Popilius tetraphyllus (Eschscholtz, 1929)

Jiménez-Ferbans, Larry, Reyes-Castillo, Pedro & Schuster, Jack C., 2015, Passalidae (Coleoptera: Scarabaeoidea) of the Greater and Lesser Antilles, Zootaxa 3956 (4), pp. 491-512 : 506-507

Popilius tetraphyllus (Eschscholtz, 1929)

( Fig. 20 )

Type locality. Guyana.

Diagnosis. 17.0–21.0 mm total length. Anterior border of labrum straight. Clypeus exposed, visible dorsally, with anterior angles well developed. Frontal-clypeal suture present. Frontal fossae punctate and pubescent. Internal tubercles large. Central tubercle with apex not free; latero-posterior tubercles marked, joined by a ridge to central tubercle. Antennal club with three or four lamellae. Apex of lacinia bidentate. Mentum with medial base protruding and glabrous. Prosternal process rhomboidal, acute. Mesosternum glabrous, with weak scars. Antero-lateral metasternum pubescent, lateral groove glabrous, disc delimited by punctures basally. Humeri and epipleura glabrous.

Distribution. Bolivia, Brazil, Colombia, French Guiana, Guyana, Trinidad and Tobago, and Venezuela ( Gillogly 2005).

Comments. We did not review material from the Antilles of this species; the diagnosis and measurements are based on Gillogly (2005) and material from Brazil and Guyana deposited in the IEXA collection.

Passalidae from Antilles Reference

Passalus (Passalus) abortivus ** Kaup (1871): Antilles

Passalus (Passalus) bucki ** Trinidad

Passalus (Passalus) coarctatus Trinidad

Passalus (Passalus) interruptus ** Arrow (1907) Trinidad

Schuster (1978): Trinidad

Passalus (Passalus) interstitialis Kaup (1871) : Cuba

Arrow (1907): Trinidad, Grenada, Cuba, Jamaica Passalus (Pertinax) affinis * Percheron (1835): Dominican Republic Arrow (1907): Santo Domingo, Haiti Reyes-Castillo et al. (1995): Dominican Republic, Haiti Passalus (Pertinax) antillarum * Arrow (1907): Grenada

Hincks & Dibb (1935)

Passalus (Pertinax) dominicanus * van Doesburg (1953): Dominican Republic Reyes-Castillo et al. (1995) Dominican Republic Passalus (Pertinax) latifrons ** Schuster (1992): Tobago

Passalus (Pertinax) pertyi * Kaup (1869): Cuba

García et al. (2009): Holguín, Cuba. Passalus (Pertinax) doesburgi * DominicanRepublic

Paxillus jamaicensis * Hincks (1950): Jamaica

Paxillus pentaphyllus * Palisot de Beauvois (1805): Dominican Republic Reyes-Castillo et al. (1995): Dominican Republic, Haiti

Spasalus crenatus Arrow (1907) : Dominica, Saint Lucia, Nevis (as Passalus puncticollis

Lepeletier & Serville)

Gravely (1918): Guadeloupe

Chalumeau (1978) Saint Lucia (as P. puncticollis ) Cartwright & Chalumeau (1978): Dominica (as P. puncticollis ) Galindo-Cardona et al. (2007) Puerto Rico Popilius tetraphyllus ** Gillogly (2005): Trinidad

Verres furcilabris ** Arrow (1907): Trinidad

Marshall (2000): Tobago, Trinidad Veturius sinuosus ** Hunt (1982): Trinidad (as Ve tur ius transversus )

Figures

FIGURES 19 – 22. Head and pronotum, dorsal view. 19. Spasalus crenatus. 20. Popilius tetraphyllus. 21. Ver res furcilabris. 22. Veturius sinuosus. Scale bars: 1 mm.