Pachyonychis paradoxus Clark 1860

Roie, Martijn Van, Clark, Shawn, Konstantinov, Alexander, Furth, David & Linzmeier, Adelita M., 2023, A paradox indeed: description of genitalia and clarification of the subtribal classification of Pachyonychis paradoxus Clark and Pachyonychus paradoxus Melsheimer (Coleoptera: Chrysomelidae: Galerucinae: Alticini), Zootaxa 5227 (1), pp. 127-136 : 129-132

Pachyonychis paradoxus Clark 1860

( Figures 2 , 3 )

Pachyonychus dimidiaticornis Dejean 1836:384 (nomen nudum)

Pachyonychis paradoxus Clark 1860:63 + Plate II Fig. 7 (type locality: North America (Pennsylvania ))

Hamletia dimidiaticornis Crotch 1873:59 (replacement name for Pachyonychis paradoxus Clark 1860 ). Mignot 1969:100 (synonymy).

Material examined: Lectotype: 67.56; [illegible]; Pachyonychis paradoxus 163 Mels [pz.ac 3] Clark [illegible] dimidiaticornis Pensylvania Dg C 3p; Hamletia dimidiaticornis Crotch (1 syntype BMNH, male). Non-type material: 1 ♀: St. Simon Island, GA; 20-VII-1931, Quersfeld; Property of USNM; USNM 2037268; Pachyonychis paradoxus H. Clark det. A.S. Konstantinov, 2005 ( USNM); 1♁: Mobile, ALA, H. P. Loding; male; Hamletia dimidiaticornis 15864 l.g.g. Cr.; Property of USNM ( USNM); 1♁: Jacksonville Fla.; Coll Hubbard & Schwarz; Hamletia dimidiaticornis Cr. ; Property of USNM ( USNM); 1♀, dissected: Myrtle Beach, Horry Co. S. C., Apr. 22. 1919; ER Kalmbach Collector; Property of USNM; Hamletia dimidiaticornis Crotch det. H. F. Wickham ( USNM).

Distribution: USA — Alabama, Florida, Georgia, Maryland, North Carolina, New Jersey, Pennsylvania, South Carolina. ( Riley et al. 2003, CSM personal data).

Body size: length 4.1–4.9 mm, width 2.1–2.5 mm, illustrated specimen— 4.9 mm long.

Description of female genitalia. Spermathecal pump clearly separated from receptacle, about two-thirds as long as receptacle, slightly arched, with top rounded. Apex of spermathecal pump flattened; length about one-third of spermathecal pump. Receptacle elongate, constricted at spermathecal pump, forming clear neck; maximum width situated at about one-third from basal part of pump, followed by slight constriction. Canal long, attached to base of receptacle, slightly widened at connection with duct, with two coils. Ramus widening near top, attached to canal via short bifurcation. Gland elongate. Posterior sclerotization of tignum broadly Y-shaped, much wider than midsection. Midsection of tignum nearly straight. Anterior sclerotization of tignum about as wide as midsection; apex blunt. Vaginal palpi shaped as conical funnels. Apex of vaginal palpus evenly rounded, facing anteromedially. Anterior sclerotization of vaginal palpus much wider than posterior sclerotization. Posterior sclerotization of vaginal palpi bearing multiple setae.

Comments. The lectotype for Pachyonychis paradoxus Clark is designated here to fix the identity of this species.

Figures

FIGURE 2. Habitus of Pachyonychis paradoxus Clark 1860. A: drawing of dorsal view (by Taina Litwak); B: Dorsal view; C: Frontal view; D: Lateral view; E: Ventral view.

FIGURE 3. Abdomen and female genitalia of Pachyonychis paradoxus Clark 1860. A: Apical tergites; B: Sternites; C: Spermatheca; D: Tignum; E: Vaginal palpi.