Perrona styriaca ( Hilber, 1879 ) Harzhauser & Landau & Janssen, 2022
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publication ID |
https://doi.org/ 10.11646/zootaxa.5123.1.1 |
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publication LSID |
lsid:zoobank.org:pub:036F6B4D-CDCC-4CD7-A914-9A1D8C7A097A |
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persistent identifier |
https://treatment.plazi.org/id/039487D1-FF00-FF2B-FFBA-FB006930F9E7 |
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treatment provided by |
Plazi |
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scientific name |
Perrona styriaca ( Hilber, 1879 ) |
| status |
comb. nov. |
Perrona styriaca ( Hilber, 1879) View in CoL nov. comb.
Figs 40A View FIGURE 40 1 –A View FIGURE 1 3 View FIGURE 3 , B 1 –B View FIGURE 1 3 View FIGURE 3 , 4V View FIGURE 4 , 5 View FIGURE 5 , 7 View FIGURE 7
* Pleurotoma (Clavatula) styriaca Auing. in coll.— Hilber 1879: 19, pl. 3, figs 6–7.
Pleurotoma (Clavatula) styriaca Auing. — Hoernes & Auinger 1891: 348 (partim), pl. 47, figs 4–5 [non figs 6–9 = Clavatula pilleri nov. sp.; non fig. 10 = Perrona lydiae (Hoernes & Auinger, 1879) View in CoL ].
Clavatula styriaca Hilber View in CoL , sp. 1879— Glibert 1960: 42.
non Clavatula styriaca Auinger View in CoL — Friedberg 1912: 198, pl. 12, fig. 7 [= Perrona ilonae View in CoL nov. nom.]. non Clavatula styriaca Auing View in CoL , var.— Friedberg 1912: 199, text-fig 4 [= Megaclavatula antoniae ( Hoernes & Auinger, 1891) View in CoL ]. non Clavatula styriaca Auing. View in CoL — Friedberg 1938: 143 [= Megaclavatula antoniae ( Hoernes & Auinger, 1891) View in CoL ]. non Clavatula styriaca Auing. View in CoL — Strausz 1954: 33, pl. 5, fig. 103 [unidentifiable from picture]. non C [lavatula]. (C [lavatula]) styriaca (Auing.) View in CoL — Sieber 1958a: 157 [= Clavatula pilleri nov. sp.]. non Clavatula (Clavatula) styriaca Auinger View in CoL — Švagrovský 1958: 12, pl. 2, figs 5–7 [= Perrona laevigata ( Eichwald, 1830) ]. non Clavatula styriaca (Hilb.) View in CoL — Kókay 1966: plate captions, pl. 9, fig. 15 [= Perrona ilonae View in CoL nov. sp.]. non Clavatula styriaca Auinger View in CoL (in Hilber)— Strausz 1966: 402, pl. 15, figs 11–14 [= Perrona laevigata ( Eichwald, 1830) ]. non Clavatula striaca [sic] [(Auinger) Hilber, 1879]— Zelinskaya et al. 1968: 221, pl. 50, figs 20–21 [not a Clavatulidae View in CoL ]. non Clavatula styriaca (Auinger) View in CoL — Bałuk 1970: 118, pl. 13, fig. 6 [= Megaclavatula antoniae ( Hoernes & Auinger, 1891) View in CoL ]. non Clavatula (Clavatula) styriaca ( Hilber, 1879) View in CoL — Švagrovský 1982: 413, pl. 8, fig. 2 [= Perrona laevigata ( Eichwald, 1830)]. non Clavatula (Clavatula) cf. styriaca ( Hilber, 1879) View in CoL — Mikuž 1998: 73, pl. 4, fig. 1 [= Perrona laevigata ( Eichwald, 1830) ]. non Clavatula (Clavatula) styriaca ( Hilber, 1879) View in CoL — Mikuž 1998: 73, pl. 3, fig. 3 [= Clavatula pilleri nov. sp.]. non Clavatula styriaca ( Hilber, 1879) View in CoL — Bałuk 2003: 38, pl. 6, fig. 4 [= maybe an aberrant Perrona laevigata ( Eichwald, 1830)]. non Clavatula styriaca ( Hilber, 1879) — Mikuž 2009: 32, pl. 10, fig. 139 [= Clavatula pilleri nov. sp.].
