W. Sarawak, II.-III. 1914, G. E. Bryant, G. Bryant Coll. 1919– 147" 1 3 1 Ƥ (NHML); "Quop, W. Sarawak, III. 1914, G. E. Bryant, G. Bryant Coll. 1919–147" 1 Ƥ (NHML)¸ "Quop, W. Sarawak, III.-IV. 1914, G. E. Bryant, G. Bryant Coll. 1919–147" 1 Ƥ (NHML); "Quop, 30. III. 14, G. Bryant Coll. 1919–147" 1 Ƥ (NHML); "Quop, W. Sarawak, G. E. Bryant, IV. 14, G. Bryant Coll. 1919– 147" 2 Ƥ (NHML); "Quop, W. Sarawak, G. E. Bryant, 8. IV. 14, G. Bryant Coll. 1919–147" 2 Ƥ (NHML); " Sarawak, Bidi. 1908–9, C. J. Brooks, B. M. 1929–551" 1 Ƥ (NHML); "Puak, Sarawak, G. E. Bryant, 3. V. 14, G. Bryant Coll. 1919–147" 1 3 (NHML); "Kuching, J.E.A. Lewis, 1910–116" 1 Ƥ (NHML); "Bornéo, Kuching, July 10, 1900 " 1 3 (MHHP); "Matang Rd., Ap 0 9, J.E.A. Lewis, 1910–116" 1 Ƥ (NHML); "Matang Rd, 3 1/ 2 miles, 8. 5. '09" 1 Ƥ (NHML); "Borneo, Xántus, Sarawak, Matang" 1 Ƥ (MTMB). LABUAN: "Borneo, Labuan, Fry Coll., 1905.100" 1 Ƥ (NHML). SABAH: "Borneo, Kinabalu, Fry Coll. 1905.100" 1 Ƥ (NHML); "Borneo, Kinabalu, Whitehead, Fry Coll. 1905.100" 3 Ƥ (NHML); "B. N. Borneo, Mt. Kinabalu, Kiau, 3000 ft. 3: 5: 1929, Ex F. M. S. Museum. B. M. 1955–354." 1 Ƥ (NHML); "British N. Borneo, Ranau, 500 m, Nov. 5–8, 1958, T. C. Maa Collector" 1 Ƥ (BIMH). BANGGI (Sabah): "Banguey b. Borneo, coll. Staudinger" 5 3 (SMTD).

SINGAPORE: " Singapore " 1 Ƥ (MNHB); " Singapore, H. N. Ridley, 97–109." 1 Ƥ (NHML).

INDONESIA: "Sunda-Inseln, v. Studt" 2 Ƥ (MNHB). SUMATRA: "Sumatra" 1 Ƥ (NHML); "Sumatra, coll. W. Wittmer" 1 Ƥ (NHMB); „Sumatra 622.9“ 1 3 (MHNG); "Sumatra, Coll. Kraatz" 1 Ƥ (DEIM); "Sumatra, Dehy" 1 Ƥ (MNHP); "Sumatra, Deli, Siboelangit, Jachan V." 1 Ƥ (MNHB); "Sumatra, Dolok Baroe" 1 Ƥ (MTMB); "Indragiri, Sumatra, v. Meckel" 1 Ƥ (NHMB); "Balighe, X. 90 – III. 91, E. Modigliani, coll. W. Wittmer" 1 Ƥ (NHMB); "Sumatra, Balighé (Modigliani)" 1 Ƥ (MNHP), "Balighe, X. 90 – III. 91, E. Modigliani" 2 Ƥ (MCGD); "Sumatra, Balighé, X. 90 – III. 91, E. Modigliani, Coll. Kraatz" 1 Ƥ (DEIM); "Pajakumbo" 1 3 1 Ƥ (MNHP); "Si Pirok" 2 3 (MNHB); "Palembang" 3 Ƥ (MNHP); "Harau valley, VII. 1991 " 1 Ƥ (NHMB); "Harau Valley 1991, nat. coll." 1 Ƥ (NHMB); "Fort de Kock, VIII. 94, E. Modigliani" 1 Ƥ (MCGD); "Fort de Kock (Sumatra), 920M, Maart 1921, leg. E. Jacobson" 1 3 (MNHP); "Fort de Kock (Sumatra) 920M, October 1921, leg. E. Jacobson" 1 3 (MNHP); "Fort de Kock (Sumatra) 920M, December 1921, leg. E. Jacobson" 1 Ƥ (MNHP); "Fort de Kock (Sumatra) 920 M, 1926, leg. E. Jacobson" 1 Ƥ (MNHP); "Fort de Kock, 920 m, Jan. 1921, E. Jacobson leg., B. M. 1927-48." 1 Ƥ (NHML); "Fort de Kock (Sumatra), 920M, Feb. 1921, leg. E. Jacobson" 1 Ƥ (DEIM); "Fort de Kock, 920 m / 192 ft., E. Jacobson leg., B. M. 1928–247" 1 3 1 Ƥ (NHML); "Fort de Kock (Sumatra), 920M, Januari 1922, leg. E. Jacobson" 1 Ƥ (MNHB); W-Sumatra, Bukkittinggi, am Licht, 7. II. 1991, Schödl leg. 1 Ƥ (NHMW); Barat, 10 km S of Bukittinggi, 1600 m, Gn. Merapi, 10.–11. I. 2005, Bolm leg. 1 Ƥ (CMB); "Bedagei int., Sumatra's O.K., +/–600', 2. Sem.

