Pedaridium maya, Vaz-De-Mello, Fernando Z., Halffter, Gonzalo & Halffter, Violeta, 2004

Vaz-De-Mello, Fernando Z., Halffter, Gonzalo & Halffter, Violeta, 2004, A New Species of Pedaridium Harold from Mexico and Guatemala (Coleoptera: Scarabaeidae: Scarabaeinae: Coprini: Ateuchina), The Coleopterists Bulletin 58 (2), pp. 247-252 : 248-251

Pedaridium maya new species

Figs. 1–9

Type Specimens. Holotype male: México: Quintana-Roo, 5 km N Carrillo Puerto, IX- 1984, A. Martínez (IEX – ex Halffter).

Allotype female: México: Quintana-Roo, Reserva de Sian Ka’an, VIII- 1984, NTP 4 A, M. A. Morón (IEX – ex Morón).

Paratypes 24 unsexed specimens as follows:

Guatemala : Petén , San Andrés, San Miguel La Palotada, 06-VI-1999, M. Tolon, EX1-L4-KK3 (1 UVG); Tikal, 23–26-VIII- 1972, S. & J. Peck (1 Halffter, 1 Vaz-de- Mello, 1 CMN) .

México: Campeche, Chicana, 10 km W Xpujil, 300 m, 12–14-VII- 1983, S. & J. Peck (1 CMN); Chiapas, Pque. Nal. Sumidero, Coyote Mirador, 1,700 m, 19-VI-1989, H. Howden (1 CMN); Pque. Nal. Sumidero, 26-V–1-VI-1990, B. Gill, dung (2 Gill); Pque. Nal. Sumidero, 21-VI-1989, E. Zuccaro & P. K. Lago (2 Gill); Quintana-Roo, 100 9 20 mi S Felipe C Puerto, 13-VIII-1971, A. Newton (1 Halffter, 1 Vaz-de-Mello); 5 km N Carrillo Puerto, IX-1984, A. Martínez (1 Halffter); Reserva de Sian Ka’an 1 km al NNE del Rancho ‘‘E124’’, 23-VIII-1985, J. F. Camal, trampa NTP 80 (1 Halffter); Reserva de Sian Ka’an 2 km al W del Rancho ‘‘El Ramonal’’, 23-VIII-1985, O. Canul, trampa NTP 80 (1 Halffter); Reserva de Sian Ka’an 5 km del Crucero del camino nuevo a Vigía Chico, 23-VIII-1985, O. Canul, trampa NTP 80 (1 Halffter); Reserva de Sian Ka’an bosque tropical subcaducifolio, 10 m, 19-VI-1984, O. Canul & S. F. Camal, trampa NTP 80 (2 Halffter, 1 Vaz-de-Mello, 1 Morón); Reserva de Sian Ka’an km 12 carretera nueva al Rancho ‘‘El Ramonal’’, 19-VI-1984, J. F. Camal, trampa NTP 80 (1 Halffter); 5 km N Carrillo Puerto, IX-1984, A. Martínez (2 CMN); Reserva de Sian Ka’an 300 m S de la unión de caminos nuevo y antiguo al Rancho Yuras, 10 m, 19-VI- 1984, O Canul & SF Camal, selva tropical subcaducifolia, NTP80 (1 Delgado).

Description. Male: Color dark brown to black, body oval, flattened. Clypeus rounded, with two small teeth separated by wide v-shaped emargination. Clypeo-frontal suture not marked, clypeo-genal suture very weak. Eyes wide dorsally, interocular space about six eye widths. Head dorsally covered with irregular setose punctures ( Fig. 1 ), setae small and erect. Posterior part of the head with scattered punctures, posterior margin lacking. Inferior clypeal process in strong angle. Mentum anteriorly emarginate, medially, longitudinally depressed; submentum separated from gula by feeble posteriorly rounded suture. Antennae with large scape (about as long as the five following segments), lamellae large.

Pronotum evenly convex, sides almost straight in dorsal view, slightly sinuated in the posterior border. Both anterior and posterior angles obtuse, posterior angle marginate by a carina ( Fig. 3 ). Anterior and posterior borders lacking marginal bead. Pronotal surface covered with small scattered setigerous punctures ( Fig. 2 ), setae small, yellow in color; surface completely microgranulated, with flattened granules. Prosternum projected as small spine between anterior coxa. Meso-and metasternum separated by horseshoe-like suture, metasternum medially with dense setigerous punctures. Fore femur ventrally setosely punctured. Fore tibia with three strong lateral teeth, middle one broader, all with acutes apices. External tibial face denticulate from base to basal tooth, apex truncated right angled. Apical spur short, narrower than in females, slightly curved ventrally, with acute apex. Fore tarsus larger than apical tibial breadth, 5th tarsomere with dorsal setose filiform projection; claws small, simple and curved, with small basal, acute denticle. Middle leg with trochanter distaly with one setigerous puncture. Ventral face of femur with short setose punctures. Middle tibia regularly broader apically, ventro-externally with a very small median tubercle, ventro- and dorso-external borders shortly bristled and with longitudinal carinae. Both spurs acutely spiniform, the ventral one larger than tarsomere 1, the dorsal one slightly smaller. Middle tarsus quite longer than tibia, 1st tarsomere subequal to 2nd in length, claws similar to fore ones, but apical seta in 5th tarsomere shorter. Hind legs with trochanter distally with many setigerous punctures, setae slender and unequal in length. Hind femur punctured like the medium one. Hind tibia apical regularly broader, laterally bristled, but without longitudinal carinae. Hind tibial spur acutely spiniform, longer than 1st tarsomere. Second tarsomere narrower than 1st, other tarsomeres and claws similar to those of middle legs.

Elytra with eight striae, including very deep lateral one. Striae slender, well defined, not punctured in the disc, seventh shorter, eighth with lateral sulcus defining discal margins. Interstriae feebly convex on disc, with one line of small setigerous punctures, parallel to internal stria. Punctures variable in deepth and size. Apically, striae much deeper and with large punctures ( Fig. 4 ).

Abdominal sternites covered by ocellate punctures laterally, 6th sternite ventrally with setae on ocellate punctures ( Fig. 5 ). Pygidium strongly convex, basally with a deep straight sulcus, apically and laterally with a feeble margin ( Fig. 6 ). Disc covered by sparse setose punctures, setae short and erect.

Aedeagus with phallobase larger than parameres, these positioned at right angle in relation to phallobase ( Fig. 7 ). Each paramere conical, curved downwards, without marginal carinae or angles, regularly tapered from basis to about two-thirds, then strongly tapered and curved inwards, with parallel round apex ( Fig. 8 ).

Total length 3.7 to 4.4 mm; pronotal width 1.1 to 1.7 mm; elytral width 1.9 to 2.1 mm.

Female: Differs from male in broader fore tibia, with lateral teeth slender and less conspicuous, fore tibial spur shorter, broader basally, shape triangular and slightly curved; abdominal sternite VI longer medially, pygidium wider than long.

Total length 4.6 to 4.8 mm; pronotal width 1.85 to 1.9 mm; elytral width 2.7 to 2.8 mm.

Figures

Figs. 1–8. Pedaridium maya new species. 1) Head; 2) punctures on pronotal disc; 3) pronotal hind angle; 4) elytral apex; 5) abdomen, ventral view; 6) pygidium; 7) aedeagus, lateral view; 8) parameres, dorsal view.

Fig. 9. Known collecting sites of Pedaridium maya new species in Mexico and Guatemala.