Patellapis (Patellapis) bidentata, Timmermann & Kuhlmann, 2009

Timmermann, Kim & Kuhlmann, Michael, 2009, Variable Selection and Inference for Multi-period Forecasting Problems, Zootaxa 2099, pp. 1-188 : 44-47

Patellapis (Patellapis) bidentata sp. n. Timmermann

( Figs. 25a–g ; 26a–h )

Diagnosis. The males of P. bidentata are characterised by the bidentate mandibles. The females of P. bidentata can be distinguished from those of P. minutior by having strong apical hair bands on the metasomal terga. Further, the surface of paraocular area of P. bidentata females is considerably more coarse sculptured than in P. minutior . The females of P. braunsi and P. schultzei are unknown.

Description

Female. Bl = 7,4–7,5mm. General habitus ( Fig. 25e ). Head. L = 2,1–2,2mm; W = 2,2–2,3mm. Head about as wide as long ( Fig. 25a ). Integument black except mandibles sometimes partly reddish-brown. Face shiny and with loose, long, brownish and yellowish, erect hairs. Mandibles bidentate. Malar area very short, linear. Clypeus about two times as wide as long; coarsely and densely punctate (i = d). Clypeoantennal distance 0,6mm. Supraclypeal area about as long as wide; strongly and densely punctate (i = 1,0–1,5d); surface sometimes finely chagreened. Clypeus and supraclypeal area strongly convex in profile. Paraocular area coarsely and densely punctate; surface partly sculptured. Antennae blackish to brownish. Mesosoma. L = 1,5–1,6mm; W (ITS) = 1,7–1,9mm. Integument black. Scutum shiny; partly chagreened anteriorly; disc densely and strongly punctate (i <d) ( Fig. 25b ). Scutellum as illustrated in Fig. 25c . Scutum, scutellum and metanotum with yellowish, erect hairs; mesepisternum and propodeum with greyish and yellowish, erect hairs. Basal area of propodeum as illustrated in Fig. 25c ; propodeum shiny; strongly and densely punctate; marginal region of posterior truncation slightly dull; loosely punctate and strongly sculptured. Wings. Hyaline. Legs. Integument brownish to black. Vestiture greyish and yellowish. Ts finely serrate ( Fig. 25f ). Bp broad and pointed at apex ( Fig. 25g ). Metasoma. L = 4,3mm; W = 2,5mm. Integument black except the margins of T pallid brownish to pallid yellowish(usually T1–T2); margins partly translucent. T1–T4 with greyish apical hair bands; the bands mostly present only laterally. Prepygidial fimbria mostly brownish. Discs and marginal zones of T1–T2 shiny and densely punctate (i = 1,0–1,5d) ( Fig. 25d ).

Male. Bl = 7,4–7,5mm. General habitus ( Fig. 26e ). Head. L = 1,8–1,9mm; W = 2,1–2,2mm. Head about as wide as long. Integument black except mandibles sometimes partly reddish-brown. Clypeus, supraclypeal area and paraocular area covered with erect, greyish to yellowish hairs; the lower half of paraocular area, supraclypeal area and clypeus also covered with appressed, whitish hairs. Mandibles bidentate. Malar area very short, linear. Antennae brownish-black. Flagellomeres knotty. A4 about 2,5 times longer than A3.

Mesosoma. L = 1,4–1,5mm; W ( ITS) = 1,6–1,7mm. Integument black. Scutum densely and strongly punctate (i = d) ( Fig. 26a ). Scutellum as illustrated in Fig. 26b . Scutum, scutellum,metanotum, mesepisternum and propodeum with very long, greyish to whitish, erect hairs. Basal area of propodeum as illustrated in Fig. 26b ; propodeum shiny, strongly and densely punctate. Marginal region of posterior truncation shiny. Wings. Hyaline. Legs. Integument brownish to black. Vestiture mostly greyish. Metasoma. L = 4,7–4,8mm; W = 2,0– 2,1mm. Integument black except the margins of T pallid brownish to whitish (usually T1 – T2 ); margins partly translucent. Discs and marginal zones of T1 – T2 shiny and densely punctate ( Fig. 26c ). Apical plate slender and angulate. Posterior margin of S4 with a dense row of coarse, erect bristles. S5 with strong, erect hair patches laterally ( Fig. 26d ). S7 and S8 as illustrated in Fig. 26f . Genitalia. Genitalia as illustrated in Fig. 26g –h ; gonocoxa dull and dorsal gonostylus with a fine and slender dorso-lateral projection .

