Palaeopsilotreta burmanica, Wichard & Espeland & Müller & Wang, 2020

Wichard, Wilfried, Espeland, Marianne, Müller, Patrick & Wang, Bo, 2020, New species of caddisflies with bipectinate antennae from Cretaceous Burmese amber (Insecta, Trichoptera: Odontoceratidae, Calamoceratidae), European Journal of Taxonomy 653, pp. 1-17 : 7

Palaeopsilotreta burmanica sp. nov.

urn:lsid:zoobank.org:act:12B72DE2-74A4-4154-ACFF-6A3B2607DBEC

Fig. 2 , 4F

Diagnosis

Antennae in males with 10 bipectinate flagellomeres and last 2 flagellomeres simple, not bipectinate; in forewing, media simple, unbranched; in male genitalia, harpago apically with 2 clear, thorn-like, black spines. Palaeopsilotreta burmanica sp. nov. is clearly distinguished from P. cretacea sp. nov. by the unbranched media in forewings, P. cretacea sp. nov. has the media branched in forewings. Palaeopsilotreta burmanica sp. nov. is distinguished from P. xiai by the harpago having 2 strong thornlike, black spines at its apex; in P. xiai , the harpago has a crown of short black spines at its apex.

Etymology

Palaeopsilotreta burmanica sp. nov. is named after Burma, the old name of the country where the amber inclusion was found.

Material examined

Holotype

MYANMAR – Kachin State • ♂; exact locality unknown; Mid-Cretaceous Burmese amber inclusion; former Patrick Müller collection, BUB 2885 ; ZFMK-TRI000813 . Body well preserved; forewing in dorsal view, hind wing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view.

Description

HEAD. Light brown, laterally with prominent eyes. Antennae partially bipectinate, nearly as long as forewings. The bipectinate antennae ( Fig. 2B ) consist of long scapus, approximately twice as long as short pedicellus, followed by short flagellomeres as long as pedicellus, then 10 successive bipectinate flagellomeres and 2 simple terminal flagellomeres without bipectinate rami. Rami originating at base of each bipectinate flagellomere slender, about as long as flagellomere. Maxillary palps 5-segmented, terminal segment not flexible or annulate; labial palps 3-segmented ( Fig. 4F ). Head with ocelli absent, setose warts present: pair of small, ovoid interantennal setose warts, pair of postantennal (anterior) setose warts and pair of large, ovoid posterior setose warts. Pro- meso- and metanotum and their setose warts destroyed.

FOREWINGS ( Fig. 2C ). Light brown, apically rounded, about 2.5 mm in length. Vein R1 running towards wing margin but turning slightly to fork R2 without reaching it. Forks I and II present; discoidal cell open, crossveins not visible. Open DC significantly shorter than forks I and II. Media simple, running to apical margin, unbranched into M1+2 and M3+4. Medial and thyridial cells absent. Cu1 running straight, apically bifurcated into fork V; at the fork junction oblique crossvein to M. Cu2 simple, apically crossvein to Cu1b and another crossvein to lateral wing margin. Anal veins complete. Hindwings light brown, dorsally covered by forewings, smaller than forewings, about 2 mm long. Radius R1 vein slightly thickened. Forks I, II and V visible, as well as media M and cubitus Cu2.

TIBIAL SPURS. 2/4/4.

MALE GENITALIA.Visible only in ventral view ( Fig. 2 D–E): inferior appendages(gonocoxites)2-segmented, with coxopodiites and harpago clear and characteristic. Coxopodites broad and scalelike, longer than harpago, which is thin and pen-shaped, slightly curved towards middle of genital area, with 2 thorn-like black spines at apex. Preanal appendages and lobes of segment X not clearly visible, concealed by lower appendages; details of phallic apparatus not sufficiently well preserved.

Figures

Fig. 2. Palaeopsilotreta burmanica sp. nov. (ZFMK-TRI000813) A. Holotype male in dorsal view. B. Head with maxillary palps and bipectinate antennae. C. Drawing of a forewing. D. Male genitalia in ventral view. E. Drawing of the male genitalia.

Fig. 4. Palaeopsilotreta xiai Wichard & Wang, 2017. A. Male and possible female in one Baltic amber piece (ZFMK-TRI000815).B. Female with simple, filiform antennae in dorsal view (ZFMK-TRI000816). C. Female genitalia, as far as visible in cloudy amber (ZFMK-TRI000816). D. Drawing of female forewing. E. Head of male (ZFMK-TRI000811) in dorsal view with antennal, posterior and pronotal setose warts (s.w.). — F. P. burmanica sp. nov. (ZFMK-TRI000813), head of male with maxillary palps (mp), labial palps (lp) and basal bipectinate antennae with scapus (sc) and pedicellus (pe).