Cochlostoma crassilabrum (Dupuy, 1849)

Gofas, Serge, 2001, The systematics of Pyrenean and Cantabrian Cochlostoma (Gastropoda, Cyclophoroidea) revisited, Journal of Natural History 35 (9), pp. 1277-1369 : 1341-1344

Cochlostoma crassilabrum (Dupuy, 1849)

ArieÁge: MNHN collection ÐAlzen ( CH 76), 2 spm. (as Pomatias neglectus ) and 1 spm. (as P. daralli ), coll. Locard; Aulus ( CH 64), 6 spm. (as P. fagoti ), 2 spm. (as P. neglectus ) and 2 spm. (as P. subobscurus ), coll. Locard; Cerizols ( CH 47), 1 spm., coll. Germain, ®gured in Germain (1931) pl. 18 ®gure 541; Foix ( CH 86), 4 spm., coll. Denis; Le Mas d’Azil ( CH 67), 21 spm., leg. Chevalier 21/8/1968; Vicdessos ( CH 73), 9 spm., coll. Mortillet, 1872.

New materialÐ Etang de Lers ( CH 6842, 1100 m), limestone rocks, 16 (7 juv)1 3* spm., 7/8/1988, MNHN .

Haute-Garonne: MNHN collection ÐCazarils ( CH 04), 41 6 spm. (as P. crassilabris ) and 41 4 spm. (as P. subobscurum ), coll. Locard; Cierp ( CH 15), 11 31 6 spm. (as P. crassilabris ), coll. Locard; Cierp, 5 spm., coll. Ad. Dollfus; Lespugues ( CH 18), 6 spm., coll. Germain; Luchon ( CH 04), 4 spm. (as P. crassilabris ), coll. Locard; Montagne d’Arri ( CH 15), 3 spm. coll. H.Fischer; Montagne d’Arri, 5 spm., purchased from Marie, 1883; St. BeÂat ( CH 15), 3 spm., leg. de Boissy, 1835; St. BeÂat, 6 spm. (as P. crassilabris ), coll. Locard; St. BeÂat, 2 spm., coll. Letellier; St. BeÂat, 4 spm., coll. Denis; Valentine ( CH 17), 1 spm., leg. Chevallier 20/8/1968.

New materialÐ Astau, woodland (BH9537, 1250 m), 10 spm., 23/10/1987, UPV .

