Orizabus mcclevei, Warner, 2011

Warner, William B., 2011, Review of the genus Orizabus Fairmaire in the United States of America (Coleoptera: Scarabaeidae: Dynastinae), Insecta Mundi 2011 (174), pp. 1-42 : 11-13

publication ID	https://doi.org/ 10.5281/zenodo.5160884
persistent identifier	https://treatment.plazi.org/id/03BA87C3-BC4B-FF88-F9E9-FBE4952A6F9D
treatment provided by	Felipe
scientific name	Orizabus mcclevei
status	sp. nov.

Treatment

Orizabus mcclevei new species

Fig. 5 View Figures 5-7 , 14, 15 View Figures 12-15 , 20, 21 View Figures 20-25 , 35 View Figures 32-35

Types. Holotype male and allotype female labeled: “NM: Hidalgo Co., Peloncillo Mts., Clanton Draw , vii- 7-1991, 5500’, UV lite [sic], S. McCleve ”.

Paratypes (36 males, 46 females) from the following localities: USA, Arizona: “Cochise Co., Douglas, vii-3-1971, at light, S. McCleve”; “Cochise Co., Rucker Canyon, vii-28-1977, D. Ahart”; “Cochise Co., Chiricahua Mts., N. Fork Rucker Cyn., vii-1-1978, at light, S. McCleve”; “Cochise Co., Guadalupe Canyon, elev. 4300 ft., vii-26-1976, P. Jump”. Colorado: “Baca Co., Sand Canyon, vii-13-1991, W. Cranshaw, M. Kippenhan, B. Kondratieff, D. & M. Leatherman, & P. Opler” (3); “Costilla Co., H Rd., 3 mi. E of Mesita, viii-21-1993, Kondratieff, Al-Ayedh & Stone”; “Las Animas Co., Purgatoire River, Lockwood Arroyo, vii-2-1989, B. Kondratieff & J. Welch”; “Powers Co., Two Buttes Dam, vii-21-1990, U.V.L., P. A. Opler”. New Mexico: same data as holotype (3); same except “ vii-6-1988, R. Gordon” (2); “Cherry Cr. campground N of Silver City, 7100 ft. elev., vii-25-1972, at light, S. McCleve”; same except: “ vii-26-1972 ”; “Grant Co., Silver City, vii-1973, R. Lenczy” (5); “Roberts Lake, 30 mi. NE Silver City, 6100’, viii-1-3- 1973, E. Lindquist”; “Corona, vi-20-1952, R.H. & L. D. Beamer, W. LaBerge, A. Wolf, C. Liaug, C. Winer” (3); “Otero Co., James Canyon campground, Lincoln National Forest, BL, viii-19-1982, R. Turnbow”; “Valencia Co., 20 mi. W Los Lunas, Carrizon Arroyo, viii-1-20-1977, S. & J. Peck” (2); “Las Vegas, Near Hot Springs, 7000 ft., vii-1882, F. H. Snow”; “Pecos, viii-18”; “Albuq., Wickham”; “Placitas, viii-9-1972, P. M. Jump”; “Sierra Co., N fork of Palomas Creek, 33 mi. N of Williamsburg, viii-10-1990, R. Durfee”; “Socorro Co., 22 mi. E San Antonio, sand hills, vii-25-1996, Wappes & Huether (5)”; same except: “ vii-25- 26-1996 ” (10); “Santa Rosa, viii-11-1973, R. Battelle”. Texas: “Tex.”; “Jeff Davis Co., Davis Mountain Resort, 5800 ft, v-25-1994, D. G. Marqua”; same except: “ v-28-1994 ”; same except: “ vi-14-1994 ” (2); same except: “ vi-16-1991 ”; same except: “ vi-22-1992 ”; same except: “ vi-26-1992; same except: “ vii-4-1993 ”; same except: “ vii-5-1992 ”; same except: “ vii-14-1994 ” (5); same except: “ vii-22-1991 ”; same except: “ vii-23-1986 ”; same except: “ vii-23-1993 ”; same except: “ viii-3-1991 ”(2); same except: “ viii-3-1990 ”; same except: “ viii-6- 1991 ”; same except: “ viii-18-1987 ”; same except: “ viii-22-1993 ”; same except: “D. Marqua house, viii-8- 1992, BL, W. Godwin, E. Riley, D. Marqua”; same except: “Davis Mtns. Resort area, vii-15-1994, C.

Wolfe”; “Jeff Davis Co., Davis Mtns., Madera Cyn., vii-9-1993, D. G. Marqua”; “Jeff Davis Co., Madera Canyon rest stop, vi-11-1991, C. S. Wolfe” (3); “Jeff Davis Co., vic. Davis Mts., vii-19-22-1995, J. Huether” (2). Mexico, Durango: “Hwy. 45, 5 mi. N Rodeo, 6 mi. W, vii- 16-1984, 4980 ft., S. McCleve & P. Jump, UVL”.

Holotype and allotype are deposited in FSCA. Paratypes are deposited in the collections of ASUT, BCRC, CNCI, CSUC, DACC, DGMC, CSWC, EGRC, EMEC, FSCA, HAHC, JHUC, MCZC, RACC, SEMC, SMCC, TAMU, UAIC, UCRC, UNSM, USNM, and WBWC.

