Orthomus barbarus subsp. penibeticus, Mateu & Colas, 1954: 53, 1828

Guéorguiev, Borislav & Wrase, David W., 2016, On the identity of Orthomus hespericus Motschulsky, 1849 (Coleoptera: Carabidae: Pterostichini), Ecologica Montenegrina 5, pp. 28-34 : 32

6.3. barbarus penibeticus Mateu & Colas, 1954: 53 Spain

Note 2: Formerly, the taxa formenterrae and penibeticus were treated as subspecies of O. abacoides by Zaballos and Jeanne (1994) and Ortuño (1996). The contrary is the view of Serrano (2003, 2013) and Bousquet (2003) who have treated them as subspecies of O. barbarus . Recently, Pupier and Coulon (2013) made a detailed study of the female genitalia of two groups of Orthomus , including the genitalia of the aforementioned three taxa. The French authors showed that the spermathecal complex of both formenterrae and penibeticus is of the type “ barbarus ”, while that of O. abacoides is of the type “ abacoides ”, as clear morphological differences exist between the two morphological types. They concluded (ibid.: 223): “Quant а Orthomus abacoides (Lucas, 1846) , il s‟agit d‟un endémique de l‟ Algérie occidentale; il ne semble pas dépasser au sud les monts de Tlemcen et les monts de Daïa, où il serait remplacée par O. lacouri . On ne peut admettre que O. penibeticus Mateu et Colas, 1954 et O. formenterrae (Breit, 1933) en soient des sousespèces, leurs genitalia femelles étant du type « barbarus ›› (voir fig. 4f et Ortuño, 1996)”.