Orthovertellopsis proteaeformis, Vachard, Krainer & Lucas, 2015

Vachard, D, Krainer, K & Lucas, SG, 2015, Late Early Permian (late Leonardian; Kungurian) algae, microproblematica, and smaller foraminifers from the Yeso Group and San Andres Formation (New Mexico; USA), Palaeontologia Electronica (English ed.) 3 (8), pp. 1-77 : 40-42

Orthovertellopsis proteaeformis n. gen. n. sp.

Figures 22.2 , 23.2, 23.4, 23.6 , 33.19, 33.20 , 35.10, 35.12

zoobank.org/ 47A263E8-C536-4705-B36C-7CEF5B7CA43D

? 1958 Orthovertella protea Cushman and Waters ; Crespin, p. 82, pl. 18, figs. 7-9.

1985 Orthovertella protea Cushman and Waters ; Foster, Palmieri and Fleming, p. 81, pl. 5, figs. 5-7.

? 1993 Neodiscus cf. permicus (Grozdilova) ; Ueno and Sakagami, fig. 3.14.

? 2006 Hemigordius sp. 1 ; Nestell, Nestell, Wardlaw and Sweatt, pl. 1, fig. 21 (only).

v. 2013 b Orthovertella ? sp. 1 (part.); Vachard,

Krainer and Lucas, p. 348 (no illustration).

v. 2013 b Orthovertella ? sp. 2; Vachard, Krainer and

Lucas, p. 348 (no illustration).

Etymology. Similar to some O. protea Cushman and Waters, 1928a of the authors.

Holotype. NMMNH P-67570; Figure 22.2 ; sample

MG 10-21.

Paratypes. NMMNH P-67571-67573; Figure 23.2,

23.4, 23.6.

Material. 10 specimens.

Locus Typicus. NMMNH locality 8901; Yeso Group, Los Vallos Formation, Torres Member, Massacre Gap in the southern Fra Cristobal Mountains (New Mexico).

Stratum Typicum. Early? Kungurian (late Leonardian).

Diagnosis. Large and elongate species of Orthovertellopsis with a large or huge proloculus and a test, inflated in the central part, entirely streptospiral or with the last or semi-last whorl semi-evolute. Wall relatively thick and lumen approximately three times wider than the wall.

Description. The test is large and elongate; D = 0.60-0.85 mm; w = (rarely 0.20)- 0.34-0.41 mm; w/ D = (rarely 0.33-0.41)-0.51-0.58. The spherical proloculus is large to huge (up to 0.09 mm). The coiling, initially weakly streptospiral, becomes oscillating and forms a thin-walled glomus, and is then sigmoidal with all the whorls thick-walled, evolute, and with deep sutures. Asymmetrical umbilici are present. Whorls: 4-5-(6), involute at the beginning, then semi-involute and finally evolute. Sutures marked between the different whorls of coiling. Lateral sides of spire irregular. Peripheral margin round. Tube walls relatively thick (0.02-0.03 mm). Lumina semi-circular (h = 0.09-0.13 mm). Uncoiling inconspicuous. Some basal lateral thickennings in the wall are obvious.

Remarks. Our sections are very similar to those of Foster et al. (1985) due to the evolute last whorls and the deep sutures that evoke the concavoangulatus stage of the archaediscoids ( Pille et al., 2007; Pille, 2008). The uncoiled last stage of coiling is inconspicuous in our thin sections. The new species differs from O. protea sensu stricto by the regular elongate test, larger dimensions, less conspicuous uncoiled last part, and different ages (early Late Carboniferous against late Early Permian). It differs from O. protea sensu Crespin, 1958 or Foster et al., 1985 by the elongate test and smaller dimensions.

Occurrence. Early Permian Australia (Carnarvon Basin, Canning Basin, Tasmania, and perhaps Sydney Basin). Early?-middle? Kungurian (Leonardian) of New Mexico (Yeso Group, Los Vallos Formation, Torres Member: samples MG(2) 10-6, MG 10-13a, MG(2) 10-14, MG 10-20, MG 10-21, MG 10-23. San Andres Formation, McLeod Hills: samples MLY 6-5, MLY 6-5a).?Early Murgabian (= Rodian) of NE Thailand.

Figures

FIGURE 22. 1, 3. Glomomidiella infrapermica n. sp. 1. NMMNH P-67600, holotype. Subtransverse section. Yeso Group. Sample MG 10-22. 3. NMMNH P-67601, paratype. Subtransverse section. Yeso Group. Sample MG 10-25. 2. Orthovertellopsis proteaeformis n. gen. n. sp., NMMNH P-67570, holotype. Axial section with relatively well-preserved wall. Yeso Group. Sample MG 10-21. 4, 7-8, 10, 12-14. Praeneodiscus convexus n. gen. n. sp. 4. NMMNH P-67606, holotype. Axial section. Yeso Group. Sample MG 10-24. 7. NMMNH P-67607, paratype. Axial section. San Andres Fm. Sample MLY 5-13 (already illustrated in Brose et al., 2013, figure 6. 11). 8. NMMNH P-67608, paratype. Axial section. San Andres Fm. Sample MLY 5-4. 10. NMMNH P-67609, paratype. Axial section with a relatively preserved wall. San Andres Fm. Sample MLY 5-7. 12. NMMNH P-67610, paratype. Axial section. San Andres Fm. Sample MLY 5-7a. 13. NMMNH P-67611, paratype. Axial section. San Andres Fm. Sample MLY 5-30. 14. NMMNH P-67612, paratype. Axial section. San Andres Fm. Sample MLY 5-21. 5. Tubiphytes epimonellaeformis n.sp. Subtransverse section. San Andres Fm. Sample SAR 8a-22. 6. Globivalvulina parapiciformis n. sp. NMMNH P-67551, holotype. Transverse section. San Andres Fm. Sample MLY 4-3. 9. Pseudospira aff. ishimbaica (Lipina, 1949). Subtransverse section. Yeso Group. Sample MG 10-26. 11. Hemigordiellina? cf. simplex (Harlton, 1928). Axial section. San Andres Fm. Sample MLY 6-28. 15. Eotuberitina reitlingerae Miklukho-Maklay, 1958. Axial section. San Andres Fm. Sample MLY 4-5. Scale bars equal 0.10 mm.