89, I. Z. Kannegieter" 1 3 (MNHP); "Sumatra, Lampung, Fry Coll. 1905.100" 1 3 1 Ƥ (NHML); "Sumatra, Lampung, Buxton, Fry Coll. 1905.100" 1 3 (NHML); "Dr. B. Hagen, Tandjong, Morawa, Serdang (N. O. Sumatra), coll. W. Wittmer" 1 3 (NHMB); "Sungei Bulu, Sett. 1878, O.Beccari" 1 Ƥ (MCGD); "Dol. Tarabugna, Modigliani, 1890" 1 Ƥ (MCGD); "Pea Ragia, X. 1890, E. Modigliani" 1 Ƥ (MCGD); "Sumatra, Pea Ragia, X. 1890, E. Modigliani, Coll. Kraatz" 1 3 (DEIM); "F. C. Drescher, Zuid-Sumatra, 500 m, degiesting G. Tanggarmoes, IV. 1934, coll. W. Wittmer" 1 Ƥ (NHMB); "F. C. Drescher, Zuid-Sumatra, 500 m, degiesting G. Tanggarmoes, IX. 1934, coll. W. Wittmer" 1 Ƥ (NHMB); Lampung Prov., Bukit Barisan Selatan N. P., 5 km SW Liwa, 5°4' S / 104°4' E, 600 m, 21. III. 2003, L. Dembický leg. 1 Ƥ (NHMB); Brastagi, Fenster, 21. VIII. 1924, coll. W. Wittmer 1 3 (NHMB); Brastagi-Sibayak, 22.–25. V. 1991, Jiři Moravec leg. 1 3 (NHMB); Brastagi, 76 km S from Medan, 30. III.–1. IV. 1996, S. Becvar leg. 1 3 (NHMB); "Corporaal, Brastagi, 10. 5. 1918, Sumatra (Corporaal)" 1 Ƥ (MNHP); "Brastagi, Sumatra: Mjöberg" 1 3 (MNHP); Utara, Brastagi, Gn Sibayak, 1600–2200, 26. I.–1. II. 2005, Bolm leg. 1 3 (CMB); "Medan, Sumatra, coll. Hayek" 2 3 1 Ƥ (ZSMU); "Medan S O.K, L. Fulmek, Tabakopslagplaats" 1 Ƥ (NHMW); W. Sumatra, Talang, II. 1991, native collector" 1 Ƥ (NHMB); Aceh, Kutacane, Ketambe, 400 m, 7.–19. X. 1991, Barries leg. 1 Ƥ (NHMB); W. Sumatra, Anai V. Nat. Res., 10 km W Padangpanjang, 250 m, 17. XI. 1989, Löbl, Agosti & Burckhardt leg. 1 Ƥ (MHNG); W-Sumatra, Padang env., 2000– 500 m, IV. 1995, S. Jakl leg. 1 3 (NHMB); "Padang, 1890, E. Modligliani" 1 Ƥ (MCGD); Liman Manis b. Padang, Schoede S. G., 8. I. 08 " 1 Ƥ (MNHB); N-Sumatra, Bukit Lawang, 27. II. 1990, Schillhammer leg. 1 Ƥ (NHMW); N-Sumatra, D. Toba, Fluß s Prapat, 17. II. 1990, Schillhammer leg. 1 3 (NHMW); Kerinci prov., Seblat N. P.: 24 km NE Tapan, Muara Sako ⊣ E env., 2°05' S / 101° 15' E, 400–550 m, 4.–18. III. 2003, Dembický leg. 1 Ƥ (NHMB); Kerinci, Pelompek, 13.–15. V. 1991, Jiři Moravec leg. 1 Ƥ (NHMB); Sungei Penok, Korinchi Valley, 2600 ft., III. 1914. 1 Ƥ (NHML); Mt. Kerintji, Aro Estate. Au Kayo. 1.–5. III. 1954, A. H. G. Alston leg., B. M. 1954-414. 2 3 1 Ƥ (NHML); Ophir mts., Gunung Talamau, 17 km E Simpangempal, 750 m, 21.–25. V. 2001, Bolm leg. 1 Ƥ (SMNS); "E Coast Sumatra, Doerian Moelan, Brindjei, Lt. R. Coughtrie, 1915–184" 1 Ƥ (NHML); "N. O. Sumatra, Tebingtinggi, Dr. Schultheiss, Coll. Schultheiss" 4 3 4 Ƥ (DEIM). NIAS: "Sumatra, Nias, Fry Coll. 1905.100" 1 Ƥ (NHML); "Nias Island, Sharp Coll. 1905–313." 1 Ƥ (NHML). "Sumatra, Nias, German Mission, Fry Coll., 1905.100" 1 Ƥ (NHML); "Nias, Coll. Kraatz" 1 3 (DEIM); "Is. Nias, 1897–98, U. Raap" 2 Ƥ (MCGD). KALIMANTAN: "S. O. Borneo, Grabowsky S. V." 1 Ƥ (MNHB); "S. O. Borneo, Grabowsky S. V., Mindai, 6. 82" 1 Ƥ (MNHB); "Pengaron, S. Borneo, Brit. Mus. 1923–320." 2 Ƥ (NHML); "Pengaron, S. E. Borneo, 1904–150." 1 Ƥ (NHML); W-Kalimantan, Nanga Ela env., 700 m, Nanga Nyeruh, 4.