Hautes-PyreÂneÂes: MNHN collection ÐArrens (YN36), 4 spm. (as Pomatias subobscurum ), coll. Locard; Artigues (BH75), 1200 m, 3 sh.; BagneÁres de Bigorre (BH77), 71 3 spm., coll. Denis; BagneÁres de Bigorre, 7 spm., coll. de Morgan; BagneÁres de Bigorre, 4 spm., purchased from Vimont, 1878; BagneÁres de Bigorre, 4 spm., coll. A.Dollfus; BagneÁres de Bigorre, 61 spm., coll. Denis; BagneÁres de Bigorre, 3 spm., coll. Staadt; BagneÁres de Bigorre, 61 51 51 2 spm., coll. Locard; BagneÁres de Bigorre, 4 spm. (as Pomatias spelaeus ), coll. Lavezzari; BagneÁres, grotte du BeÂdat (BH77), 2 spm. (as P. frossardi ), coll. Jousseaume; BagneÁres, grotte du BeÂdat, 2 spm. (as P. frossardi ), coll. Denis; BagneÁres, grotte du BeÂdat, 3 spm. (as P. spelaeus ), coll. A.Dollfus; BagneÁres, grotte du BeÂdat, 3 spm. (as P. spelaeus ), purchased from Marie, 1883; BagneÁres, grotte du BeÂdat, 2 spm. (as P. spelaeus ), coll. Locard; BareÁges (BH65), 14 spm., coll. H. Fischer; BareÁges, 2 spm. (as P. crassilabris ), and 4 spm., coll. Locard (as P. subobscurum ); BareÁges, 19 spm., coll. Michaud, 1841; Bigorre, 9 spm., coll. Letellier; Bigorre, 11 spm., coll. Jousseaume; Bigorre, 6 spm., coll. Locard; Cauterets (YN35), 29 spm. (as P. obscurus , det. Dautzenberg) [1 6 C. partioti ], coll. Lavezzari; Cauterets, 15 spm., leg. Bayan; Cauterets, 6 spm., coll. H.Fischer, leg. Dautzenberg 1901; Cauterets, 5 spm., coll Bertrand; Cauterets, 17 spm., coll. Arnoul, leg. 1880; Cauterets, 31 31 61 11 21 51 51 6 spm (as P. crassilabris ), 4 spm. (as P. nouleti ), 2 spm., (as P. berilloni ) and 3 spm. (as P. subobscurus ), coll. Locard; Col d’Aubisque, road towards Soulor (YN26, 1360 m), 14 sh., leg. Chevallier 9/8/1968; FerrieÁres: val de l’ Ouzom (YN26), 4 spm., leg. Chevallier; Gerde (BH77), 19 spm. (as Pomatias gerdensis ), coll. A.Dollfus, 1903; Gerde, 6 spm. (as P. gerdensis ), coll. Locard; Gripp (BH75), 13 spm., leg. E.Batten, 1953; Gripp (BH75), leg. Tillier; Lourdes (YN47), 3 spm. ex Dupuy; Lourdes, 1 spm. (label by Dupuy); Lourdes, 4 spm., coll. Soyer; Lourdes, 41 3 spm. (as P. crassilabris ), coll. Locard; 2 spm., (as Pomatias mabillianus ), coll. Locard ex Saint-Simon; Luz (YN45), 7 spm., coll. Denis; Paiolle (BH86, 1100 m), 4 spm. (as Cochlostoma crassilabris ), coll. Germain; Peyrouse, (YN47), 5 sh., leg. Chevallier; St. Sauveur (YN45), 3 spm. (as Pomatias crassilabris ), coll. Locard; Tramezaigues (BH84), 5 spm., coll. Germain.

New materialÐ Arrens, rocks ( YN 2658, 950 m), 6 spm. 28/10/1987, UPV ; BeaudeÂan (BH6967), 301 68* (5 juv) spm., 10/1989, MNHN ; Cadeac-Govaux (BH 8351, 700 m), 24/10/1987, UPV ; Cauterets ( YN 3655, 800 m), 25/10/1987, UPV ; Gavarnie, limestone mounds ( YN4536 , 1300 m), 4 juv., 4/1988 and 11/1988, MNHN ; GeÁdre (BH5541, 1100 m), 4 spm., 10/1989, MNHN ; La Vialette (BH 6766, 775 m), 3 spm., 24/10/1987, UPV ; Lourdes ( YN47 ), 291 6 spm., 15/6/1985, coll. J. Pino, Barcelona ; Ost, birch woodland ( YN 3667, 450 m), 5 spm., 25/10/1987, UPV ; Pic du Jer, limestone in grass ( YN 4174, 900 m), 31 58* spm., 29/3/1988, MNHN ; St. Pe de Bigorre, limestone ( YN 3175, 400 m), 31* spm., 6/1987, MNHN ; Soulam (BH7945, 1300 m), 6 spm., 24/10/1987, UPV .

