Plocamocera procera, OPITZ, 2004

OPITZ, WESTON, 2004, Classification, Natural History, And Evolution Of The Epiphloeinae (Coleoptera: Cleridae). Part Ii. The Genera Chaetophloeus Opitz And Plocamocera Spinola, Bulletin of the American Museum of Natural History 2004 (280), pp. 1-82 : 51-53

Plocamocera procera , new species Figures 182 , 214 ; map 9

HOLOTYPE: ( LACM). Colombia: Amazonas : PNN, Amacayacu, Matamata, 3 ° 23 ̍ S, 70 ° 6 ̍ W, 150 m, Malaise, 4/2/01–4/16/01, D. Chota, leg., M. 1609. (Specimen pin mount­ ed; paper point with antenna; sex label affixed to white support card; locality label, white, outlined in black, machine printed; collection method, date of collection, collector label, white, outlined in black; LACM repository label, white, machine printed, holotype label, red, machine printed; plastic vial with abdomen and aedeagus.)

PARATYPES: None.

DIAGNOSIS: Distinguishable from specimens of the sister species, P. confrater Kuwert , by the shape of the 8th antennomere whose anterodistal margin is not angular (compare figs. 193, 182). Also, the aedeagus is more elongated in the members of this species.

DESCRIPTION: Size: Length 6.0 mm; width 2.4 mm. Integument: Cranium castaneous; frons and epicranium darker; pronotum castaneous, disc feebly infuscated in front of subapical depression; remainder of thorax castaneous except pro–mesofemora infuscat­ ed on anterior disc, metafemur black in distal half, tibiae variously infuscated; elytral color variegated, predominantly dark­cataneous, flavotestaceous trigonal posthumeral macula well developed. Head: Antennal club as in figure 182. Thorax: Pronotal anterior margin notably projected at middle; elytral trichobothria not discernible; protibial anterior margin with 4 spines. Abdomen: Male pygidium broad scutiform, aedeagus (fig. 214) lanceolate.

VARIATION: One specimen examined.

NATURAL HISTORY: The available specimen was collected during April with a Malaise trap at 150 m.

DISTRIBUTION (map 9): This species is known only from the type locality.

ETYMOLOGY: The specific epithet procera is a Latin adjectival and means slender. I refer to the extended length of the aedeagus when compared to the aedeagus of males of P. confrater Kuwert.

COACTILIS GROUP Plocamocera coactilis , new species Figures 6–8 , 12 , 46 , 64, 65, 95, 104, 116, 136, 163, 198; map 3

HOLOTYPE: Male. Brazil: Matto Grosso : X­ 1973, M. Alvarenga ( MZSP). (Specimen point amounted, sex label affixed to paper point, white, machine printed; support card, white; locality label, white machine and hand printed; holotype label, red, machine and hand printed.) Opitz, 1997: 70.

PARATYPES: Forty­three specimens. Seventeen specimens from the same locality as the holotype ( AMNH, 1; BMNH, 1; JNRC, 1; LACM, 1; OSUC, 1; USNM, 1; WFBM, 1; WOPC, 10). Bolivia: Cochabamba: Chapare : 5­X­ 1949, 400 m, R. Zischlka ( FMNH, 1). Brazil: Matto Grosso : Sinop ( CASC, 1; CNCI, 1; FMNH, 1; MCZC, 1; WOPC, 3): Goias ( DEIC, 1; WOPC, 3): Rondonia ( WOPC, 1): Amazonas : Manaus , 1 km W Taruma Falls, 19­1­1981, W. Opitz ( WOPC, 1) ; 28­II­1981, C.W. Young ( CMNH, 1): Amazonas ( MNHN, 8; DEIC, 2; ZMHB, 1): Para: 70 km W. Monte Dourado, malaise at edge of natural forest, 27–29­XII­1980, R. Krell ( UCAG, 1) .

DIAGNOSIS: P. coactilis and P. confrater share the tripartite fascia behind the humerus of the elytral disc (compare figs. 64 and 56). In P. coactilis specimens, however, the eighth antennomere is rectangulate and not subquadrate (compare figs. 163 and 152) and the aedeagus is considerably more elongate (compare figs. 112 and 116).

DESCRIPTION: Size: Length 3.5–5.2 mm; width 1.2–2.0 mm. Integument: Cranium piceous, castaneous along ocular margins; pronotum castaneous, with broad discal macula; elytral coloration variegated as in figure 64; legs flavotestaceous, infuscated or not. Head: Antenna as in figure 46. Thorax: Pronotal anterior margin moderately projected at middle; anterior margin of protibia with four spines (fig. 8); elytral epipleural margin of elytra with five conspicuous trichobothria (fig. 65). Abdomen: Male pygidium broadscutiform, very convex in posterior margin (fig. 95), female pygidium broad­scutiform; aedeagus conspicuously elongate (fig. 116). Internal Reproductive Organs: Male as in figure 104.

VARIATION: The legs vary in color, being entirely flavotestaceous in some specimens and infuscated in others.

NATURAL HISTORY: Brazilian specimens were collected from Villa Vera during October, from Goias during November, and from the environs of Manaus during January. I collected the Manaus specimen on a recently felled tree of Manilkara .

DISTRIBUTION ( Map 3 ): Known from the Matto Grosso highlands and Amazon Basin of Brazil, and from northern Bolivia.

ETYMOLOGY: The Latin adjective coactilis (thick). I refer to the robust body of these beetles.