–10. VII. 1993, Schneider leg. 1 Ƥ (NHMW); "Ponteanak, Borneo, VII. 07 " 4 3 (BIMH); "Borneo, Mahakam" 1 3 1 Ƥ (MNHB); "Bandjermasin, L. W. Schaufuss G." 3 Ƥ (MNHB); "Long Navang, Borneo, Mjöberg" 11 Ƥ (NRMS), 1 Ƥ (CMG). JAVA: "Java" 1 Ƥ (DEIM), 1 3 (MTMB), 2 Ƥ (NHMB), 1 Ƥ (NHML); "Java 1921" 1 Ƥ (MNHB); "Java, 40 4 1 2850" 1 Ƥ (NHML); "Java, coll. W. Wittmer" 1 Ƥ (NHMB); " India or., coll. E. Friv., Java" 1 Ƥ (MTMB); "Horsfield, 60–15" 1 3 1 Ƥ (NHML); "Java, Bowring, 63 47*" 3 Ƥ (NHML); "Java, coll. Staudinger" 3 Ƥ (SMTD); "Java, Coll. Kraatz" 4 3 11 Ƥ (DEIM); "Java, Aurivillius" 1 Ƥ (NRMS); "Fruhstorfer, Java, Coll. Kraatz" 3 Ƥ (DEIM); "Java, Oberthur, 100–96" 1 3 (MNHP); "Java, J. D. Pasteur 268–94" 3 3 4 Ƥ (MNHP); "Ullr." 1 Ƥ (NHMW); "Warsberg, 868" 1 Ƥ (NHMW); "Pfeiffer" 1 3 (NHMW); Pfeiffer, 852" 1 3 (NHMW); "Pfeiffer, 853" 1 Ƥ (NHMW); "Pfeiffer, 854" 1 Ƥ (NHMW); "Pfeiffer, 854" 1 Ƥ (NHMW); "Java, Doleschall" 1 Ƥ (MTMB); "Schmidt-Göbel, 1884" 1 3 1 Ƥ (NHMW); "Leyden, d. H., Java" 1 Ƥ (ZSMU); "Ploem, Java, 1870" 1 3 1 Ƥ (NHMW); "Java, XII. 1879, 36." 1 3 2 Ƥ (NHMW); "Java merid., 1500, H. Fruhstorfer 1891" 1 Ƥ (MCGD); "Java, Wijnkoops b, Kemner" 1 Ƥ (NRMS); Pagilaran Estate, 1940, K. Landberg leg. 1 Ƥ (NRMS); "Buitenzorg, F. Muir" 1 Ƥ (BIMH); "Buitenzorg, +/–1000' Aug. 95, I. Z. Kannegieter" 1 Ƥ (MNHP); "Buitenzorg, Axel Preyer S. G." 1 Ƥ (MNHB); "Buitenzorg, G. B. Ferrari 1875" 5 3 6 Ƥ (MCGD); "G. Papandajan, Garoet Preanger, Jan. 91. 4– 6000' I. Z. Kannegieter" 1 Ƥ (MNHP); "Preanger, Java" 1 3 1 Ƥ (SMTD); "Java merid., Palabuan, 1892, H. Fruhstorfer" 1 Ƥ (MNHP); "Sipayon, R. Oberthür 97–97" 1 Ƥ (MNHP); "Java (Meuwen Bay), détr de la Sonde, Raffray & Maindron, 1878" 1 Ƥ (MNHP); " Batavia, P. Serre 1904" 1 3 2 Ƥ (MNHP); "V. Java, Bandoeng, 700 m, 2/ XII. 1943, Kurt Landberg" 1 Ƥ (NRMS); "Idjen Plateau, V.-VI. 1920, H. Lucht, Blair d.t." 2 Ƥ (DEIM); "Megamenbong, Java, Xántus" 1 3 1 Ƥ (MTMB); "Moegamendong, leg. Xantus" 1 3 3 Ƥ (MTMB); "Sinagar, Java, Xántus" 2 3 (MTMB); "Simagar, leg. Xantus" 1 3 1 Ƥ (MTMB); "Palaboen, Java, Xántus" 3 Ƥ (MTMB); "Sindanglaja, Java, Xántus" 9 3 5 Ƥ (MTMB); "Sindanglaja, leg. Xantus" 1 Ƥ (MTMB); "Java, Xantus, Simdanglaja" 2 Ƥ (MTMB); "Java occident., Mons Gede, 8000`, Aug. 1892, H. Fruhstorfer" 1 Ƥ (MNHB); "Batoe, 1923" 1 Ƥ (MNHB); "Malang, XI. 21 " 1 Ƥ (MNHB); "Sinagar, Giava, Beccari, IV. 1876 " 3 3 2 Ƥ (MCGD); "Giava, Tcibodas, Ott. 1879, O. Beccari" 2 Ƥ (MCGD); "Java occident., Sukabumi 2000, H. Fruhstorfer 1893" 1 Ƥ (MCGD); "Depok, VIII. 31 " 1 Ƥ (NHMB); "Tankoeban, Prahoe, Lembong, 1100 m, 25. 7. 36, coll. W. Wittmer" 3 Ƥ (NHMB); "Silosanen, E. Java, coll. W. Wittmer" 1 Ƥ (NHMB); "Batoerraden, G. Slamet, Java, F. C. Drescher, II. 1936, 800 m, coll. W. Wittmer" 2 Ƥ (NHMB); "C. O. Nandjoeng-Djaja, G. Simpal, 600 Mr., Preanger, Java, II. 1934, leg. C. P. J. de Haas, coll. W. Wittmer" 2 Ƥ (NHMB); "Garahan, Bes Besoeki, E. Java" 1 Ƥ (NHMB); "Boeloe Lawang, Res. Pasoeroean (Holz)" 1 3 (NHMB); "Soekaboemi, Java, 3. 8. 