Diagnosis. Head with clypeal carina transverse, straight; antennal club longer than distance between frontoclypeal tubercle and clypeal carina; anterior pronotal margin at middle with bead posteriorly prolonged into obtuse angle; protibia tridentate, with apex transversely rounded to subtruncate; paramera laterally dentate at about apical third.

Description. Holotype male, length 15.5 mm, width at humeri 7.5 mm, widest width (apical one-third of elytra) 9.5 mm. Color reddish brown; body convex, pyriform ( Fig. 14 View Figures 12-15 ). Head with frons and clypeus punctatorugulose; frontoclypeal tubercle transverse, about two-thirds as wide as frontoclypeal suture, in anterior view obtusely angulate to medial summit. Clypeus parabolic, expanding behind antennal insertions, apex reflexed; clypeal carina transverse, edentate, obviously subapical (slightly proximal to apical fourth of clypeus). Antennal club five-fourths length of clypeus, elongate-luniform in apical view. Pronotum half as long as elytra, disc confluently variolately punctate anteriorly, punctures becoming smaller and sparser posteriorly, mostly impunctate along midline; anterior angles subobtuse, marginal bead complete except posterior margin anterior to scutellum where bead is continued as line of discrete punctures; posterior edge of anterior marginal bead posteriorly broadened to obtuse angle at midline. Scutellum parabolic, impunctate, midline weakly impressed. Elytra broad (length 11:9 width), distinctly umbilicately striate-punctate, impunctate between striae except in more or less single rows in basal two-thirds of first interval, posterior half of second interval, between humeral and apical umbones in third interval, and in posterior half of fourth and fifth intervals. Epipleuron distinct to about level of pygidium, of nearly even width throughout. Pygidium convex, transversely subtriangular, basal margin convexly arcuate; apex narrowly truncate; subapical edge of disc forming transverse gibbosity overhanging excavated apical submargin; free (=apical) margin with bead complete; disc scabriculously punctulate in basal half, shallowly sparsely punctulate in apical half. Venter with prosternite glabrous except anteromedial apex and on conical but laterally compressed prosternal process, other thoracic sternites, episternites, mesepimera and posterior coxae densely villously punctulate, setae ferrugineous; metasternite glabrous medially. Abdominal sternites each with transverse row of setigerous punctures, row mediodiscal in basal sternites, apicomarginal in apical sternite; penultimate sternite about twice as long as each more proximal sternite; apical sternite apicomedially emarginate. Legs: protibia with apex narrowly rounded-truncate ( Fig. 5 View Figures 5-7 ), ventral spur arising at about middle, nearly half as long as tibia; dorsal margin tridentate, apical 2 teeth closer together than middle and basal teeth; basal tooth slightly proximal to middle of tibia. Metatibia with submedial transverse carina distinct, but with ventral terminus not forming angular tooth. Paramera ( Fig. 20, 21 View Figures 20-25 ) in lateral view narrowed to about half width of base and deflexed in apical half, with weak dorsal tooth at about apical third; in dorsal view strongly narrowed and very thin in apical half.

Allotype female: length 17.0 mm, width at humeri 8.0 mm, widest width 10.5 mm. Similar to male except abdomen more convex; pygidium less convex, with free submargin impressed each side of apex; apical abdominal sternite longer and not apicomedially emarginate.

Variation. Length (males and females) 15.5 to 18.0 mm, width at humeri 7.0 to 9.5 mm, widest width 9.0 to 11.5 mm. Other than in size and slight differences in surface sculpture, the specimens examined show little variation. The male from Durango is less heavily punctate and has the lateral pronotal margins slightly more arcuate than in US specimens, but otherwise (including genitalia) is similar.

Etymology. I am pleased to name this new species in honor of Scott McCleve of Douglas, Arizona, who first provided me with specimens of this new species.

Remarks. This species is similar in size, shape, and general genital form to O. pyriformis , however it is easily differentiated from that species by its longer antennal clubs, obviously subapical and straight clypeal carina, broadly transverse frontoclypeal tubercle, shorter, more deflexed paramera, and pronotum with (usually) subobtuse anterior angles, angularly broadened (at midline) anterior marginal bead, and a more or less complete posterior marginal bead. Aphonus texanus Gill and Howden is also similar in size and dorsal facies, but has a more finely and densely sculptured (“shagreened”) pygidium, different male genitalia, obsolescent frontoclypeal tubercle, and tridentate (when not worn) clypeal carina. Orizabus mcclevei also is somewhat similar to O. isodonoides ; however, O. isodonoides has broader antennal clubs (in apical view), a transverse carina instead of a clypeal (albeit transverse) tubercle, pronotum lacking the medially broadened anterior marginal bead, in males a medial pronotal tubercle separate from and immediately posterior to the anterior marginal bead with a more or less distinct and broad concavity posterior to the tubercle ( Fig. 18 View Figures 16-19 ), and less strongly deflexed paramera which in dorsal view are more strongly “bowed” and have the external tooth more developed ( Fig. 23, 24 View Figures 20-25 ).

Nothing is known of the biology of O. mcclevei other than that it is attracted to lights at night, and that it is primarily (but not entirely) a montane species.