PyreÂneÂes-Atlantiques: MNHN collectionÐ Assat, 2 spm., coll. Denis; Aste (YN 17, 400 m), 2 coq.; Bayonne (XP31), 3 spm., coll. de Morgan; Bayonne, 5 spm. (as Pomatias nouleti ), coll. Locard; Biarritz, Chambre d’Amour (XP 11, 110 m): 2 coq., leg. Chevallier, 7/4/1966; CoÃte de Behobie (XP00), 8 spm. (as P. hidalgoi ), purchase from Marie, 1883; Callaber, 2 spm., coll. A.Dollfus; Cambo (XP30), 2 spm., coll. A.Dollfus; Cambo, 4 spm. (as P. berilloni ), coll. Locard; Castet (YN 17, 400 m), 2 spm. leg. Chevallier; Col d’Osquich (XN 68, 500 m), 1 spm., leg. Chevallier; Eaux- Bonnes (YN16), 3 spm. (as P. fagoti ) and 4 spm. (as P. subobscurus ), coll. Locard; Eaux-Chaudes (YN16), 4 spm., coll. Staadt; Hendaye (XP00), 3 spm. (as P. berilloni ), coll. Locard; Laruns (YN 16, 530 m), 21 sh., leg. Chevallier 2/8/1968; Mousserolles, 3 spm. (as P. angustus ), coll. Locard; Salies de BeÂarn (XP72), 5 spm. (as P. crassilabris ), coll. Locard; St. Jean de Luz (XP10), 2 spm., coll. A. Dollfus; St. Pierre d’Irube (XP31), 5 spm. (as P. nouleti ), coll. Locard; Urrugne (XP10), 4 spm. (as P. berilloni ), coll. Locard.

New materialÐ Arthez d’Asson ( YN 2374, 400 m), 28/10/1987, UPV ; Arudy, limestone cliOE ( YN0875 , 450 m), 71 33* spm., leg. Gofas 5/1981 and 29* spm. (7 juv), 11/1989, MNHN ; Arudy, ( YN0976 , 500 m), 29/10/1987, UPV ; Atherey, ( XN 7372, 250 m), 25 spm., 28/2/1982, and 7 spm., 30/10/1987, UPV ; Aydius ( YN0063 , 700 m), 3 spm., 29/10/1987, UPV ; Barcus ( XN8184 ), 1 spm., 28/2/1982, UPV ; Bedous ( XN 9562, 400 m), 2 spm., 29/10/1987, UPV ; Beherobie, rocks in grass ( XN 4870, 600 m), 67* spm., leg. Gofas 11/1980, MNHN ; Castet ( YN 1072, 500 m), 11 spm., 29/10/1987, UPV ; Cette-Eygun ( XN9755 ), 10/198/1985, UPV ; Col d’Escale ( XN9743 , 1100 m), 7 spm., 29/10/1987, UPV ; Col d’Haltza, rocks in grass ( XN 5472, 750 m), 104 spm., 11/1980 and 39* spm., leg. Gofas 6/1987, MNHN ; Eaux-Bonnes, limestone ( YN 1361, 850 m), 89 (24 juv.)1 133*1 117*1 54#spm., leg. Bouchet and Gofas 6/1989, MNHN ; Eaux-Bonnes ( YN 1261, 850 m), 