32 " 1 Ƥ (NHMB); "Bogor" 1 3 1 Ƥ (MNHP). SUMBAWA: " Insel Sumbawa, coll. Staudinger" 1 Ƥ (SMTD); "B. Aroe Hassa, Sambawa, 2–5000', Doherty IX. X., Brit. Mus. 1923–320 1 3 1 Ƥ (NHML); Tambora, 600 m, 9. XI. 1941, Rolf Blomberg leg. 3 Ƥ (NRMS). FLORES: C- Flores, Moni, Wolowaru, 11. XI. 1949, Dr. Bühler & Dr. Sutter leg. 1 Ƥ (NHMB). SUMBA: " Insel Sumba" 1 Ƥ (NHMB); W. Sumba, Kodi, 30. VII. 1949, Dr. Bühler & Dr. Sutter leg. 1 3 (NHMB); W. Sumba, Pogobina, 17. IX. 1949, Dr. Bühler & Dr. Sutter leg. 1 Ƥ (NHMB); C. Sumba, Lokojengo, 23. IX. 1949, Dr. Bühler & Dr. Sutter leg. 1 Ƥ (NHMB); O. Sumba, Baing, 1. VII. 1949, Dr. Bühler & Dr. Sutter leg. 1 Ƥ (NHMB). ALOR: "Maroe, W. Allor, 2000', Doherty, X., Brit. Mus. 1923–320." 1 Ƥ (NHML). TIMOR (west): "Koepang, Timor, 90–80." 1 Ƥ (NHML); "Siva, West-Timor, Sunda Exp. Rensch, 1. VII. 1927 " 1 Ƥ (MNHB). DAMAR: "Dammer Insel " 2 Ƥ (MNHB); " Insel Dammer, coll. Staudinger" 1 Ƥ (SMTD). SULAWESI: "Celebes" 1 Ƥ (NHML); „Celebes, P. de Borre“ 1 Ƥ (MHNG); "Célébes, De la Savinière 1877" 1 Ƥ (MNHP); "S. Celebes, 250 m, 3. 1941, coll. W. Wittmer" 1 Ƥ (NHMB); "Celebes, Kandari, III. 74, O. Beccari" 1 Ƥ (MCGD); "Tondano, Celebes, coll. Staudinger" 10 3 4 Ƥ (SMTD); "Tondano, Celebes" 4 3 (NHMB); "Drs. Sarasin, N. Celebes, Tomohon, M. 49." 3 3 5 Ƥ (NHMB); Minahassa, Tomohon, 31. V. 1954, A. H. G. Alston leg., B. M. 1954-414. 1 3 (NHML); Minahassa, Tomohon, 10. VII. 1954, A. H. G. Alston leg., B. M. 1954- 414. 1 Ƥ (NHML); Sulawesi Utara, Gunung Ambang F. R., nr. Kotamobagu, flood debris by stream, 11. I. 1985, R. Ent. Soc. London, Project Wallace, B. M. 1985–10. 1 Ƥ (NHML); Sulawesi Utara, Dumoga-Bone N. P., Base camp area, ca. 190 m, at light, X. 1985, R. Ent. Soc. London, Project Wallace, B. M. 1985–10. 1 Ƥ (NHML); Sulawesi Utara, Dumoga-Bone N. P., Fog 11, 230 m, BMNH Plot A, 10. III. 1985, R. Ent. Soc. London, Project Wallace, B. M. 1985–10. 1 3 (NHML); S-Sulawesi, 8 km W Mamasa, 950 m, 2°56'13'' S / 119°20'32'' E, 18.–21. VII. 1999, Bolm leg. 1 Ƥ (SMNS); N.E. of Makassar, Rantepao– 340 km, V. 1966, R. Straatman leg., 3 Ƥ (BIMH), 1 3 1 Ƥ (CMG); "Süd-Celebes, Bonthain, C. Ribbe 1883" 1 Ƥ (MNHB); "Bonthain, Z. W. Celebes, J. P. A. Kalis, 9. 1938, coll. W. Wittmer" 1 3 1 Ƥ (NHMB); "Bonthain, Z. W. Celebes, J. P. A. Kalis, 10. 1938, coll. W. Wittmer" 1 3 (NHMB). TUKANGBESI: "Toekan Besi-Ins., Kalidupa" 1 Ƥ (MNHB). TANIMBAR: "Jandema, Tenimber, Doherty, VI. VII., Brit. Mus. 1923–320" 1 Ƥ (NHML). BURU: "Ins. Buru" 1 Ƥ (MNHB). AMBON: "Waai, P. Ambon, 5. VII. 1959, leg. A. M. R. Wegner" 1 Ƥ (NHMB). CERAM: Umg. Manusela, 700–900 m, 16.–18. II. 1989, Schödl leg. 6 3 (NHMW). HALMAHERA: "Dodinga, Halmaheira, Doherty, VIII., Brit. Mus. 1923–320" 1 Ƥ (NHML). KAI: "Key- Ins." 4 Ƥ (MNHB); "Ins. Key" 1 Ƥ (MNHB); "Isole Key, 1873, O. Beccari" 1 Ƥ (MCGD); "Key Inseln, coll. Plason, coll. W. Wittmer" 1 3 (NHMB), "Ins. Key (Planten)" 1 3 (MNHP); "Ins. Key, Bandai" 1 Ƥ (MNHB); " Ile de Tonal, Groupe de Key" 1 Ƥ (MNHP). IRIAN JAYA: "Nuova Guinea, Andai, Ag. 72, L. M. D'Albertis" 2 Ƥ (MCGD); "N. Guinea, Hatam, Beccari 1875" 2 Ƥ (MCGD); "N. Guinea, Hatam, Beccari, VI. 1875 " 1 Ƥ (MCGD); "N. Guinea, Hatam, Beccari, VII. 1875 " 1 Ƥ (MCGD); Testega, 1100–1300 m, 30. III.–2. IV. 1993, A. Riedel leg. 1 3 1 Ƥ (SMNS); Iba, 1300 m, 7.–8. IV. 1993, A. Riedel leg. 1 Ƥ (SMNS); Westpapua, Manokwai Prov., vic. Mokwam, Siyoubrig, 1400–1800 m NN, 01°06.26' S / 133°54.41' E, 24.–28. II. 2007, A. Skale leg. 1 Ƥ (NKME).

