Paracrossidius radekcervenkai Nakládal, 2017

Červenka, Radek & Nakládal, Oto, 2017, Paracrossidius radekcervenkai (Coleoptera: Scarabaeidae: Aphodiinae), a new species from China, Zootaxa 4268 (4), pp. 597-600 : 597-598

Paracrossidius radekcervenkai Nakládal , new species Figs. 1–4.

Type locality. CHINA, Yunnan Province, pass 20 km NW of Zhongdian.

Type material. Holotype (♂), allotype (♀), and 26 paratypes (♂♂) labelled “ CHINA mer., Yunnan prov., pass 20 km NW from Zhongdian , 15.–16. 6. 2005, Oto Nakládal lgt.” ; 142 paratypes (127♂♂, 15♀♀) labelled “N YUNNAN 22.–24.VI.1994, 27.49N 99.34E 3800–4200m, mts. 15 km W of ZHONGDIAN, lgt. D. Král & J. Farkač ”. Holotype , allotype, and five paratypes deposited in National Museum, Prague, Czech Republic ( Jiří Hájek ) ; 142 paratypes David Král collection (deposited in National Museum, Prague collection) ; 21 paratypes deposited in Oto Nakládal personal collection, Prague, Czech Republic.

Description. Length: males 6.5–8.8 mm (holotype 8.5 mm), female: 6.8–7.7 mm; length of elytra and pronotum: 5.5–7.2 mm.

Male ( Figs. 1–3). Feebly convex; head and pronotum moderately glossy; elytra shagreened, less glossy than head and pronotum. Head, pronotum, and basal two thirds of elytra glabrous, the apical third and the lateral intervals up to the midlength of elytra setaceous. Body oblong-oval, only slightly widened away from head. Colour of elytra and pronotum brown; head darker; sides of pronotum and sutural area of elytra lighter, reddish brown; legs lighter brown.

Head transverse, almost semi-circular, anteriorly rounded. Maximum width of the head slightly narrower than maximum width of the pronotum. Frontal suture thin, almost indistinct, V-shaped. Surface flat, without trace of tubercles. Clypeus anteriorly rounded, not emarginate, sides towards genae straight. Entire clypeal margin distinctly bordered, border thickened on anterior margin. Genae large, strongly protruding beyond eyes, directed laterally with tips forming sharp angles, with several fine setae. Punctation of clypeus fine, not dense. Punctures of vertex slightly larger than those of clypeus.

Anterior margin and base of pronotum without border, sides distinctly bordered. Lateral border of pronotum thick, keel-shaped, rimmed. Sides of pronotum rounded, pronotum widest approximately at the midlength. Anterior angles rounded, posterior angles widely rounded. Punctation of pronotum almost imperceptible, very fine and shallow; punctures scarcely distributed, distance between punctures equal to several puncture diameters. Laterally with denser, double punctation with medium size punctures intermixed with small punctures; medium size punctures more abundant. Small punctures only on surface near anterior margin, with sparse distribution. Base of pronotum slightly narrower than base of elytra. Pronotum at widest point distinctly wider than maximum width of elytra.

Elytra shagreened (35x), feebly glossy; strongly shagreened, matt near apex. Scutellum triangular, elongate; sides arcuate; surface shagreened, with several fine, shallow punctures. Elytral striae narrow and shallow, punctures of striae fine and shallow, margins of intervals feebly crenate. Distance between punctures approximately twice their width. All intervals elevated, rounded; with distinct, irregularly distributed punctures. In 3 lateral intervals the punctures small; near apex each puncture bears a distinct, yellow seta. Epipleura visible from base to apex in dorsal view, narrowly bordered and distinctly rimmed. Elytra with small, almost indistinct humeral bulges; without humeral teeth.

Protibiae with 3 outer teeth, outer margin from base of tibia to the basal tooth serrate; with long, narrow yellow setae. Apical spur of protibiae narrowed towards tip, the tip curved downwards (visible in lateral view) reaching base of the protarsomere 2. Protarsal spur inserted opposite to the apical outer tooth. Lower terminal mesotibial spur arcuate, not reduced, less than twice length of upper spur; upper spur longer than mesotarsomere 1, thus reaching base of the mesotarsomere 2. Apical setae of mesotibiae and metatibiae unequal in length. Metatarsus longer than the upper terminal spur of metatibiae and equal in length to the 3 following tarsomeres combined.

Aedeagus as in Figs. 2–3.

Female ( Fig. 4). Body more convex. Head smaller, border of clypeus not thickened in the middle. Elytra apex impunctate and completely glabrous. Base of elytra relatively wider, wider than base of pronotum. Pronotum at widest point distinctly narrower than maximum width of elytra.

Distribution. China (Yunnan).

Name derivation. The paper is a collective work of both authors and we had previously agreed on another specific name for the new species. Unfortunately, Radek Červenka, my friend and first author of this paper, died tragically while we were finalizing the manuscript. As a tribute to his memory, I have unilaterally decided to change the species name to “ radekcervenkai ”.

Differential diagnosis. The new species differs from the similar species Paracrossidius transmontanus ( Balthasar, 1932) as follows. For this comparison, we used a syntype of Paracrossidius transmontanus from Chongqing Province, China deposited in the National Museum, Prague, Czech Republic.

Paracrossidius transmontanus ( Figs. 5–8 ) Paracrossidius radekcervenkai ( Figs. 1–4) Body larger (8.7–11.4 mm) Body smaller (6.5–8.8 mm)

Head conspicuously large, relatively wider, maximum width Head relatively narrower, width at the widest point distinctly approximately as wide as that of pronotum narrower than maximum width of pronotum Clypeus almost semicircular, anterior margin rounded Clypeus trapezoidal, anterior margin almost straight Elytra in males setaceous on entire surface except for narrow Elytral in males on most of the surface glabrous, with setae at strip at the base apex only

Punctation of elytra fine, shallow, sparse; distance between Punctation of elytra coarse and dense, distance of punctures equal punctures equal to or larger than two times their diameter or subequal to their diameter

Metatarsus relatively longer, approximately ¼ longer than upper Metatarsus relatively shorter, only slightly longer than the upper terminal spur of the metatibiae terminal spur of the metatibiae

Aedeagus as in Figs. 6–7 Aedeagus as in Figs. 2–3

Distribution: China (Sichuan, Chongqing) Distribution: China (Yunnan)

Figures

FIGURES 5 – 8. Paracrossidius transmontanus (Balthasar, 1932). 5, male; 6, aedeagus in dorsal view; 7, aedeagus in lateral view; 8, female.