FIGURE 23. 1, 3, 7-8. Glomomidiella infrapermica n. sp. 1. NMMNH P-67602, paratype. Subtransverse section. Yeso Group. Sample MG 10-12. 3. NMMNH P-67603, paratype. Oblique subaxial section. Yeso Group. Sample MG 10-15. 7. NMMNH P-67604, paratype. Two subtransverse sections. Yeso Group. Sample MG 10-2. 8. NMMNH P-67605, paratype. Subtransverse section. Yeso Group. Sample MG 10-16. 2, 4, 6. Orthovertellopsis proteaeformis n. gen. n. sp. 2. NMMNH P-67571, paratype. Subaxial section. Yeso Group. Sample MG 10-23. 4. NMMNH P-67572, paratype. Subaxial section. Yeso Group. Sample MG 10-13a. 6. NMMNH P-67573, two paratypes. Subtransverse section. Yeso Group. Sample MG 10-20. 5. Calcitornella cf. elongata Cushman and Waters, 1928a. Subaxial section. Yeso Group. Sample MG 10-20a. Scale bars equal 0.10 mm.

FIGURE 33. 1-14, 16, 18. Hemigordiellina? cf. simplex (Harlton, 1928). 1. Oblique subtransverse section. San Andres Fm. Sample MLY 3-1. 2. Oblique subaxial section. San Andres Fm. Sample MLY 3-11. 3. Subtransverse section. San Andres Fm. Sample MLY 4-8. 4. Subaxial section. San Andres Fm. Sample MLY 4-11. 5. Transverse section. San Andres Fm. Sample MLY 4-15. 6. Subtransverse section. San Andres Fm. Sample MLY 4-16. 7. Transverse section. San Andres Fm. Sample MLY 4-18. 8. Subaxial section. San Andres Fm. Sample MLY 4-19. 9. Subtransverse section. San Andres Fm. Sample MLY 8-7. 10. Subtransverse section. San Andres Fm. Sample MLY 4-20. 11. Subtransverse section. San Andres Fm. Sample MLY 5-15. 12. Subaxial section. San Andres Fm. Sample MLY 5-16a. 13. Transverse section. San Andres Fm. Sample MLY 5-5. 14. Subtranverse section. San Andres Fm. Sample MLY 4-12. 16. Oblique section. San Andres Fm. Sample MLY 6-25a. 18. Transverse section. San Andres Fm. Sample MLY 4-21. 15, 21. Pseudospira aff. ishimbaica (Lipina, 1949). 15. Subaxial section. San Andres Fm. Sample MLY 6-22. 21. Axial section with Hemigordiellina? cf. simplex (left). San Andres Fm. Sample MLY 5-17. 17, 22, 26. Praeneodiscus convexus n. gen. n. sp. 17. Oblique section. San Andres Fm. Sample MLY 5-16. 22. Transverse section. San Andres Fm. Sample MLY 5-8a. 26. Subaxial section. San Andres Fm. Sample MLY 5-9. 19-20. Orthovertellopsis protaeformis n. gen. n. sp. 19. Subaxial section. San Andres Fm. Sample MLY 6-5. 20. Oblique subaxial section. San Andres Fm. Sample MLY 6-5a. 23-24. Hemigordiellina? aff. pseudopusilla (Baryshnikov in Zolotova and Baryshnikov, 1980). 23. Oblique section. San Andres Fm. Sample MLY 5-25. 24. Axial section. San Andres Fm. Sample MLY 5-24. 25. Palaeonubecularia? sp. Axial section. Perhaps also related with Ellesmerella (see Vachard et al., 2013b). San Andres Fm. Sample MLY 5-29. Scale bars equal 0.10 mm.

FIGURE 35. 1-5, 8. Olgaorlovella davydovi n. gen. n. sp. 1. Axial section. San Andres Fm. Sample SAR 11a-11. 2. Axial section. San Andres Fm. Sample SAR 11a-8 (already illustrated in Brose et al., 2013, figure 6. 12). 3. Subaxial section. San Andres Fm. Sample SAR 11a-15. 4. Axial section. San Andres Fm. Sample SAR 11a-14. 5. Axial section. San Andres Fm. Sample SAR 11a-3. 8. Axial section. San Andres Fm. Sample SAR 11-6. 6-7, 13. Orthovertellopsis calcitornellaeformis n. gen. n. sp. 6. Subtransverse section. San Andres Fm. Sample SAR 11a-16. 7. Transverse section. San Andres Fm. Sample SAR 11-16. 13. Oblique section. San Andres Fm. Sample SAR 11a-5. 9, 11, 14. Glomomidiella infrapermica n. sp. 9. Axial section showing the proloculus. San Andres Fm. Sample SAR 11-14. 11. Subtransverse section. Yeso Group. Sample MG(2) 10-7b. 14. Axial section relatively well preserved (dark wall). Yeso Group. Sample MG(2) 12-7. 10, 12. Orthovertellopsis protaeformis n. gen. n. sp. 10. Axial section. Yeso Group. Sample MG(2) 10-14. 12. Subaxial section. Yeso Group. Sample MG(2) 10-6. Scale bars equal 0.10 mm.