Penthetria luberonica, Skartveit, John & Nel, André, 2017

Skartveit, John & Nel, André, 2017, Revision of fossil Bibionidae (Insecta: Diptera) from French Oligocene deposits, Zootaxa 4225 (1), pp. 1-83 : 29-31

Penthetria luberonica sp. n.

(Figs. 12 – 13, 90 – 94)

Material examined. Holotype (male) FSL 140048b Paratype (males) FSL 140050, FSL 140053g.

Additional material (7 males 20 females): FSL 140009, 140010, 140021, 140023, 140027, 140034, 140036, 140038, 140041, 140046, 140048a, 140049, 140053, 140279, 140307, 140312.

Etymology. The species is named after the Lubéron Massive, where the outcrop is situated.

Localities. Vitrolles.

Diagnosis. A small Penthetria , total length 7 – 9 millimetres. Antennal flagellum 9-segmented. Wing light brownish fumose, R2+3 of medium length, slightly sinuate. Male gonostylus robust, curved and pointed, epandrium dome-shaped. Most similar to P. claripennis , but recognized by basal part of Rs being nearly parallel to M.

Description. Male: Total length 7.0 – 9.5 mm (N=8).

Head (fig. 91): Length 0.92 – 1.21 mm (N=5), width 0.96 – 1.21 mm (N=5). Brown. Compound eye rather small, subspherical, prominent. Flagellum 9-segmented, length 0.47 – 0.65 mm (N=3).

Thorax: Length 2.0 – 2.7 mm (N=8), width 1.8 mm (N=1). Dark brown, presumably black in life. Haltere pale brown.

Wing (fig. 90): Length 5.3 – 7.0 mm (N=7), width 2.0 – 2.8 mm (N=7), length/width =2.52 – 2.82 (N=7). Light brown fumose, anterior part slightly darker, pterostigma inconspicuous. Veins light brown throughout. Costa extends to half-way between apices of R4+5 and M1. Humeral vein present. Subcosta relatively strong, terminating in Costa just above origin of R2+3. Basal part of Rs subparallel to M, crossvein R-M nearly vertical. R2+3 of medium length, nearly straight to slightly sinuous. Fork of M rounded. Fork of CuA relatively short, just a little longer than stem. Wing vein measurements in Table 1 .

Legs: Pale brown, not well preserved in any of the specimens at hand.

Abdomen: Length 4.4 – 6.4 mm (N=7), width 0.9 – 1.0 mm (N=4). Brown, quite slender.

Terminalia (fig. 92): Hypopygium width 0.58 – 0.93 mm (N=6). Gonostylus length 0.43 – 0.47 mm (N=2). Blackish-brown. Epandrium presumably dome-shaped. Gonostylus robust, apically curved and sharply pointed.

Fig. 91 . Male, head and thorax. ECL 140.050. Fig. 92 . Male, terminalia. ECL 140.053/7. Fig. 93 . Female, wing. ECL 140.046.

Fig. 94 . Female, head and thorax. ECL 140.053/5.

Female: Total length 6.7 – 8.2 mm (N=11).

Head (fig. 94): Length 0.67 – 0.76 mm (N=2), width 0.67 – 0.94 mm (N=8). Dark brown, presumably black in life. Flagellum probably 9-segmented, length 0.54–0.62 mm (N=3).

Thorax: Length 1.9 – 2.4 mm (N=12), width 1.5 – 1.9 mm (N=7). Dark brown, presumably black in life. Haltere pale brownish.

Wing (fig. 93): Length 6.1 – 7.5 mm (N=12), width 2.6 – 3.0 mm (N=11), length/width = 2.29 – 2.57 (N=11). Light brownish, slightly darkened anteriorly, pterostigma indistinctive. Humeral vein present, subcosta distinctive, long and straight. Costa extends to nearly half-way between apices of R4+5 and M1. Basal part of Radial sector subparallel to M. R2+3 oblique, nearly straight. Fork of M rounded. Wing vein measurements in Table 1 . Fork of CuA not much longer than stem. CuP distinctive, straight. Wing vein measurements in Table 1 .

Legs: Not well preserved in any of the specimens at hand. Pale brown, tarsi a little darker. Fore tibia length 1.8 – 1.9 mm (N=4).

Abdomen: Length 4.7 – 6.1 mm (N=11), width 1.5 – 2.3 mm (N=8). Brown, apically darkened.

Terminalia: Dark brown, presumably black in life. Tergite VIII rectangular with slightly protruding hind corners. Sternite VIII mesally with a small, rounded indentation.

Figures

FIGURES 90 – 94. Penthetria luberonica sp. n. Fig. 90. Male, wing. ECL 140.050.