28/10/1987, UPV ; Eaux- Chaudes ( YN0958 ), 5 spm. leg. Gofas 5/1989, MNHN ; Eaux-Chaudes ( YN0959 , 650 m), 28/10/1987, UPV ; Escot ( XN9572 ), 2 spm., UPV ; Gave de Brousset ( YN1346 1300 m), 1 spm., 28/10/1987, UPV ; Gorges de Kakoueta ( XN 7561, 500 m), 4*1 30 spm., 1 30 juv., leg. Gofas 7/1990, MNHN ; Gorges de Kakoueta ( XN7561 ), 7 spm., 17/2/1980, and 8 spm., 30/7/1982, UPV ; Gourette ( YN1760 , 1200 m), 49 (15 juv), 1 76* (12 juv.)1 35#spm., leg. Bouchet and Gofas 5/1989, MNHN ; Gourette ( YN1860 , 1400 m), 60 (3 juv), 1 90* spm., leg. Bouchet and Gofas 5/1989, MNHN ; Gourette, beech woods ( YN1759 , 1400 m), 28/10/1987, UPV ; Grotte de Sare , small limestone blocks ( XN 1692, 200 m), 95* spm., leg. Gofas 11/1980, MNHN ; Holzarte ( XN6964 ), 3 spm., 27/7/1980, UPV ; Iraty, limestome mounds on grass ( XN5767 , 1000 m), 76*1 19 spm., (12 juv), leg. Gofas 6/1987, MNHN ; Iriberri ( XN 4978, 150 m), 34* spm., leg. Gofas 11/1980, MNHN ; Issor ( XN 9074, 300 m), 1 spm., 29/10/1987, UPV ; Kolorotz, limestone in grass ( XP4600 ), 46 spm., leg. Gofas 11/1980, MNHN ; Larrau, beech woods on karst ( XN6460 , 1500 m), 30/10/1987, UPV ; Lescun ( XN 9256, 900 m), 6 spm., 9/4/1980, UPV ; Massif des Arbailles , limestone in grass ( XN 6274, 900 m), 12* spm., leg. Gofas 7/1987, MNHN ; Mauleon ( XN 7187, 200 m), 3 spm., 27/02/1982, UPV ; Montagne du Rey, limestone ( YN1273 ), 4 spm., leg. Gofas 5/1981, MNHN ; Muskildi ( XN6584 ), 2 spm., 21/2/1982, UPV ; Pic de Gillers , beech woods on karst ( XN8463 1250 m), 7 spm., 30/10/1987, UPV ; Pierre St Martin, karsti®ed limestone ( XN 8461, 660 m), 211 (25 juv.)1 20*1 35#spm., 7/1990, MNHN ; PragneÁres, woodland (BH 5545, 920 m), 6 spm., 25/10/1987, UPV ; Prat-Lahite (BH 8271, 600 m), 24/10/1987, UPV ; St. Christau ( XN9576 ), 9 spm., leg. Bouchet 6/1989, MNHN ; Sainte-EngraÃce, beech woods ( XN 8262, 500 m), 16 spm., 28/2/1982, and 8 spm., 28/7/1982, UPV ; Sainte-EngraÃce: village ( XN 7762, 500 m), 3 spm., 28/7/1982, UPV ; St Martin d’Arberoue ( XP6605 , 80 m), 5 spm., 14/10/1987, UPV .