Type material. Lectotype: NHMW 1846/0037/0309a, Pöls ( Austria), SL: 21.8 mm, MD: 9.3 mm, illustrated in Hilber (1879, pl. 3, figs 7a–c) and Hoernes & Auinger (1891, pl. 47, figs 5a–c), designated by Švagrovský (1982: 412), figs 40A 1 –A 3, 4 V. Paralectotypes: NHMW 1846/0037/0309b, Pöls ( Austria), SL: 40.5 mm, MD: 16.1 mm, illustrated in Hilber (1879, pl. 3, fig. 6) and Hoernes & Auinger (1891, pl. 47, fig. 4, figs 40B 1 –B 3; NHMW 1949/0005/0051f, Pöls ( Austria), SL: 44.7 mm, MD: 18.8 mm, illustrated in Hoernes & Auinger (1891, pl. 47, fig. 8); NHMW 1949/0005/0051g, Pöls ( Austria), SL: 49.9 mm, MD: 18.2 mm, illustrated in Hoernes & Auinger (1891, pl. 47, fig. 9); NHMW 1861/0001/0266, 2 spec., Pöls ( Austria).
Revised description. Shell small, solid, moderately broad fusiform with high spire; apical angle 30–33°. Protoconch not preserved. Teleoconch of ten whorls. Early teleoconch whorls flat-sided with two smooth, narrow adsutural spiral cords separated by broad mid-portion bearing fine, opisthocline axial riblets. Riblets fading out on fourth or fifth teleoconch whorls. Later whorls concave, with periphery at abapical suture. Prominent subsutural collar bearing widely-spaced pointed tubercles placed at mid-collar, developed from eighth whorl. Suprasutural cord almost completely obscured by succeeding whorl. Spiral sculpture of delicate spiral threads weakening on last whorl. Suture narrowly incised, superficial, linear. Last whorl ~60% of total height; of medium width. Weakly coronate subsutural collar bearing 10–12 pointed tubercles. Subsutural ramp wide, concave, delimited by angular shoulder placed at level of insertion of outer lip. Base placed high, in short distance below shoulder, weakly angled, strongly constricted. Weak siphonal fasciole. Entire surface covered in fine spiral cords of alternating strength and slightly stronger over base and fasciole. Aperture wide, pyriform. Outer lip not thickened, smooth within. Anal sinus moderately wide and deep, asymmetrically U-shaped, with apex mid-ramp; siphonal canal moderately long, narrow, slightly deflected to the left, shallowly notched at tip. Columella strongly excavated, twisted at fasciole, smooth. Columellar and parietal callus moderately thickened, adherent, forming broad callus rim.
Discussion. This species was until now placed in the genus Clavatula by all authors. However, the commashaped axial riblets on early teleoconch whorls suggest a placement in Perrona . Perrona styriaca ( Hilber, 1879) is surprisingly rare and the wide distribution suggested by the literature records is based on numerous misidentifications. The concept of this species was also blurred by Hoernes & Auinger (1891) by including an aberrant specimen of Perrona lydiae ( Hoernes & Auinger, 1891) and by including specimens of Megaclavatula pilleri nov. sp. from Baden-Sooss ( Austria) in their description. Perrona lydiae , however, is clearly separated from P. styriaca by its swollen adsutural cords, higher spire and more slender outline. Megaclavatula pilleri is distinguished by its tripartite sculpture on the early teleoconch whorls, as opposed to the comma-shaped axial riblets typical of Perrona species. Several authors placed small sized and subadult shells of Megaclavatula laevigata ( Eichwald, 1830) in P. styriaca (e.g., Švagrovský 1958, Strausz 1966), which all differ from P. styriaca in having tripartite early teleoconch whorl sculpture. Similarly, the single specimen from Korytnica, reported by Bałuk (2003) as P. styriaca has a reduced peribasal spiral cord and differs from P. styriaca also in its shorter spire and wider apical angle. Specimens from the Carpathian Foredeep, referred to as Clavatula styriaca by Friedberg (1912, 1938) and Bałuk (1970) were described as Clavatula kowalewskii by Bałuk (2003) [= Megaclavatula antoniae ( Hoernes & Auinger, 1891) ], which differs from P. styriaca in its slender fusiform outline, a conical spire and strongly reduced sculpture on the subsutural collar.
Paleoenvironment. Shallow marine, inner neritic, based on the mollusc assemblage from Pöls ( Austria) (own data).
Distribution in Central Paratethys. Badenian (middle Miocene): Styrian Basin: Pöls ( Austria) ( Hoernes & Auinger 1891).
| NHMW |
Naturhistorisches Museum, Wien |
| MD |
Museum Donaueschingen |
| V |
Royal British Columbia Museum - Herbarium |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Perrona styriaca ( Hilber, 1879 )
| Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald 2022 |
Clavatula styriaca
| Glibert, M. 1960: 42 |
Pleurotoma (Clavatula) styriaca Auing.
| Hoernes, R. & Auinger, M. 1891: 348 |
Pleurotoma (Clavatula) styriaca Auing.
| Hilber, V. 1879: 19 |