Type material ( 118 specimens). Holotype, male, South Africa, Northern Cape, Nieuwoudtville Wild Flower Reserve East, S 31°21'56" / E19°08'52", 735m, 07.ix.2003, on Oxalis spec. , leg. M. Kuhlmann, SANC.

Paratypes: 83♀, 34♂. South Africa. Western Cape : 1♀, Uniondale District , x.1952, leg. Mus. Exp., SAM – HYM – B009613 , SAMC ; 26♀, Kunje farm, 28 km SE Citrusdal, 23.ix.2001, leg. C.D. Eardley, SANC ; 2♀, Citrusdal , 08.ix.1966, leg. C.D. Michener, SEMC ; 1♀, 10 km W Algeria, Clanwilliam road, 04.ix.1987, leg. C.D. Eardley, Database No. HYMA 07285 , SANC ; 1♀, Clanwilliam , 08.ix.1966, leg. M. and C.D. Michener, SEMC ; 1♀, Sparaxis Hill, S of Clanwilliam , 06.ix.1997, leg. P. Goldblatt, SEMC ; 1♀, Clanwilliam 20 km E, 08.ix.2001, leg. C.D. Eardley, SANC ; Northern Cape: 1♀, 20km S Nieuwoudtville, Farm Papkuilsfontein, Fynbos , 09.ix.2007, leg. K. Timmermann, KTPC ; 1♀, Papkuilsfontein , 15 km S Nieuwoudtville, malaise trap, 09.x.2002, leg. F.D. Parker and M.E. Irwin, BLCU ; 1♂, Nieuwoudtville, Farm Glen Lyon, Renosterveld , 03.vi.1987, leg. P. Goldblatt, SEMC ; 1♀, 1♂, idem., 23.viii.2003, leg. M. Kuhlmann, KTPC ; 1♂, 10.ix.2003, KTPC ; 3♂, idem., 12.ix.2003, KTPC ; 1♂, idem., 07.viii.2004, KTPC ; 2♀, Nieuwoudtville, Farm Glen Lyon, Renosterveld , Pf , 25.viii.2006, leg. K. Timmermann, KTPC ; 1♀, Nieuwoudtville Wild Flower Reserve , 26.viii.1984, leg. V . B. Whitehead and M. Macpherson , SAM – HYM – B009692 , SAMC ; 2♀, idem., 27.viii.1984, leg. V . B. Whitehead and M. Macpherson , SAM – HYM – B009688 , SAMC ; 1♀, idem., 29.viii.1984, leg. V . B. Whitehead and M. Macpherson , SAM – HYM – B009686 , SAMC ; 1♀, idem., 28.viii.1985, leg. V . B. Whitehead and M. Macpherson , SAM – HYM – B009636 , SAMC ; 2♀, idem., 15.viii.1996, leg. V . B. Whitehead , SAM – HYM – B009690 , SAMC ; 3♂, idem., 16.viii.2007, leg. K. Timmermann, KTPC ; 1♀, Nieuwoudtville Wild Flower Reserve, Car Park East , yellow pan trap, 28.ix.2006, leg. K. Timmermann, KTPC ; 2♀, idem., 04.x.2006, leg. M. Kuhlmann, KTPC ; 3♂, idem., 20.viii.2007, leg. K. Timmermann, KTPC ; 1♀, Nieuwoudtville Wild Flower Reserve , dolerite hills, 10.ix.2006, leg. K. Timmermann, KTPC ; 1♀, idem., 18.ix.2006, KTPC ; 1♀, 21.viii.2007, KTPC ; 1♀, idem., 29.viii.2007, KTPC ; 1♀, idem., 08.ix.2007, KTPC ; 1♀, Nieuwoudtville Wild Flower Reserve East, 06.ix.2002, leg. M. Kuhlmann, KTPC ; 1♀, idem., 11.ix.2002, KTPC ; 1♀, idem., 13.ix.2002, KTPC ; 2♂, idem., 04.ix.2003, KTPC ; 3♀, 5♂, idem., 07.ix.2003, KTPC ; 5♀, 4♂, idem., 11.ix.2003, KTPC ; 2♀, 6♂, idem., 10.viii.2004, KTPC ; 1♀, 2♂, idem., 12.viii.2004, KTPC ; 1♀, idem., 21.viii.2004, KTPC ; 1♀, Nieuwoudtville Wild Flower Reserve West, 31.viii.2002, leg. M. Kuhlmann, KTPC ; 1♀, Farm Toren , river bank, 25.viii.1985, leg. V . B. Whitehead and M. Macpherson , SAM – HYM – B009633 , SAMC ; 1♀, Nieuwoudtville Waterfall, 23.–30.ix.1994, leg. F.W. and S.K. Gess, AMGS ; 4♀, 15 km NW Nieuwoudtville, Farm Engelsepunt, Fynbos , 24.ix.2003, leg. K. Timmermann, KTPC ; 1♀, idem., 31.viii.2007, KTPC ; 1♀, 15 km NW Nieuwoudtville, near farm Engelsepunt, Fynbos , 23.ix.2003, leg. K. Timmermann, KTPC ; 1♀, Groenekloof , 7,5 km SE of Leliefontein, 13.ix.1992, leg. F.W. and S.K. Gess, No. 92/93/76, AMGS ; 2♂, Kamiesberg , 11.ix.1987, leg. M.W. Mansell, SANC ; 1♀, Kamiesberg , 2 km S of Leliefontein, 13.ix.1992, leg. F.W. and S.K. Gess, No. 92/93/73, AMGS ; 2♀, Springbok, Goegap Nature Reserve (hills), 8.–10.ix.1992, leg. F.W. and S.K. Gess, No. 92/93/37, AMGS ; 1♀, Springbok, Goegap Nature Reserve (plains), 07.ix.1992, leg. F.W. and S.K. Gess, No. 92/93/21, AMGS ; 1♂, Hester Malan Nature Reserve , 30.vii.1986, leg. M. Struck, SANC ; 1♀, Bowesdorp , ix. 1941, leg. Mus Exp., SAM – HYM – B009596 , SAMC .