Huesca: New materialÐ Bielsa (BH7125, 1200 m), 49 (20 juv) spm. 1 eggs 1 9* spm., leg. Bouchet and Gofas 10/1989, MNHN; Candanchu ( YN 0039 , 1360 m), 3 spm., 3/8/1981 , UPV; Canfranc ( YN0231 ), numerous spm., leg. Serrano 8/1985, coll. L. Dantart, Barcelona; Formigal , ( YN1638 , 1500 m), 9 spm., 8/1985 , UPV; Formigal ( XN1640 , 1580 m), 361 10*1 15#spm., leg. Bouchet and Gofas 10/1989, MNHN; Formigal , beech woods ( YN1638 , 1500 m), 8 spm., 30/11/1988 , UPV; Plan de EscuÂn (BH 7716, 1300 m), 14 spm., 25/9/1987 , UPV; Tramacastilla de Tena ( XN2132 ), limestone mounds in shade 21 12* spm., leg. Gofas 8/1987, MNHN; Zuriza ( XN7845 ), 2 spm., leg. Serrano 8/1985, coll. L. Dantart, Barcelona.

Navarra: New material: Alto Lizarraga ( WN8047 , 1000 m), 2 spm., 24/5/1986, UPV ; Alsasua: Sorozarreta ( WN6849 ), 1 spm., 2/3/1980, UPV ; Aralar: Altxueta ( WN 8456 ), 10 spm., 2/11/1980, UPV ; SastruÂstegui: BeriaõÂn ( WN8349 ), 2 spm., 5/8/1981, UPV ; Belagua ( XN 7756 ), 4 spm. 26/6/1982, UPV ; Belagua : mountain refuge, 8 spm., 14/8/1980, UPV ; BiguÈezal : near water spring ( XN5127 ), 1 spm., 8/7/1981, UPV ; Cilveti ( XN 2560, 700 m), 8 spm., 12/8/1981, UPV ; Ezcaroz ( XN 5550 ), 6 spm., 10/7/1981, UPV ; Foz de ArbayuÂn ( XN 4928 ), 8 spm., 8/7/1981, UPV ; Foz de Lumbier ( XN3721 ), 48 (24 juv)1 16* (2 juv)1 11# spm., leg. Gofas and Salas 8/1990, MNHN ; Gamoeta (1700 m), 6 spm., leg. Serrano 8/1985, coll. L. Dantart, Barcelona; Huarte Arakil ( WN85 ), 2 spm., 12/4/1982, UPV ; Iraty ( XN 5561 ), 191 25 spm., 11/10/1981, UPV ; Irurzun: rio Ega ( WN95 ), 2 spm., 20/8/1979, UPV ; Isaba , 5 spm., 15/8/1980, UPV ; Lumbier ( XN 3923 ), 1 spm., 14/2/1982, UPV ; Monreal: Elomendi ( XN 2229 ), 1 spm., 6/1/1982, UPV ; Nagore ( XN3343 ), 4 spm., 11/4/1982, UPV ; Ochagavia ( XN5651 ), 17 spm., leg. Serrano 8/1985, coll. L. Dantart, Barcelona; Orbaiceta ( XN 4459 ), 1 spm., 10/8/1980, UPV ; Orotz-Betelu ( WN 8163, 220 m), 2 spm., 12/9/1981, UPV ; San Miguel de Aralar , limestone in grass ( WN8554 ), 221 58*1 14#spm., 7/1989, MNHN ; Selva de Oza ( XN8746 , 1220 m), 1 spm., 1/12/1988, UPV ; UsuÂn ( XN4625 ), 6 spm., 29/7/1981, UPV ; Urbasa , km 28 heÃtraie ( WN 6743, 900 m), 2 spm., 23/7/1987, UPV ; Urdax: LeorlaÂs , rocks ( XN 2092, 100 m), 9 spm., 3/12/1987, UPV ; Vera de Bidassoa ( XN0693 ), 7 spm., 24/7/1982, UPV ; Yanci , woodland ( XN0586 , 160 m), 7 spm., 3/12/1987, UPV .

Guipuzcoa: MNHN collection ÐSan SebastiaÂn: Monte Urgull, numerous spm. coll.Staadt.

New materialÐ Ataun ( WN6762 ), 3 spm., 19/3/19831 4 spm., 15/4/1983, UPV ; Aralar: Ausa Gaztelu ( WN 7163 ), 4 spm., 17/04/1981, UPV ; Aranzazu ( WN5060 ), limestone in grass 101 28* spm. (4j), leg. Gofas 22/7/1989, MNHN ; Alkiza ( WN 7280, 340 m), 5 spm., 4/12/1987, UPV ; Azcoitia, road to Vergara at 3 km ( WN4780 ), 1 spm., 29/6/1985, UPV ; Deva-Motrico, limestone in grass ( WN5394 ): 51 12*1 7* spm., leg. Gofas 1/7/1987, MNHN ; Hernialde ( WN 7478, 250 m), 5 spm., 23/3/1986, UPV ; Ibarra ( WN7676 ), 3 spm., 25/10/1979, UPV ; Itziar, among small limestome blocks ( WN 5791, 500 m): 51 16*1 34* spm. (2 juv), leg. Gofas 1/7/1980, MNHN ; Itziar ( WN5791 ), 1 spm., 29/6/1985, UPV ; Loyola, limestone in grass ( WN 5982, 800 m): 31 45* spm., leg. Gofas 4/2/1989, MNHN ; Otzaurte ( WN6054 ), 5 spm., 22/3/1986, UPV ; PenÄa Madariaga ( WN5684 ), 9 spm., 29/6/1985, UPV ; ReÂgil ( WN6781 ), 4 spm., 15/3/19801 6 spm., 23/3/1986, UPV ; San SebastiaÂn: Gladis Enea ( WN8296 ), 1 spm., 3/2/1981, UPV ; San SebastiaÂn: Monte Urgull ( WN8197 ), 7 spm., 23/2/19801 1 spm., 22/2/1981, UPV .