Plocamocera aspera , new species

Figures 153, 199 ; map 7

HOLOTYPE: Female. Brazil: Amazonas : Paraque , 30 km east of Manaus , Amazon River, 18­II­ 1981, 34 m Chen­wen Young ( MZSP). (Specimen point mounted, antenna and sex label affixed to paper point, white, machine printed; support card, white; locality label, white, machine printed; MZSP repository label; holotype label, red machine printed.)

PARATYPES: None.

DIAGNOSIS: Within the coactilis group specimens of P. aspera most closely resemble those of P. paris . However, only in P. aspera specimens is the cranium distinctly rugose and antennomere eight more ovoid than angulate; also, antennomere nine is more narrowly prolonged distally (compare figs. 153, 164).

DESCRIPTION: Size: Length 6.0 mm; width 2.2 mm. Integument: Cranium piceous; pronotum predominantly piceous, castaneous along margins; elytra variegated, with flavotestaceous humeral macula that extends posteriorly, then divides, medialmost flavotestaceous macula prominent; three irregular patches of light setae that connect to abundance of light setae along sutural margin, setae dark in piceous regions; legs flavotestaceous, metafemora faintly infuscated, tarsus flavotestaceous. Head: Antennal club as in figure 153; cranium rugose. Thorax: Pronotal anterior margin prominently projected at middle, discal swelling shallow; epipleural margin with five conspicuous trichobothria. Abdomen: Female pygidium broad­scutiform.

VARIATION: Not observed.

NATURAL HISTORY: The holotype was collected during February, at 34 m.

DISTRIBUTION (map 7): Known only from the type locality.

ETYMOLOGY: The trivial name stems from the Latin adjective aspera (rough). I refer to the rugulose cranium.

Plocamocera paris , new species Figures 164, 204 ; map 7 HOLOTYPE: Female. Brazil: Distrito Federal : Parque Nacional, III– II–1970, J. M. &

B. Z. Campbell (CNCI). (Specimen point mounted; sex label affixed to paper point, white, machine printed; support card, white; locality label, white, machine printed; CNCI repository label white, machine printed; holotype label, red, machine printed; plastic vial with pygidium and sixth visible sternum.)

PARATYPES: None.

DIAGNOSIS: The members of this species resemble members of P. confrater , from which they may be distinguished by the shape of the eighth antennomere (compare figs. 152, 164). Also, in P. paris specimens the humeral and postmedian maculae are more pronounced.

DESCRIPTION: Size: Length 5.3 mm; width 2.2 mm. Integument: Cranium piceous; pronotum piceous; elytra variegated, with flavous humeral macula that extends and divides posteriorly, mostmedial fascia angulate, transverse, three irregular aggregates of white and golden setae; legs bicolorous, pro– mesofemur predominantly flavotestaceous, boldly infuscated, metafemur flavotestaceous in basal half, piceous in remainder. Head: Antenna as in figure 164. Thorax: Pronotal anterior margin moderately projected at middle, discal swelling shallow, pronotal arch scabrous; elytral epipleural margin with four conspicuous trichobothria; protibial anterior margin with four spines. Abdomen: Female pygidium broad­scutiform.

VARIATION: Not observed.

NATURAL HISTORY: The holotype specimen was collected during March from the type locality, at 1000 m.

DISTRIBUTION (map 7): Known only from the type locality.

ETYMOLOGY: The trivial name paris is a Latin adjective that means ‘‘equal’’. I refer to the superficial similarity of this beetle to other members of the P. confrater group.

Figures

Figs. 181–195. Antennal clubs. 181. Plocamocera confrater. 182. P. procera. 183. P. manausensis.

Figs. 213–218. Aedeagi and antenna. Aedeagi. 213. Plocamocera castanea. 214. P. procera. 215. P. confrater. 216. P. buenavista. Antenna. 217. P. buenavista. 218. P. aura.

Figs. 6–11. Plocamocera coactilis; 6. Head (1, labrum, 2, ocular notch, 3, scape, 4, pedicel). 7. Pronotum (5, trichosome, 6, discal trichbothrium). 8. Distal portion of protibiae (7, spine). P. confrater. A. antenna, 90× (11, filamentous sensilla trichodea). B. Ninth antennomere. 9. Pronotal discal trichobothrium (8, bothrium, 9, trabiculae, 10, trich). 10. Antenna. 11. Antennomere 1200× (12, sensilla chaetica I; 13, filamentous sensilla trichodea, 14, sensilla basonica).

Figs. 12–15. 12. Plocamocera coactilis; tenth antennomere (1, filamentous sensilla trichodea; 2, sensilla chaetica II). 13–15. P. confrater. 13. pronotal lateral trichobothrium (3, trich; 4, bothrium). 14. Pronotum (5, sensilla trichodea I; 6, basal peg of sensilla trichodea II). 15. Pronotum (7, basal peg of sensilla trichodea II).

Figs. 40–51. Antennae. 40, Plocamocera castanea. 41, P. pupula. 42, P. confrater. 43, P. aliguantula. 44, P. minima. 45. P. manausensis. 46, P. coactilis. 47, P. sericella. 48, P. argentea. 49, P. auratilis. 50, P. sesquipedalis. 51, P. lucis.

Maps 1–3. (1) Distribution of Plocamocera castanea; (2) Distribution of Plocamocera confrater;

Figs. 196–212. Antennal clubs. 196. Plocamocera sesquipedalis. 197. P. similis. 198. P. coactilis. 199. P. aspera. 200. P. santa. 201. P. onorei. 202. P. bispina. 203. P. carnegei. 204. P. paris. 205. P. ambra. 206. P. bolivari. 207. P. auratilis. 208. P. insula. 209. P. selva. 210. P. cericellopsis. 211. P.