TIMOR-LESTE: Baaguia, VIII. 1935, C. Bühler & Meyer 1 Ƥ (NHMB); "Dili, Port. Timor, 2500', Doherty IV. V. Brit. Mus. 1923–320" 1 Ƥ (NHML).

PHILIPPINES: "Phil Isla " 1 Ƥ (NHML); "Phil. Isl., 350" 1 3 (NHML); " Philippinen, Juli 1982, Fr. Schramm" 1 Ƥ (SMNS); "Phillip. Islands, Semper, Fry Coll., 1905.100" 1 3 2 Ƥ (NHML); "Acc. No. 15453, Bu. of Sci. P. I., collected by C. R. Jones, coll. W. Schultze" 2 3 (SMTD). LUZON: "Luzon, P. I., coll. W. Schultze" 2 Ƥ (SMTD). PANAY: "N. W. Panay, Baker, Philipp. Is., C. F. Baker, Brit. Mus. 1922–57 " 2 Ƥ (NHML). MINDANAO: "Dansalan, 9. II. 1915 " 1 3 (MNHB); "Momungan, coll. Staudinger" 1 3 1 Ƥ (SMTD). PALAWAN: "N. Palawan, Binaluan, Nov.-Dez. 1913, leg. G. Boettcher" 1 Ƥ (MNHB); Cleopatra Needle Nat. P., Tanabank Riv. Val., 300 m, 20.–22. XII. 1990, Bolm leg. 2 3 (NHMB).