Alava: New materialÐ Angosto ( VN 9445), 6 spm., 24/8/1981 and 3 spm., 26/9/1981, UPV ; DeÂlica ( WN0157 ), 6 spm., 23/07/1982, UPV ; Santa Cruz de Campezo : Ega ( WN 5624 ), 1 spm., UPV .

Vizcaya: MNHN collection ÐBilbao: Paseo de los CanÄos, 12 spm., coll. H. Fischer ex Hidalgo; Bilbao, 4 spm., coll. H. Fischer ex Hidalgo; OrdunÄa, 3 spm., coll. Staadt.

New materialÐ Axpe-Busturia ( WP2303 ), 1 spm., 2/3/1980, UPV ; Bilbao S: Pagasarri ( WN0584 ), 13/10/1979, UPV ; Bilbao N: La Cadena, 2 spm., 24/8/1980, UPV ; Bengoetxea ( WN0577 ), 3 spm., 25/10/1979, UPV ; Durango : Untzillaitz ( WN2975 ), 1 spm., 18/10/1981, UPV ; Erenno ( WP3102 ), 25/4/1980, UPV ; Galdames : Cueva Los Cuervos ( VN 9289), 3 spm., 24/2/1980, UPV ; Galdames : Cueva Los Cuervos ( VN 9289), 2 spm., 4/1/1981, UPV ; LendonÄo de Arriba ( VN 9562), 5 spm., 7/1/1982, UPV ; Gorbea ( WN1766 ), limestone rocks with beech litter, 291 17*1 13#spm., leg. Gofas and MartõÂn 5/1989, MNHN ; Gorbea : Atxas, pastures with beech trees ( WN1569 ), 4 spm., 3/10/1981, UPV ; Gorbea : AldaminÄape ( WN1865 ), 3 spm., 7/3/1982 and 3 spm., 18/3/1982, UPV ; Gorbea : Arimekorta ( WN1866 ), 7 spm., 15/3/1981 and 6 spm., 5/4/1981, UPV ; Gorbea : Atxuri ( WN2065 ), 8 spm., 5/4/1981, UPV ; Gorbea : Rio Padrobaso, beech woods on karst ( WN2164 ), 6 spm., 18/4/1982, UPV ; Gorbea : Urkizu ( WN1981 ), 7 spm., 23/3/1980, UPV ; Gorbea : Urkizu, cueva de Otxas ( WN1981 ), 3 spm., 23/8/1981, UPV ; Gorbea : Aginalde, 1 spm., 15/2/1981, UPV ; Gorbea : S de Artalarra 1 spm., 7/4/1985, UPV ; Itxina ( WN1469 ), 3 spm., 8/7/1979, UPV ; ManÄaria ( WN2876 ), 2 spm., 8/11/1979, UPV ; OrdunÄa : Txarlazo ( VN 9759), 3 spm., 3/5/81, UPV ; PenÄas de Ranero ( VN 6991, 800 m), 61 28*1 11#spm., leg. Gofas and Gomez, 5/1989, MNHN ; PenÄas de Ranero ( VN 6991), 3 spm., 12/8/1981, UPV ; Udalaitz ( WN3871 ), 3 spm., 2/2/1980, UPV ; Urquiola ( WN2771 ), 1 spm., 30/9/1979, UPV .