Etymology. Named after the bidentate mandibles of the males.

Distribution ( Fig. 28). Records from different locations of the winter rainfall area.

Floral visitation. Senecio arenarius , Senecio spec. (Asteraceae) , Heliophila spec. (Brassicaceae) , Lebeckia sericea (Fabaceae) , Homeria bifida , Romulea montana , Sparaxis fragrans (Iridaceae) , Oxalis spec. (Oxalidaceae) , Nemesia leipoldtii (Scrophulariaceae) , Selago minutissima (Selaginaceae) , Hermanni a spec. ( Sterculiaceae ),

Seasonal activity (first–last observations). vi–x.

Remarks. This species is treated as Patellapis new species 2 in Timmermann and Kuhlmann (2008b).

Figures

FIGURE 25. Patellapis (P.) bidentata, female: a. Head (scale=1mm); b. Scutum (scale=1mm); c. Scutellum and propodeum (scale=0,5mm); d. Metasomal terga (scale=1mm); e. Body (scale=1mm); f. Inner hind tibial spur (scale=100µm); g. Basitibial plate (scale=100µm).

FIGURE 26. Patellapis (P.) bidentata, male: a. Scutum (scale=1mm); b. Scutellum and propodeum (scale=0,5mm); c. Metasomal terga (scale=1mm); d. Metasomal sterna (scale=1mm); e. Body (scale=1mm); f. Metasomal sterna S7 and S8, ventral view (scale=0,5mm); g. Genitalia, dorsal view (scale=0,5mm); h. Genitalia, lateral view (scale=0,5mm).