UNCLEAR OR IMPRECISE LOCALITIES: " India or." 1 3 (MTMB); "E. Ind." 1 3 (NHML); "12. 1, 225, Blair d.t." 1 Ƥ (DEIM); "7710–85" 1 3 (MNHP); "Khouang, 1923, Indo Chine, coll. Dussault" 1 3 1 Ƥ (NHMB); Budeuyory? [illegible] 1 3 1 Ƥ (MNHP); "Borneo, Pascoe Coll., 93–60." 1 Ƥ (NHML); "Borneo, ... [illegible], German Mission, Fry Coll., 1905.100" 1 Ƥ (NHML); "SAR, Bowring 63 47*" 2 Ƥ (NHML); "Borneo, SAR" 2 Ƥ (NHML); "Borneo, Wallace, Fry Coll., 1905.100" 2 Ƥ (NHML); "Borneo" 1 Ƥ (NHML); "Borneo, coll. Staudinger" 1 3 (SMTD); "Nord-Borneo, coll. Staudinger" 2 Ƥ (SMTD); "Borneo, Chaper 1891" 8 Ƥ (MNHP); "246–78" 1 3 3 Ƥ (MNHP); "Hattam, Vraz" 1 Ƥ (NMPC).

NO LOCALITY: 2 3 (NHMW); „coll. Melly“ 1 Ƥ (MHNG); "Hist. Coll. ( Coleoptera ), Nr. ????" 4 3 (MNHB).

Measurements 3 (n = 595): TBL 8.3–12.2 mm, L-h 7.1–10.4 mm

HL 1.2–2.1 mm, PL 1.5–2.1 mm, EL 5.6–8.3 mm

Measurements Ƥ (n = 850): TBL 9.8–13.3 mm, L-h 8.2–11.4 mm

HL 1.5–2.3 mm, PL 1.5–2.1 mm, EL 6.7–9.4 mm

Differential diagnosis: This species is very similar to P. coeruleipennis , but easily recognized by the constant differences in coloration: P. b i c o l o r always has orange yellow or brownish elytra, while in P. coeruleipennis , the elytra are always metallic blue or green. Other important differences are the shape of the median lobe in the male aedeagus, as well as the phallobase, which is more strongly bent downwards in P. bicolor , but never as strong as in P. wittmeri . The other species of Prionocerus clearly differ from P. b i c o l o r by coloration, shape of aedeagus and the last abdominal sternite in males.

Redescription: Habitus as in figs. 33 (3) and 34 (Ƥ). Body metallic blue, sometimes slightly greenish. Pronotum usually bright reddish orange, elytra usually orange yellow (see under "Variability"). Antennae almost black, with at least the largest part of the terminal segment brownish orange or dark brown, this coloration can extend to large parts of segments 9–10. The maxillary palpi are usually almost completely black, but can be partly reddish to a very variable extent. Tarsi black or bluish black, claws orange brown, sometimes infuscate.

Head without any obvious differences to P. coeruleipennis . Antennae of very similar shape to P. coeruleipennis (including sexual dimorphism), but the last segment is a bit longer, about 1.5 times as long as the first or slightly longer.

Pronotum similar in shape to P. coeruleipennis , maximal length: maximal width 1: 0.9–1: 1.1; with two shallow, rather indistinct oblique impressions in the basal half, at both sides of the disc, finely bordered, but bordering interrupted along the front margin; shining, without any microsculpture; regularly, but sparsely covered with some inconspicuous suberect black or subrecumbent yellow hairs and with a few longer black setae along the outer margin.

Elytral shape very similar to P. coeruleipennis , usually a little bit more elongate, and slightly more parallel in males; humeral area convex, slightly flattened towards the apex; finely and regularly punctate over their whole surface, but punctures getting shallower towards the apex; regularly covered with fine, short recumbent hairs of the same colour as the elytra, and bearing some few, blackish, long, erect hairs, partly arranged into rows; granular rows hardly visible or completely absent in most specimens. Margins crenulate, more strongly in humeral area, getting slightly weaker towards the apex, the crenules all bearing blackish setae. Scutellum as in P. coeruleipennis .

Femora metallic, shining, with fine punctures and sparse, greyish pubescence. Tibiae also metallic, densely covered with brownish setae. Tarsi not metallic, with blackish or brownish setae and some greyish hairs.

Abdomen with rather short, not very dense, greyish, recumbent pubescence and some longer black, suberect setae; with rather dense, fine punctures and somewhat rugose texture in some parts.

Male: Last abdominal sternite of similar shape as in P. coeruleipennis , slightly larger and broader; basal margin of the same shape (fig. 23). Apical margin not quite as deeply incised, the opening of more transverse shape, the last tergite almost not emarginate.

Aedeagus (figs. 2, 9, 16): Process of phallobase slightly more tilted downwards than in P. coeruleipennis . Paramera rather flat, but with basal part thicker than in P. coeruleipennis ; arcuate; their tip often (but not always) with a very small dorsal tooth, always with long brownish hairs along the ventral edge, as well as a row of short, minute, black teeth before the apex. Seen in dorsal view, they are straight and almost parallel as in P. coeruleipennis , only a bit more narrowed towards the apex, making the gap between them slightly broader. Median lobe straight after the basal part, very weakly narrowed until after the endophallus insertion, then dorsally with a small tooth and ventrally abruptly narrowed, tip produced into a very flat disc. In dorsal view with oval, elongate ostium; its tip subtriangular and similar to P. coeruleipennis .

Sexual dimorphism: The elytra are more parallel and generally slightly narrower in males, broader and with more rounded sides in females. Female are usually slightly larger in total body size. The eyes are larger in males and the frons is narrower between the eyes. The antennal joints 5–10 are more dilated in males than they are in

Variability: As already noted, the coloration of the last antennal segments and the maxillary palpi is subject to strong individual variation.

The colour of the scutellum is metallic blue in most individuals, but very rarely orange yellow as the elytra (1 3 from Mae Hong Son, Ban Huai Po at NHMB and 1 Ƥ from Myanmar, Tharrawaddy at NHML). This variation has already been noticed by Gorham (1895) and Champion (1919) but has not been named.

A dark form was described by Pic under the name of " var. notaticollis ". In this form, the elytra are infuscate reddish brown in variable gradations, sometimes almost dark brown. The pronotum is often infuscate at the disc, sometimes forming a diffuse black discal macula. This colour form seems to occur only in Peninsular Malaysia and Sumatra, where it coexists with the typical form. As the shape of the aedeagus and all other examined characters are absolutely identical to the typical form, it cannot be regarded as a subspecies or even as a different species. P. b i c o l o r var. notaticollis Pic, 1910 is here treated as a synonym of P. bicolor , as it was already suggested by Champion.

Another "aberrative" specimen was found at NHML, collected in Myanmar ("Tenasserim, Tavoy"). It has two large, oblong, comma-shaped black patches on the pronotum, one at each of the normally coloured middle line. The patches are not reaching the pronotal margins. This specimen is a male, but it is damaged and had lost its abdomen, so its aedeagus could not be examined.

Distribution (fig. 44): This species is very widely distributed in a large part of the Oriental region, including the border regions to the Palearctic (Himalaya, South China, Taiwan) and New Guinea (see discussion).

Nomenclatural note: According to the original description, there is little doubt that P. pertii Castelnau, 1836 is identical to P. b i c o l o r Redtenbacher, 1868, even though the type of P. p e r t i i was not examined. This, however, makes P. b i c o l o r a junior synonym of P. pertii , a species which was never cited by anyone and omitted from all catalogues after its description in 1836. Every time since 1868, P. b i c o l o r was used as valid name for this common and widespread species. Article 23.9.1.2. of the ICZN (1999) does not allow the automatic consideration of P. p e r t i i a nomen oblitum, so an application will be sent to the International Commission of Zoological Nomenclature, in order to maintain the usage of P. b i c o l o r as a valid species.

Prionocerus paiensis n. sp.

Holotype 3: "NW Thailand, Mae Hong Son, Ban Huai Po, 1600 m, 8.–17. V, 1992, S. Bily leg."; NHMB.

Paratypes (1 3, 6 Ƥ): 1 Ƥ: same data as the holotype ( NHMB); 1 3: " Thailand –nord, Pai, 22.–29. 5. 1996, A. Kudrna jr. Lgt." (CMG); 1 Ƥ: "NW Thailand, 19.19° N, 97.59° E, Mae Hong Son, Ban Si Lang, 1200 m, 23.–31. 5. 1991, L. Dembický leg." ( NHMB); 1 Ƥ: "NW Thai, 19.19° N, 97.59° E, Mae Hong Son, Huai Sua Tao, 11.–17. 5. 1992, L. Dembický leg." ( NHMB); 3 Ƥ: " Thailand bor. occ., Pai, 6. 6.–9. 6. 1997, Lgt. M. Snizek" ( NHMB).

Type locality: Ban Huai Po, Mae Hong Son, Thailand.

Measurements 3 (n = 2): TBL 11.8–12.5 mm, L-h 9.7–10.3 mm

HL 2.1–2.2 mm, PL 1.8–1.9 mm, EL 7.9–8.4 mm

Measurements Ƥ (n = 6): TBL 12.2–13.6 mm, L-h 10.1–11.5 mm

HL 2.0– 2.4 mm, PL 1.8–2–0 mm, EL 8.3–9.6 mm

Differential diagnosis: Easily distinguished from all other species of the genus by the coloration of the femora and antennae and the shape of the paramera.

Description: Habitus as in figs. 45 (3) and 46 (Ƥ). Body metallic dark blue. Pronotum, pro- and mesothorax, trochanters, basal part of all femora, antennae, maxillary and labial palpi and central part of scutellum yellowish orange. The yellowish coloration extends to about 2/3 of the fore femora, half of the middle femora, but only the extreme base of the hind femora. The trochanters are usually darkened along their outer (posterior) edge. Blue metallic lustre of the tibiae and outer part of the femora somewhat indistinct, often almost black, tarsi somewhat brownish. Elytra shining metallic dark blue.

Head behind the eyes about half as wide as the middle part of the pronotum. Vertex shining, very sparsely and very finely punctate and with a few long, erect black hairs. Frons between eyes shining, sparsely punctate, at its narrowest part about half as broad as the length of the first antennal joint in males, slightly narrower than the length of the whole joint in females; in front of the eyes distinctly depressed and slightly rugose. Clypeus trapezoidal, almost half as long as wide at the base and very slightly convex, with a few fine punctures and some longer hairs. Labrum about as long as wide, rather flat and more coarsely punctate than the head, with a few long blackish hairs.

Male antennae reaching the first quarter of the elytra in length, until slightly after the humeral callus. First three segments subfiliform (basal one very slightly incrassate), segments 4 slightly widened, 5 widened and flattened, 6–10 of subtriangular shape, very strongly widened and with a sharp outer angle. Last segment robust, not conspicuously widened, but clearly emarginate. First segment long, second very short, less than one third as long, third again long, almost like the first, the following a bit shorter, slightly decreasing in length, but increasing in width, the last segment shorter than 1 and 3. Female antennae slightly shorter, only scarcely reaching the humeral callus; median and subapical segments a bit less strongly widened and their outer angle not as sharp.

Pronotum slightly wider than long, widest in apical half, before middle, maximal length: maximal width 1: 1.05–1: 1.2; shining, with few sparse punctures in the basal half. Hind angles somewhat oblique and slightly rounded, but still forming an angle of approximately 100 degrees; front angles rounded; basal and lateral margins distinctly bordered, bordering broad at hind angles, becoming finer, more indistinct and then completely disappearing towards the fore margin, basal half distinctly, obliquely impressed at both sides between disc and hind angles, disc and part before the hind angles slightly convex. Pubescence of pronotum yellow, sparse, decumbent, denser and clearly visible along the base and near the apical angles, sparser near the anterior margin and almost completely lacking on the disc. Lateral margins with some long, black, hairlike setae, almost lacking (fallen off?) in some individuals.

Elytra of about the same shape as in P. coeruleipennis but slightly more elongate (2.5–2.6 times as long as wide in males, 2.3–2.4 times in females), widest in apical third; humeral area convex, apical half less flattened than in other species; finely, densely and regularly punctate over their whole surface, the punctures getting slightly shallower and denser towards the apex; regularly covered with fine, short recumbent, pale pubescence, sparsely intermixed with long, erect hairs which are partly arranged into rows; with three more or less distinct, parallel rows of granules starting in the basal third and continuing until shortly before the apex. Margins finely crenulate, the crenules all bearing blackish setae. Scutellum like in P. coeruleipennis .

Femora and tibiae not metallic (or only slightly), covered with black setae, intermixed with recumbent greyish hairs. Tarsi also not metallic, with blackish setae.

Abdomen with fine, silky, recumbent, greyish, rather dense pubescence and some longer black or brownish, suberect setae; with small, fissure-like punctures and somewhat rugose texture.

Male: Last abdominal sternite of similar shape as in P. coeruleipennis and P. b i c o l o r, somewhat intermediate in width; basal margin of similar shape (fig. 24). Apical margin not quite as deeply incised as in P. coeruleipennis , the emargination more similar to P. b i c o l o r, but less rounded, the opening of subtriangular, slightly transverse shape, the last tergite almost not emarginate.

Aedeagus (figs. 3, 10, 17): Process of phallobase pointing straight backwards. Paramera less flattened than in the two preceding species and strongly sclerotised, almost straight, not arcuate, with slightly thickened apex. Seen in dorsal view, they are very broad, straight, and only very slightly diverging towards the apex. At the tip with a small dorsal tooth and some minute teeth or granulae, as well as some long, brownish hairs, along the ventral edge. Median lobe like in P. coeruleipennis , but a bit less curved base and subapical more arcuate.

Sexual dimorphism: Females have broader, less parallel elytra and are usually slightly larger than males. The eyes are larger and more protruding in males than in females and therefore the frons between the eyes is also slightly narrower in males. The antennae are much more filigrane and their joints 5–10 are much less dilated in females than they are in males.

Variability: The limited material seems quite uniformous in coloration and other characters.

Distribution (fig. 45): So far known only from some localities in north-west Thailand (province Mae Hong Son). Its occurrence in the adjacent parts of Myanmar is possible.

Derivatio nominis: Named after the village Pai in Mae Hong Son province, North-West Thailand, where one half of the type series has been collected.